Corresponding author: Roger A. Burks (
Academic editor: Michael Engel
The Australasian and southwest Pacific species of
The genus
This is the second of three studies aimed at revising the global fauna of
Previously, 14 species were described from the region, all from the Australian mainland and all prior to 1930.
The contributions of the individual authors are as follows; R.A. Burks: character definition, species concept development; key development, imaging, capture of specimen data, manuscript preparation, phylogenetic analysis and illustration; L. Masner: specimen acquisition, and generic overview; N.F. Johnson: generic concept development and manuscript preparation; A.D. Austin: project planning, species concept discussions, manuscript preparation, and taxonomic overview.
Specimens examined were provided by the following collections: The American Entomological Institute, Gainesville, Florida, USA Australian Museum, Sydney, Australia Australian National Insect Collection, Canberra, Australia New South Wales Department of Primary Industries, Agricultural Scientific Collections Unit, Orange, Australia The Natural History Museum, London, United Kingdom Canadian National Collection of Insects, Arachnids and Nematodes, Ottawa, Canada Illinois Natural History Survey, Urbana, IL Illinois, USA Institut Royal des Sciences Naturelles de Belgique, Bruxelles, Belgium Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, USA Muséum National d’Histoire Naturelle, Paris, France Museum of Victoria, Entomology, Melbourne, Australia National Museum of Natural History, Washington, DC, USA C.A. Triplehorn Insect Collection, Ohio State University, Columbus, Ohio, USA Queensland Primary Industries and Fisheries Insect Collection, Indooroopilly, Australia Queensland Museum, Brisbane, Australia Queensland Primary Industries Insect Collection, Mareeba, Australia Royal Museum of Central Africa, Tervuren, Belgium South Australian Museum, Adelaide, Australia South African National Collection of Insects, Pretoria, South Africa Tasmanian Department of Primary Industries and Water, Hobart, Australia University of California, Riverside, California, USA University of Queensland Insect Collection, Brisbane, Australia National Museum of Natural History, Washington, DC, USA Western Australian Museum, Perth, Australia Waite Insect and Nematode Collection, Adelaide, Australia
This revision is a product of the
Morphological terminology follows
Surface sculpture terminology follows
Diminutive variant terms (such as “foveolate” or “rugulose”) were avoided because of a lack of criteria for separating them from non-diminutive alternatives. “Major” surface sculpture refers to repeated sculptural patterns that interact with seta placement. It does not include non-repeated elements or those which are repeated only once due to bilateral symmetry. “Umbilicate-foveate” sculpture refers to rounded crater-like sculptural elements, each surrounding a setiferous punctum (and thus interacting with a seta), with each fovea being much larger than its setiferous punctum and spatially separated from it (
“Microsculpture” refers to repeated sculptural elements that do not interact with seta placement. Microsculpture can occur on “major” sculptural elements, such as on rugae and on all surfaces of foveae. Punctate microsculpture refers to tiny round pits that do not bear setae (
Examples of surface sculpture in
One of the difficulties in defining surface sculpture using hierarchies and genus-differentia formats is that there is more than one valid way of classifying surface sculpture into Aristotelian genera. We maintain that the art of creating genus-differentia definitions, although ancient, is not yet at a methodologically mature state. Another problem is that variation can render some commonly-used sculptural classifications problematic. Finally, surface sculpture has typically been named and defined using only vague shape or pattern-based comparisons to other, better known entities (such as reticulate = net-like), and not through more purely logical means. Umbilicate-foveate sculpture could be called “reticulate,” for instance, because of a net-like pattern of foveae. However, in some Australian
We assert that the word “pit” is a word but not a specialized term, having terminological importance only as part of a two-word term that includes a specialized term as a qualifier, as in “tentorial pit.” For that reason, we have changed from using “pit” to using “punctum” (the Latin word for “puncture”) here. We make no distinction between types of puncti other than to describe them as setiferous or not, but other criteria for classifying puncti probably exist for
carina: The process that is linear and is regular.
fovea: The depression that is ovate and is shallow relative to its diameters.
granulate: The sculptural pattern that is a network of branched impressions.
interspace: The area that is between sculptural elements.
major sculpture: The sculptural pattern that is associated with seta placement.
microsculpture: The sculptural pattern that is not associated with seta placement.
punctate microsculpture: The sculptural pattern that is comprised of puncta.
punctum: The depression that is ovate and is deep relative to its diameters.
ruga: The process that is linear and is irregular.
sculptural element: The anatomical structure that is one unit in a sculptural pattern.
sculptural pattern: The pattern that is formed by repeated adjacent surface sculpture.
setiferous fovea: The fovea that surrounds a seta.
setiferous punctum: The punctum that surrounds a seta.
sculptural septum: The septum that separates sculptural elements.
surface sculpture: The anatomical structure that is texture.
umbilicate-foveate: The sculptural pattern that consists of setiferous foveae.
umbilicate-punctate: The sculptural pattern that consists of only setiferous puncta.
Body length: 2.6–7.1 mm.
Head shape in dorsal view: weakly transverse, width approximately 1.5x greatest length; subquadrate. Hyperoccipital carina: absent; present. Occipital carina: present, complete medially; present, broadly interrupted medially. Occipital carina sculpture: crenulate. Ocular ocellar line (OOL): OOL < 0.5 ocellar diameter (OD). Dorsal area of frons: convex, without frontal shelf. Antennal scrobe shape: present, unmargined; scrobe margined by carina. Frons sculpture: umbilicate-punctate, with transverse carinae within scrobe; scrobe largely smooth, otherwise with transverse carinae. Submedian carina: absent. Orbital carina: absent. Inner orbits: diverging ventrally. Interocular space(IOS)/Eye height (EH): IOS distinctly less than EH. Interantennal process: triangular in lateral view. Central keel: absent. Antennal foramen opening: oriented laterally on interantennal process. Facial striae: present. Malar sulcus: present. Compound eye size: not significantly reduced. Compound eye setation: absent. Gena: weakly convex, receding behind posterior orbit; convex, distinctly produced behind eye. Clypeus shape: narrow, slightly convex medially, lateral corner not produced. Apical margin of clypeus: with small median point. Labrum: not visible. Mandibular teeth: apex with 2, acute, subequal teeth. Arrangement of mandibular teeth: transverse. Number of maxillary palpomeres: 4. Shape of maxillary palpomeres: cylindrical. Number of labial palpomeres: 2. Number of antennomeres in female: 12.
Number of antennomeres in male: 12. Insertion of radicle into A1: parallel to longitudinal axis of A1. Shape of A1: more or less cylindrical, not flattened. Length of A3 of female: subequal to length of A2; distinctly longer than A2. Number of clavomeres in female antenna: 7; 0. Claval formula of female antenna: A12-A7/1-2-2-2-2-1; A12-A6/1-2-2-2-2-2-2. Arrangement of doubled multiporous plate sensilla on female clava: in longitudinal pairs. Tyloid distribution on male antenna: A5 only. Shape of male flagellum: filiform.
Mesosoma shape in dorsal view: longer than wide. Mesosoma shape in lateral view: longer than high. Medial portion of transverse pronotal carina: weakly indicated laterally; absent. Posterior apex of pronotum in dorsal view: straight, bifid apically to articulate with tegula. Vertical epomial carina: present. Dorsal epomial carina (lateral portion of transverse pronotal carina of
Number of mid tibial spurs: 1. Number of hind tibial spurs: 1. Dorsal surface of hind coxa: smooth. Hind tibia shape: cylindrical, ecarinate. Trochantellus: present.
Wing size of female: macropterous. Wing size of male: macropterous. Tubular veins in fore wing: present. Bulla of fore wing R: absent. Extent of marginal venation of fore wing: R1 reaching and ending at costal margin; distinct marginal or postmarginal veins present. Origin of r-rs in fore wing: arising before (basad of) R/R1 attains costal margin. Structure of basal vein (Rs+M) in fore wing: spectral. Structure of R in hind wing: elongate, extending to costal margin; abbreviated, not attaining costal margin.
Number of externally visible terga in female: 6. Number of externally visible sterna in female: 6. Number of externally visible terga in male: 8. Number of externally visible sterna in male: 7. Shape of metasoma: acuminate, widest submedially. Laterotergites: present, narrow. Laterosternites: present. T1 of female: raised medially into low, rectangular or subelliptical platform, laterally depressed. Relative size of metasomal segments: T2 distinctly largest; T2 and T3 distinctly larger, subequal in size. Terga with basal crenulae: T1, T2. Sublateral carinae on tergites: present on T1. Median longitudinal carina on metasomal terga: absent. Shape of female T6: flattened. Shape of posterior margin of male T7: straight; incised medially. Anterior margin of S1: protruding anteriorly as short sharp extension of median longitudinal carina. Felt fields: absent. Ovipositor type:
The genus is highly diverse and comprises in excess of 200 species worldwide. It has been recorded from Africa, the Indo-Malayan and eastern Palearctic regions (
1 | Male (female unknown): T7 with elongate and flat subparallel apical projections, but metasomal flanges absent from other terga; mesoscutal midlobe anteriorly granulate and without median carina but with very strong notauli ( |
|
– | Female, or if T7 with such apical projections ( |
2 |
2 | Gena in lateral view longer than eye ( |
|
– | Gena in lateral view shorter than eye ( |
3 |
3 | T2 and T3 in females with sublateral depressions present, medially defined by sharp carina ( |
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– | T2 and T3 in females without sublateral depressions. T1 midlobe in males with distinct carinae or T7 apex either truncate ( |
4 |
4 | Metasomal depression setose ( |
|
– | Metasomal depression without setae ( |
5 |
5 | Metascutellum with a median carina that ends in a granulate area posteriorly ( |
|
– | Metascutellum if with median carina (rare), then without posterior granulate area ( |
6 |
6 | Metascutellum convex, lateral rims directed ventrally and not visible in dorsal view ( |
7 |
– | Metascutellum concave, with raised lateral rims ( |
10 |
7 | Metascutellum much longer than broad, fitting in groove formed by lateral pronotal areas ( |
|
– | Metascutellum as broad as or broader than long, not in contact with propodeum ( |
8 |
8 | Face without oblique carina between eye and frontal depression ( |
|
– | Face with oblique carina between eye and frontal depression ( |
9 |
9 | Mesoscutellum with granulate sculpture ( |
|
– | Mesoscutellum without granulate sculpture ( |
|
10 | Interantennal process elongate ( |
|
– | Interantennal process not elongate ( |
11 |
11 | Hyperoccipital carina indicated by a sharp carina ( |
12 |
– | Hyperoccipital carina incomplete ( |
19 |
12 | Mesoscutellum with oblique elevated sculpture in female ( |
|
– | Mesoscutellum in female without oblique elevated sculpture ( |
13 |
13 | Mesoscutal midlobe with median carina ( |
|
– | Mesoscutal midlobe without a median carina ( |
14 |
14 | Lower face crossed by many curved rugae extending from eye across frontal depression ( |
|
– | Lower face not entirely crossed by curved rugae, any transverse rugae present are restricted to frontal depression. Submedian carina variable, but often indicated by a single elevation ( |
15 |
15 | Metascutellum with only a small median concave area, laterally with broad longitudinally striate areas ( |
16 |
– | Metascutellum with broad median concave area, laterally with narrow or no longitudinally striate areas ( |
17 |
16 | Female: occiput completely smooth ( |
|
– | Female: occiput with elevated sculpture ( |
|
17. | Occiput strongly sculptured, with transverse rugae ( |
|
– | Occiput weakly sculptured, almost entirely smooth ( |
8 |
18 | Metascutellum with median carina ( |
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– | Metascutellum without median carina ( |
|
19 | T1 lateral carina expanded and laterally lobe-like ( |
20 |
– | T1 lateral carina simple, not expanded, carina-like ( |
72 |
20 | T6 in female [for male, see couplet 50] with subapical spine-like (cornicle-shaped) dorsally projecting spines ( |
|
– | T6 in female either lacking spines or with flat, apical spines ( |
21 |
21 | Hyperoccipital carina indicated by a strong carina; occiput almost entirely smooth ( |
22 |
– | If hyperoccipital carina strong, then occiput densely sculptured ( |
23 |
22 | Female: mesoscutellum with strong longitudinal carinae and/or rugae ( |
|
– | Female: mesoscutellum with oblique raised sculpture ( |
|
23 | Female: Main body of T6 forming a complete ledge above apical rim, hiding it from dorsal view, instead usually concave and revealing apex of S6 to dorsal view ( |
24 |
– | Female: Main body of T6 either not forming ledge, or with ledge interrupted medially and descending to a clearly visible apical rim, not revealing S6 to dorsal view ( |
35 |
24 | Female: T6 with complete peripheral ledge formed by thin cuticle ( |
|
– | Female: T6 with lateral metasomal flanges or a posterior metasomal flange that is interrupted medially ( |
25 |
25 | Mesoscutal midlobe with transverse carinae ( |
|
– | Mesoscutal midlobe without transverse carinae ( |
26 |
26 | Frontal depression with transverse carinae present in dorsal portion ( |
27 |
– | Frontal depression without transverse carinae in dorsal portion; submedian carina strong, sharp ( |
32 |
27 | Female: A4 longer than broad, A5 as long as or longer than broad ( |
|
– | Female: A4 not longer than broad, A5 broader than long. Male: T6 metasomal flanges (when male known) with rounded projecting lobes ( |
28 |
28 | Mesoscutellar rim without median notch ( |
29 |
– | Mesoscutellar rim with median notch ( |
30 |
29 | Metascutellum tiny ( |
|
– | Metascutellum broad ( |
|
30 | Female: T5 without metasomal flanges, T6 weakly or not at all emarginate apically ( |
|
– | Female: T5 with metasomal flanges, T6 deeply emarginate apically ( |
31 |
31 | Mesoscutal midlobe with strong, coarse foveate sculpture, without smooth areas ( |
|
– | Mesoscutal midlobe with weaker sculpture and with smooth area near midlength ( |
|
32 | Female: T6 deeply emarginate but medially truncate ( |
|
– | Female: T6 V-shaped or rounded medially if deeply emarginate ( |
33 |
33 | Mesoscutellar rim expanded ( |
|
– | Mesoscutellar rim not expanded ( |
34 |
34 | Coxa darker than rest of leg ( |
|
– | Coxa same color as rest of leg ( |
|
35 | T6 abruptly narrower than T5 (sometimes T5 also abruptly narrower than T4), these with strong, straight posterolateral corners ( |
36 |
– | If apical terga abruptly narrower than preceding terga, then some of them other than apical tergum with metasomal flanges ( |
39 |
36 | T5 with subapical rounded protrusions ( |
|
– | T5 without subapical protrusions ( |
37 |
37 | Apical tergum medially blunt or roundly emarginate, with elongate narrow metasomal flanges ( |
38 |
– | Apical tergum medially angular, with sharp posterior corners that are not elongate ( |
|
38 | Submedian carina distinct ( |
|
– | Submedian carina absent medially ( |
|
39 | Female: T6 with lateral ledges that medially slope down to the apical rim ( |
40 |
– | Female: T6 without lateral ledges, without metasomal flanges ( |
48 |
40 | Female: T6 with tiny sharp subapical metasomal flanges, tergum abruptly narrowed posterior to them ( |
41 |
– | Female: T6 with lateral margins forming shelf-like areas ( |
42 |
41 | Coxa darker than rest of leg ( |
|
– | Coxa same color as most of femur ( |
|
42 | Female: T6 apex with strongly protruding posterior rim, main body of tergum anterior to it forming a narrowed shelf laterally, median carina extending from main body of tergum to apical rim; all this together forming an almost trident-shaped apex ( |
43 |
– | Female: T6 with hardly protruding posterior rim, main body of tergum forming broad shelf laterally that is partially raised above apical rim and does not have a median carina; all this together forming an essentially truncate apex ( |
44 |
43 | Female: T4 with broad, flat metasomal flanges ( |
|
– | Female: T4 without metasomal flanges, or with very tiny and sharply pointed metasomal flanges ( |
|
44 | Frontal depression without carinae in dorsal portion ( |
45 |
– | Frontal depression with some carinae in dorsal portion, which may be interrupted medially ( |
46 |
45 | Female (males unknown): Fore wings not long enough to extend beyond T5 ( |
|
– | Female: Fore wings extending beyond T5 ( |
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46 | Female: A4 broader than long. [Males unknown] | 47 |
– | Female: A4 longer than broad. [Male: ( |
|
47 | Female: T6 only shallowly emarginate medially, not sloping to apical rim ( |
|
– | Female: T6 deeply emarginate medially, with a median slope leading down to apical rim ( |
|
48 | Males only: T7 with long posterior spines ( |
49 |
– | Male or female: apical tergum without spines ( |
51 |
49 | A4 longer than broad ( |
|
– | A4 broader than long ( |
50 |
50 | Head and mesosoma weakly metallic blue or green ( |
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– | Entire body non-metallic dark brown, including legs ( |
|
51 | Frontal depression deep and nearly parallel-sided, with many transverse carinae above and below dorsal separator, but submedian carina weak or absent medially ( |
|
– | Frontal depression, if with many carinae, then strongly broadening ventrally and not deep ( |
52 |
52 | Tibiae with spines ( |
53 |
– | Tibiae without spines | 54 |
53 | Frontal depression very broad, with strong submedian carina ( |
|
– | Frontal depression not very broad, submedian carina weak or absent ( |
|
54 | Mesoscutal midlobe posteriorly with many strong longitudinal rugae | |
– | Mesoscutal midlobe posteriorly with few or no longitudinal rugae | 55 |
55 | Hyperoccipital carina indicated by sharp carina ( |
|
– | If hyperoccipital carina sharp and submedian carina accompanied by additional carinae, then mesoscutellum without elevated sculpture ( |
56 |
56 | Frontal depression with carinae or transverse elevations, which may be medially interrupted, above dorsal separator ( |
57 |
– | Frontal depression smooth dorsally ( |
61 |
57 | Coxa darker than rest of leg ( |
58 |
– | Coxa not darker than rest of leg, apical part of legs yellowish ( |
|
58 | Female only (male cannot be reliably keyed past this point): A4 longer than broad | 59 |
– | Female: A4 broader than long | 60 |
59 | Submedian carina complete medially as a distinct carina ( |
|
– | Submedian carina absent medially ( |
|
60 | Postmarginal vein present, venation reaching or nearly reaching anterior wing margin ( |
|
– | Postmarginal vein absent, venation not closely approaching anterior wing margin ( |
|
61 | Entire body dark brown, including antennae and legs | 62 |
– | Some part(s) of legs and antennae yellowish | 64 |
62 | Occipital carina with flat transverse sections connected median arch to lateral areas ( |
|
– | Occipital carina nearly uniformly arched, without flat transverse sublateral sections. ( |
63 |
63 | Female: T1 midlobe longitudinal carinae obscured by smooth area anteriorly. Male: most flagellomeres between A4 and A11 broader than long, T7 blunt apically and without lobes | |
– | Female: T1 midlobe with 4 distinct longitudinal carinae. Male: all flagellomeres longer than broad, T7 emarginate apically and with rounded posterior lobes | |
64 | Coxa darker than rest of leg ( |
65 |
– | Coxa not darker than rest of leg ( |
67 |
65 | Female: fore wing not long enough to exceed metasomal apex; A4 longer than broad; T6 not concave apically ( |
|
– | Female: fore wing long enough to reach or exceed metasomal apex, A4 as broad or broader than long; T6 at least mildly concave apically ( |
66 |
66 | Female: mesoscutellum very sparsely foveate, with broad smooth areas between foveae ( |
|
– | Female: mesoscutellum densely foveate ( |
|
67 | Occipital carina omicron-shaped, with narrow and strongly convex median section connected to lateral sections by a transverse carina ( |
68 |
– | Occipital carina broadly rounded, with no differentiation medially ( |
69 |
68 | Metanotum and posterior rim of mesoscutellum pale, in contrast with surrounding areas ( |
|
– | Metanotum and posterior rim of mesoscutellum not pale ( |
|
69 | Occipital carina absent medially | |
– | Occipital carina present medially | 70 |
70 | Submedian carina weak and indicated by rounded ruga, frontal depression shallow ( |
|
– | Submedian carina sharp and strong, and frontal depression deep ( |
71 |
71 | Dorsal portion of frontal depression without transverse carinae ( |
|
– | Female: Dorsal portion of frontal depression with interrupted transverse carinae ( |
|
72 | Hyperoccipital carina and submedian carina represented by several (more than 3) sharp and distinct rugae ( |
73 |
– | Hyperoccipital carina and submedian carina not both represented by so many rugae ( |
74 |
73 | Female: Fore wing long enough to reach middle of T5; T1 with very strong anterior horn; T6 longer than broad ( |
|
– | Female: Fore wing long enough to reach beyond metasomal apex; T1 with very weak anterior swelling; T6 broader than long ( |
|
74 | Metasomal sterna entirely smooth aside from setiferous puncta ( |
75 |
– | Metasomal sterna with some longitudinal rugae or carinae ( |
76 |
75 | Clypeus elongate ( |
|
– | Clypeus not elongate ( |
|
76 | Epomial corner projecting laterally ( |
77 |
– | Epomial corner not projecting laterally ( |
78 |
77 | Submedian carina present, indicated by rounded ruga ( |
|
– | Submedian carina absent, the area with some convergent heart-shaped sculpture ( |
|
78 | Mesoscutum and mesoscutellum with sunken median longitudinal depression where median carinae would otherwise be; areas lateral to these depressions strongly raised ( |
|
– | Mesoscutum and mesoscutellum without longitudinal depression, without raised submedian areas ( |
79 |
79 | Submedian carina and hyperoccipital carina absent ( |
|
– | Submedian carina and/or hyperoccipital carina indicated by carinae or rugae ( |
80 |
80 | Mesoscutellum, posterior portion of mesoscutal midlobe with many strong longitudinal rugae ( |
|
– | Mesoscutellum especially, and usually mesoscutal midlobe, without longitudinal rugae ( |
81 |
81 | Metascutellum broad and convex, rugose centrally ( |
|
– | Metascutellum about as long as broad, concave dorsally, without rugae centrally. ( |
82 |
82 | Postmarginal vein absent or extremely short. Entire body, including antennae and legs, dark brown ( |
|
– | Postmarginal vein long, thick. At least part of legs yellowish or reddish ( |
83 |
83 | Hyperoccipital carina sharp and strongly developed ( |
|
– | Either: hyperoccipital carina irregular or weakly developed, or: frontal depression with transverse carinae that may be interrupted medially ( |
84 |
84 | Metasomal depression roughly sculptured, with anterior carinae or submedian foveae ( |
85 |
– | Metasomal depression smooth anteriorly ( |
86 |
85 | T1 midlobe with 4 longitudinal carinae | |
– | T1 midlobe with 5 longitudinal carinae | |
86 | Occiput concave dorsally, outlined by carinae because of weak connection between hyperoccipital carina and occipital carina ( |
87 |
– | Occiput convex dorsally, with no connection between occipital carina and hyperoccipital carina ( |
89 |
87 | Median carina of frontal depression reaching submedian carina ( |
|
– | Median carina of frontal depression not reaching submedian carina ( |
88 |
88 | Female: T1 horn weakly developed, not nearly reaching metascutellum ( |
|
– | Female: T1 horn strongly developed, nearly reaching metascutellum ( |
|
89 | Metascutellum medially with a branched ruga ( |
|
– | Metascutellum medially with 3 longitudinal carinae between the lateral margins ( |
These groups are provided here to indicate intuitively perceived structure within the genus, and to provide an aid for identification. They are characterized in the above key and are diagnosed here in a succinct way. Some characters are omitted from certain species group diagnoses because those characters are variable within the group or are otherwise unhelpful for that particular group’s identification. Individual species descriptions can be consulted regarding characters omitted from these diagnoses.
This group resembles the and
Characteristics:Hyperoccipital carina absent or represented by weak rugae. Occipital carina incomplete medially, with strong lateral corners. Metascutellum nearly flat. Postmarginal vein present. T1 lateral carina not expanded. Metasomal flanges absent. T2 with sublateral depressions, set off by carinae medially. Main body of T6 in females not separated from apical rim. T7 in males weakly emarginate apically.
Female. Body length 3.1–4.15 mm (n=20).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: shorter than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: present. Occipital carina: present laterally, absent medially. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate; rugose; umbilicate punctate. Major sculpture of gena posteroventrally: rugose; umbilicate punctate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: irregularly sculptured. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a ruga. Major sculpture of mesoscutellum centrally: umbilicate foveate; umbilicate punctate. Major sculpture of mesoscutellum peripherally: umbilicate foveate; umbilicate punctate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T5; reaching near apex of T5; reaching middle of T6. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: present. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: straight. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: densely foveate, longitudinal sculpture irregular. T4 metasomal flanges: absent. T5 sculpture: densely foveate, longitudinal sculpture irregular. T5 metasomal flanges: absent. T6: longer than broad; broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression flat, with oblique interrupted carinae and sometimes with an incomplete longitudinal carina; submedian carina weak. Hyperoccipital carina indicated by sharp rugae. Occipital carina connected to hyperoccipital carina by a weak longitudinal carina or irregular ruga, laterally with strong corners and medially sinuate; area between occipital and hyperoccipital carinae densely sculptured. Mesoscutellum densely foveolate. Metascutellum flat, rugose. Postmarginal vein present. Coxa not darker than rest of leg. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female: A3 not longer than pedicel. A4 slightly longer than broad, A5 broader than long. T1 with slight horn obscuring longitudinal carinae. Fore wing long enough to reach middle of T6. T6 broader than long. Male: A4 slightly longer than broad, A11 longer than broad. T1 midlobe with 3 longitudinal carinae. Fore wing long enough to exceed metasomal apex. T7 truncate or slightly emarginate apically.
Latin noun, genitive case, meaning “needle.”
[
Holotype, female: AUSTRALIA: QLD, Mount Glorious, 1977, malaise trap, OSUC 368117 (deposited in QMBA). Paratypes: AUSTRALIA: 37 females, 37 males, Australian Museum K245256, K245257, K245263, K245273, K245275 (AMSA); ANIC DB 32-020074, 32-020886, OSUC 368113, OSUC 368114, OSUC 368135, OSUC 368137, OSUC 368138, OSUC 368142, OSUC 368159 (ANIC); OSUC 227551, 227611, 462583-462584 (CNCI); OSUC 368141, 368143, 368156, 368158, QDPC 0-165753 (QDPC); OSUC 368111 (QMBA); OSUC 448965 (UQIC); OSUC 368107-368110, 368112, 368115-368116, 368118-368123, 368131-368134, 368136, 368139-368140, 368145-368150, 368152-368155, 368157, 368160-368165, 368172, 368174, 448946-448948, 448951-448953, 448961-448964 (WINC).
Female. Body length 4.25 mm (n=1).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: concave. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: with oblique carina extending towards mouth corner. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: present. Occipital carina: present laterally, absent medially. Occiput sculpture: umbilicate foveate. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate; transversely rugose. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a flattened or rounded elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: granulate. Microsculpture of mesoscutellum peripherally: granulate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: not emarginate, convex dorsally. Metascutellar setae: absent. Metascutellum sculpture: with many longitudinal rugae. Postmarginal vein: present. Fore wing apex at rest: not reaching base of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: unknown. Anterior sculpture of metasomal depression: unknown. Median propodeal carina: unknown.
T1 horn: present. Number of longitudinal carinae of T1 midlobe: 6. T1 lateral carina: straight. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: longer than broad. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression concave, transverse carinae present ventrally; dorsal separator present as a complete carina; submedian carina weakly indicated by a single carina. Oblique carina extending from bottom of frontal depression towards lower margin of eye. Head directed downward. Hyperoccipital carina indicated by rugae. Occipital carina absent medially, with protruding lateral corners. Metascutellum convex. Postmarginal vein present. Coxa not darker than rest of leg. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female: A3 longer than pedicel. A4, A5 broader than long. Mesoscutellum with granulate sculpture. T1 midlobe with strong anterior horn, 6 strong longitudinal carinae nevertheless distinct. Fore wing long enough to reach middle of T4. T6 longer than broad.
Latin noun, genitive case, meaning “a curve.”
[
Holotype, female: PAPUA NEW GUINEA: East Sepik Prov., New Guinea Isl., Sepik River, Korogo, 8.III.1964, D. H. Colless, OSUC 448564 (deposited in ANIC).
Female. Body length 3.55–8.7 mm (n=20).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: longer than broad. A5: longer than broad; as long as broad.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: with medially interrupted transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: granulate. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: umbilicate foveate. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: reaching near apex of T5; reaching middle of T6. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 5. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate foveate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression shallow, not parallel-sided, many interrupted transverse carinae present, including some above dorsal separator; submedian carina weakly indicated but complete. Hyperoccipital carina indicated by weak rugae. Occipital carina complete, weakly convex medially. Metascutellum deeply concave, emarginate apically, projecting dorsally. Postmarginal vein absent. Coxa darker than rest of leg. T1 lateral carina expanded laterally. Metasomal flanges absent. Female only: A3 much longer than pedicel. A4 longer than broad, A5 nearly as long as broad. Mesoscutellum without granulate sculpture. T1 midlobe with 4 longitudinal carinae or with raised sculpture obscuring them. T6 without metasomal flanges, main body of tergum not separated from apical rim. Fore wing long enough to reach middle of T5 or middle of T6.
[
Holotype, female, S. atricoxa: AUSTRALIA: NSW, Sydney, Elizabeth Bay, no date, ANIC DB 32-012551 (deposited in ANIC). Other material: AUSTRALIA: 67 females, OSUC 429912-429917, 429919 (ANIC); OSUC 462560 (CNCI); OSUC 281687-281688, 309399 (OSUC); QDPC 0-165655, 0-165726, 0-165754 (QDPC); OSUC 429918 (QMBA); OSUC 437012 (UQIC); OSUC 429876-429911, 429920-429921, 448968-448980 (WINC).
Female. Body length 4.5–4.9 mm (n=12).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with transverse carinae. Dorsal portion of frontal depression: with some transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: absent. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: umbilicate foveate. Extra carina ventral to occipital carina: present, medially incomplete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate; rugose. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: with shallow irregular carinae, posterodorsal corner with dense microsculpture. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: with large smooth areas. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent; punctate. Mesoscutellar rim: expanded. Mesoscutellar rim medially: with notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: reaching base of T5. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: present. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: present as slightly protruding sharp corners. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present, blade-like with rounded margins. T6: broader than long. Major sculpture of T6: longitudinally striate; umbilicate foveate. Microsculpture of T6: absent. T6 medially: with deep emargination that is V-shaped medially, separated from apical rim. T6 metasomal flanges: present, very broadly rounded, with rounded apices. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: present, posteriorly complete. S6 apex in relation to T6: exposed to dorsal view by T6 emargination. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep, all carinae except dorsal separator interrupted medially, carinae present above dorsal separator; submedian carina weakly defined or absent medially. Hyperoccipital carina absent. Occipital carina complete, medially sinuate. Mesoscutal midlobe weakly sculptured with some smooth areas near midlength. Mesoscutellar rim not expanded, with median notch. Metascutellum scoop-like, projecting dorsally. Coxa darker than rest of leg. Postmarginal vein absent. Female: A3 longer than pedicel. A4 not longer than broad, A5 broader than long. T4 with weak metasomal flanges. T5 with lobe-like metasomal flanges. T6 with broad, wavy lateral metasomal flanges. T6 broadly and deeply emarginate, roundly concave medially. S6 exposed to dorsal view, rounded apically. This species is similar to
Latin noun, genitive case, meaning “confectioneries.”
[
Holotype, female: AUSTRALIA: NSW, Pilliga, 1.II.1931, A. P. Dodd, OSUC 435957 (deposited in ANIC). Paratypes: AUSTRALIA: 11 females, OSUC 435950-435956, 435958-435960 (ANIC); OSUC 449096 (WINC).
Female. Body length 3.25–3.4 mm (n=2).
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: absent; present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: punctate. Hyperoccipital carina: present as a single carina. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: smooth; umbilicate punctate. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate punctate. Major sculpture of gena posteroventrally: umbilicate punctate; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with a median carina, otherwise weakly sculptured. Postmarginal vein: present. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: present. Anterior sculpture of metasomal depression: with median areole or pair of pits. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: straight. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: densely foveate, longitudinal sculpture irregular. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression shallow; submedian carina weakly indicated by a rounded carina. Hyperoccipital carina indicated by one complete carina that may be accompanied by an additional ruga. Occipital carina incomplete, lateral portions approaching hyperoccipital carina. Occiput umbilicate punctate. Metascutellum broad and short, concave, with a distinct median carina. Postmarginal vein present. Coxa and rest of leg bicolored. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female: A3 longer than pedicel. A4, A5 broader than long. Mesoscutellum with small umbilicate foveae. T1 midlobe without anterior horn, with 4 longitudinal carinae. Fore wing long enough to reach beyond metasomal apex. T6 broader than long.
Latin noun, genitive case, meaning “bicolored foot.”
[
Holotype, female: NEW CALEDONIA: Sud Prov., Riviere Bleue Territorial Park,
Female. Body length 2.4–3.75 mm (n=6).
Radicle color and shade: darker than scape. Pedicel color: at least partially darker than scape. A3: shorter than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: present as a single carina; indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: omicron-shaped, with sharp corners where median portion meets lateral portions. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate; rugose. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent; granulate. Microsculpture of gena posteroventrally: absent; granulate.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: punctate. Microsculpture of mesoscutellum peripherally: punctate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: reaching near apex of T5; reaching middle of T6. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, but parallel for a short distance anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as strong posterior corners. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: strongly convex, tapering and sloping down to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Mesoscutum and mesoscutellum black. Frontal depression deep, transverse carinae absent; submedian carina strong. Hyperoccipital carina indicated by strong rugae. Occipital carina complete, convex medially. Metascutellum concave, broad and short, projecting dorsally. Coxa not darker than rest of leg. T1 lateral carina expanded laterally. Female only: A3 shorter than pedicel. A4, A5 much broader than long. T1 midlobe with 4 longitudinal carinae. T6 without metasomal flanges. Fore wing long enough to reach or exceed metasomal apex.
Latin noun in apposition, meaning “the short.”
[
Holotype, female: AUSTRALIA: ACT, Canberra, 4.II.1964, I. D. Galloway, OSUC 441375 (deposited in ANIC). Paratypes: AUSTRALIA: 6 females, OSUC 437878, 441376-441378 (ANIC); OSUC 441374, 451319 (WINC).
This species can be recognized by its extremely short A3, A4, and A5. Additionally, the strongly defined frontal depression and submedian carina are unusual for a short-bodied species.
Female. Body length 3.5–4.05 mm (n=11).
Radicle color and shade: darker than scape; same as scape, both dark brown. Pedicel color: same as scape. A3: longer than pedicel. A4: longer than broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: granulate. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: uniformly rounded dorsally. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, medially incomplete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate; rugose. Major sculpture of gena posteroventrally: umbilicate foveate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: densely covered with setiferous puncta. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: with large smooth areas. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: absent; umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: punctate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: reaching base of T5. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, but parallel for a short distance anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: with median areole or pair of pits. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: straight. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate; longitudinally striate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: truncate.
Latin noun, genitive case, meaning “chain.”
[
Holotype, female: AUSTRALIA: WA, Mount Cooke, 28.I-17.II.1991, malaise trap, M. S. Harvey & J. M. Waldock, OSUC 448987 (deposited in WAMP). Paratypes: AUSTRALIA: 10 females, 4 males, OSUC 439567-439569, 448986, 448988-448997 (WINC).
Female. Body length 2.8–3.1 mm (n=4).
Radicle color and shade: same as scape, both yellowish or reddish. Pedicel color: same as scape. A3: shorter than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: granulate. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: broadly angular, with rounded median peak. Occiput sculpture: umbilicate foveate. Extra carina ventral to occipital carina: present, medially incomplete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: with shallow irregular carinae, posterodorsal corner with dense microsculpture. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: longitudinally rugose. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: reaching near apex of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 5. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as slightly protruding sharp corners. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: with medially truncate emargination, sloping down to apical rim. T6 metasomal flanges: present as spine-like structures posterolaterally. T6 raised peripheral rim: absent. S4 sculpture: densely setose, setal pits between very weak longitudinal rugae. S5 sculpture: densely setose, setal pits between very weak longitudinal rugae. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: exposed to dorsal view by T6 emargination. S6 apex: truncate.
Both sexes: Frontal depression shallow, without carinae ventrally, dorsal separator interrupted medially, weakly indicated carinae present above dorsal separator; submedian carina absent. Hyperoccipital carina indicated by vague rugae. Occipital carina complete, medially sinuate. Mesoscutellar rim not expanded, without median notch. Metascutellum small, weakly emarginate apically, projecting dorsally. Coxa same color as rest of leg. Postmarginal vein absent. Female: A4, A5 broader than long. T4, T5 without distinct metasomal flanges, but T5 with sharp posterior corners. T6 abruptly narrower than T5, without expanded lateral margins but with narrow and sharp elongate posterior metasomal flanges. Main surface of T6 strongly emarginate medially, sloping down to apical rim. S6 exposed to dorsal view, rounded apically. It differs from
Latin noun, genitive case, meaning “tails.”
[
Holotype, female: AUSTRALIA: QLD, open forest, 3.4km S Port Douglas, 17.XI.1979, E. C. Dahms, J. B. Woolley & J. LaSalle, OSUC 437874 (deposited in QMBA). Paratypes: AUSTRALIA: 3 females, OSUC 437873, 437876 (ANIC); OSUC 437875 (WINC).
Female. Body length 3.25 mm (n=1).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: longer than broad. A5: longer than broad.
Ventral clypeal margin: concave. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: with transverse ledge, face sharply receding below it. Transverse curved rugae extending from frontal depression to eye: present. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: present as multiple regular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate punctate; absent. Major sculpture of gena posteroventrally: umbilicate punctate; absent. Microsculpture of gena anteroventrally: absent; granulate. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: with narrow curved smooth elevations. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; transversely rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent. Major sculpture of mesoscutellum centrally: umbilicate punctate. Major sculpture of mesoscutellum peripherally: umbilicate punctate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with a median carina, otherwise weakly sculptured. Postmarginal vein: present. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: present. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: straight. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression nearly flat, surrounded by many transverse curved rugae; submedian carina indicated by a set of rugae. Hyperoccipital carina indicated by many transverse carinae. Occipital carina incomplete, lateral portions approaching hyperoccipital carina. Occiput irregularly sculptured. Metascutellum broad and very short, concave, with a distinct median carina. Postmarginal vein present. Coxa and rest of leg bicolored. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female: A3 longer than pedicel. A4, A5 longer than broad. Mesoscutellum with setiferous puncta. T1 midlobe with very weak anterior horn, no complete longitudinal carinae. Fore wing long enough to reach beyond metasomal apex. T6 broader than long. Male: Fore wing long enough to exceed metasomal apex. T7 tiny, truncate.
Latin noun, genitive case, meaning “rings.”
[
Holotype, female: NEW CALEDONIA: Nord Prov., 5km WSW Pouebo, Mount Mandjelia,
Female. Body length 3.3–4.75 mm (n=20).
Radicle color and shade: darker than scape. Pedicel color: at least partially darker than scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: uniformly convex. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: broadly angular, with rounded median peak. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent; granulate.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a ruga. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: punctate. Microsculpture of mesoscutellum peripherally: punctate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: reaching base of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: present as slightly protruding sharp corners. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as strongly protruding acuminate flanges. T6: broader than long. Major sculpture of T6: longitudinally striate; umbilicate foveate. Microsculpture of T6: absent. T6 medially: with a median projection set off by an abrupt narrowing posterior to tiny metasomal flanges, not separated from apical rim. T6 metasomal flanges: present subapically. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep and broad, carinae not present above dorsal separator; submedian carina distinct, sharp. Hyperoccipital carina indicated by vague rugae. Occipital carina complete, medially weakly convex. Metascutellum concave dorsally, weakly emarginate apically, projecting dorsally. Coxa color variable. Postmarginal vein absent. Female: A3 longer than pedicel. A4, A5 broader than long. T1 midlobe with 5 longitudinal carinae or obscured by raised sculpture. T4 with or without small metasomal flanges, T5 with metasomal flanges, not otherwise abruptly narrower than preceding tergum. Main body of T6 raised above apical rim, but sloping down to it medially, not abruptly separated from it. S6 not exposed to dorsal view.
Latin noun, genitive case, meaning “slope.”
[
Holotype, female: AUSTRALIA: SA, 79km NNW Renmark, Calperum Station, Bookmark Biosphere Reserve,
Female. Body length 3.7–4.15 mm (n=7).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate; longitudinally rugose. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a ruga. Major sculpture of mesoscutellum centrally: absent; umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent; punctate. Microsculpture of mesoscutellum peripherally: punctate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: reaching base of T5. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: present as slightly protruding sharp corners. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as strongly protruding acuminate flanges. T6: broader than long. Major sculpture of T6: umbilicate punctate; longitudinally striate. Microsculpture of T6: absent. T6 medially: with a median projection set off by an abrupt narrowing posterior to tiny metasomal flanges, not separated from apical rim. T6 metasomal flanges: present subapically. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep and broad, carinae not present above dorsal separator; submedian carina distinct, sharp. Hyperoccipital carina indicated by strong rugae. Occipital carina complete, medially weakly convex. Metascutellum concave dorsally, weakly emarginate apically, projecting dorsally. Coxa darker than rest of leg. Postmarginal vein absent. Female: A3 longer than pedicel. A4, A5 broader than long. T4, T5 with sharp metasomal flanges, not otherwise abruptly narrower than preceding tergum. Main body of T6 raised above apical rim, but sloping down to it medially, not abruptly separated from it; T6 apex forming a trident-like shape. S6 not exposed to dorsal view.
Latin noun, genitive case, meaning “bronze shield.”
[
Holotype, female: AUSTRALIA: NSW, Fowlers Gap Research Station,
Female. Body length 3.55–4.15 mm (n=20).
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: shorter than pedicel; as long as pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: uniformly rounded dorsally; weakly arched dorsally, with rounded lateral corners. Occiput sculpture: transversely rugose. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate punctate; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; irregularly rugose. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a ruga. Major sculpture of mesoscutellum centrally: absent; umbilicate punctate. Major sculpture of mesoscutellum peripherally: umbilicate foveate; umbilicate punctate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, but parallel for a short distance anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T6. Coxae color brightness: same color as femora.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 5; obscured by other raised sculpture. T1 lateral carina: straight. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate; longitudinally striate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Body, including antennae and legs, entirely dark brown. Frontal depression small and nearly flat, transverse carinae absent; submedian carina indicated by weak rugae. Hyperoccipital carina indicated by one or more sharp rugae. Occipital carina incomplete, weakly convex medially; occiput with many transverse rugae. Metascutellum deeply concave, emarginate apically, projecting dorsally. Postmarginal vein absent or extremely short. T1 lateral carina not expanded laterally. Female: A3 longer than pedicel. A4, A5 broader than long. T1 midlobe with 5 longitudinal carinae that may be obscured by a smooth raised area. Fore wing long enough to reach middle of T6 or beyond metasomal apex. T6 broader than long. Male: A4, A11 about as long as broad. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach beyond T7. T7 truncate or weakly concave.
[
Holotype, female, S. concoloripes: AUSTRALIA: NSW, no date, ANIC DB 32-012463 (deposited in ANIC). Other material: AUSTRALIA: 76 females, 99 males, OSUC 438799-438808, 438811-438814, 439384-439387, 439389-439395, 439409-439442, 439445-439481, 439483-439485, 445326 (ANIC); OSUC 462587-462589 (CNCI); OSUC 438810, 439388, 439396-439408, 439443-439444, 439482, 439486-439539 (WINC).
Our concept of this species suggests that it is variable in female metasomal length. The holotype has a shorter metasoma, in which T6 is broader than long, but some specimens have a longer metasoma, where T6 is longer than broad. Specimens of both forms have been found from the same collection events, and some specimens appear to be intermediate between the two states. Several other Australian species are entirely dark brown, but these species differ in many other respects, and do not appear to form a monophyletic group.
Female. Body length 3.35–4.85 mm (n=20).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: longer than broad; as long as broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: with medially interrupted transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: granulate. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate; rugose. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: absent; granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: reaching middle of T5. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate foveate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
A3: longer than pedicel. A5 tyloid shape: narrow, linear. A6: broader than long. A11: longer than broad. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Fore wing apex at rest: reaching middle of T5. T1 midlobe longitudinal carinae: 4. T3 metasomal flanges: absent. T4 metasomal flanges: absent. T5 metasomal flanges: absent. T6 metasomal flanges: absent. T7: broadly and deeply emarginate, with rounded posterolateral margins.
Both sexes: Frontal depression deep and broad but nearly parallel-sided, many interrupted transverse carinae present, including some above dorsal separator; submedian carina weakly indicated or absent medially. Hyperoccipital carina indicated by weak rugae. Occipital carina complete, sinuate medially. Metascutellum deeply concave, weakly emarginate apically, projecting dorsally. Postmarginal vein absent. Coxa darker than rest of leg. T1 lateral carina expanded laterally. Metasomal flanges absent. Female: A3 much longer than pedicel. A4 about as long as broad. Mesoscutellum without granulate sculpture. T1 midlobe with 4 longitudinal carinae. T6 without metasomal flanges, main body of tergum raised but not always clearly separated from apical rim. Fore wing long enough to reach middle of T4 or middle of T5. Male: A4 as broad or broader than long, A11 longer than broad. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach middle of T5. T7 medially emarginate, with rounded lobes laterally.
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Holotype, female: AUSTRALIA: QLD, 5729 ‘amphitheatre’ yards, open forest, Expedition National Park,
Female. Body length 2.95–3.95 mm (n=20).
Radicle color and shade: same as scape, both yellowish or reddish. Pedicel color: same as scape. A3: as long as pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: uniformly convex. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: with shallow irregular carinae, posterodorsal corner with dense microsculpture. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: reaching middle of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: present as slightly protruding sharp corners. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as strong posterior corners. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: with deep emargination that is V-shaped medially, separated from apical rim. T6 metasomal flanges: present as slightly expanded lateral rims, rounded posteriorly. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep, with interrupted carinae ventrally, dorsal separator complete medially, weakly indicated carinae present above dorsal separator; submedian carina distinct but not strongly projecting. Hyperoccipital carina indicated by vague rugae. Occipital carina complete, medially weakly convex. Mesoscutellar rim not expanded, without median notch. Metascutellum small, weakly emarginate apically, projecting dorsally. Coxa same color as rest of leg. Postmarginal vein absent. Female: A3 longer than pedicel. A4, A5 broader than long. T4, T5 without distinct metasomal flanges. T6 abruptly narrower than T5, without expanded lateral margins. Main surface of T6 strongly emarginate medially, with sharp posterior corners, sharply separated from apical rim. S6 exposed to dorsal view, rounded apically.
Latin noun, genitive case, meaning “contusion.”
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Collected on
Holotype, female: AUSTRALIA: QLD, Batavia Downs,
Female. Body length 4.3–4.85 mm (n=7).
Radicle color and shade: darker than scape. Pedicel color: at least partially darker than scape. A3: longer than pedicel. A4: longer than broad. A5: longer than broad.
Ventral clypeal margin: concave. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate punctate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: mostly granulate, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally and ventrally. Metascutellum shape: not emarginate, convex dorsally. Metascutellar setae: absent. Metascutellum sculpture: with many longitudinal rugae. Postmarginal vein: present. Fore wing apex at rest: reaching base of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: present. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: straight. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: longer than broad. Major sculpture of T6: umbilicate punctate; longitudinally striate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression shallow, transverse carinae absent; submedian carina sharp. Hyperoccipital carina indicated by rugae. Occipital carina absent medially, lateral corners not protruding. Postmarginal vein present. Coxa not darker than rest of leg. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female: A4 longer than broad, A5 about as long as broad. Mesoscutellum without granulate sculpture. Metascutellum broad and short, slightly convex, irregularly rugose. Fore wing long enough to reach base of T5. T6 longer than broad.
Latin noun, genitive case, meaning “corrugation.”
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Holotype, female: AUSTRALIA: QLD, GS2, Mount Edith,
Female. Body length 2.9–3.55 mm (n=19).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: longer than broad. A5: longer than broad.
Ventral clypeal margin: concave. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: granulate. Hyperoccipital carina: present as a single carina. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: transversely rugose. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate punctate. Major sculpture of gena posteroventrally: umbilicate punctate. Microsculpture of gena anteroventrally: granulate. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: irregularly sculptured. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate; umbilicate punctate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate; irregularly rugose. Major sculpture of mesoscutellum peripherally: umbilicate foveate; irregularly rugose. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally and ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T6. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: meeting near propodeal midlength. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: with series of longitudinal carinae extending to lateral propodeal carinae. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 5. T1 lateral carina: straight. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: granulate. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression nearly parallel-sided, transverse carinae complete but weakly indicated; submedian carina sharp or weak and irregular. Hyperoccipital carina sharp and strong. Occipital carina incomplete, lateral portions almost reaching hyperoccipital carina; occiput with many weak rugae and a row of strong setiferous puncta. Metascutellum with a concave postero-medial area, laterally with broad longitudinally striate area, weakly emarginate, projecting dorsally. Postmarginal vein present. Coxa not darker than rest of leg. T1 lateral carina not expanded laterally. Female: A3 longer than pedicel. A4 longer than broad, A5 as long or longer than broad. Mesoscutellum densely and coarsely sculptured. T1 midlobe with 5 longitudinal carinae. Fore wing long enough to reach or exceed T6. Male: All flagellomeres longer than broad. T1 midlobe with 3 or 4 longitudinal carinae, hardly raised above sidelobes. Mesoscutellum with strong irregular rugae and some granulate sculpture. Fore wing long enough to reach beyond T7. T7 truncate, steeply sloping.
Latin noun, genitive case, meaning “saffron.”
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Holotype, female: AUSTRALIA: QLD, via Canungra, forest edge, O’Reilley’s Guest House, 3.II-2.III.1980, malaise trap, OSUC 442318 (deposited in QMBA). Paratypes: AUSTRALIA: 18 females, 23 males, OSUC 442321, 442340 (ANIC); OSUC 451285 (BMNH); OSUC 227541, 227610, 227628, 462567-462570 (CNCI); OSUC 442324, 442329, QDPC 0-165788 (QDPC); OSUC 442319 (QPIM); OSUC 442311-442317, 442320, 442323, 442325-442328, 442330-442339, 451284, 451286-451288 (WINC).
Female. Body length 3.2–3.95 mm (n=14).
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: granulate. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, medially incomplete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: with large smooth areas. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a narrow carina; present as a flattened or rounded elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate; irregularly rugose. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: reaching base of T5. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 6. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: present. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: present as slightly protruding sharp corners. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as slightly protruding sharp corners. T6: broader than long. Major sculpture of T6: longitudinally striate; umbilicate foveate. Microsculpture of T6: absent. T6 medially: with a median projection set off by an abrupt narrowing posterior to tiny metasomal flanges, not separated from apical rim. T6 metasomal flanges: present subapically. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: truncate.
Both sexes: Frontal depression deep, carinae not present above dorsal separator; submedian carina distinct, sharp. Hyperoccipital carina indicated by rugae. Occipital carina complete, medially weakly convex. Metascutellum concave dorsally, weakly emarginate apically, projecting dorsally. Coxa darker than rest of leg. Postmarginal vein absent. Female: A3 longer than pedicel. A4, A5 broader than long. T4 with sharp corners, T5 with small sharp metasomal flanges, not otherwise abruptly narrower than preceding tergum. Main body of T6 not raised above apical rim, not abruptly separated from it; T6 with small and sharp subapical metasomal flanges. S6 not exposed to dorsal view.
Latin noun, genitive case, meaning “point.”
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Holotype, female: AUSTRALIA: QLD, Mount Moffatt Section, 3km SE park headquarters, Carnarvon National Park,
Females of
Female. Body length 2.75–4 mm (n=20).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: longer than broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: with medially interrupted transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: absent. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: umbilicate foveate. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate; umbilicate punctate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent; punctate. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: deeply emarginate, with the resulting pair of posterior processes subtriangular and directed dorsally. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: reaching middle of T6. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate foveate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression shallow, transverse carinae absent or broadly interrupted ventrally, with sometimes a few faint carinae dorsally; submedian carina absent medially. Hyperoccipital carina indicated by weak rugae. Occipital carina complete, weakly convex or omicron-shaped medially. Metascutellum deeply concave, truncate or weakly emarginate apically, projecting dorsally. Postmarginal vein absent. Coxa darker than rest of leg. T1 lateral carina expanded laterally. Metasomal flanges absent. Female: A3 longer than pedicel. A4 longer than broad, A5 nearly as long as broad. Mesoscutellum without granulate sculpture. T1 midlobe with 4 longitudinal carinae or with raised sculpture obscuring them. T6 without metasomal flanges, main body of tergum sloping down to apical rim. Fore wing long enough to reach middle of T5 or middle of T6.
Latin noun, genitive case, meaning “density, abundance.” Refers to the abundance of mesoscutal midlobe foveae.
[
Inhabits brigalow: [
Holotype, female: AUSTRALIA: QLD, via Taroom, FIT 057, Boggomoss No. 30,
Female. Body length 4–4.15 mm (n=2).
Radicle color and shade: same as scape, both yellowish or reddish. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: concave. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: granulate. Hyperoccipital carina: present as a single carina. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: transversely rugose; umbilicate punctate. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent; punctate.
Lateral pronotal area sculpture: irregularly foveate, with smooth area dorsally. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; irregularly rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate; umbilicate punctate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally and ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T5; reaching near apex of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: narrowly separated, angled anteriorly to become parallel. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: with median areole or pair of pits. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: strongly convex, tapering and sloping down to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep and broad, transverse carinae interrupted or absent; submedian carina present. Hyperoccipital carina indicated by strong and sharp ruga. Occipital carina incomplete medially. Metascutellum deeply concave, not emarginate apically, projecting dorsally. Postmarginal vein present, short. Coxa not darker than rest of leg. T1 lateral carina expanded laterally. Metasomal flanges absent. Female: A3 longer than pedicel. A4, A5 much broader than long. Mesoscutellum without granulate sculpture. T1 midlobe with 4 longitudinal carinae or with these obscured by raised sculpture. T6 without metasomal flanges, not concave apically. Fore wing long enough to reach middle or apex of T5. Male: Flagellomeres between A3 and A11 broader than long. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach to or beyond T7. T7 weakly emarginate, without projections.
Latin noun, genitive case, meaning “concealment.”
[
Holotype, female: AUSTRALIA: QLD, via Taroom, FIT 057, Boggomoss No. 30,
Female. Body length 3.2–3.5 mm (n=11).
Radicle color and shade: darker than scape. Pedicel color: at least partially darker than scape. A3: shorter than pedicel. A4: longer than broad. A5: longer than broad; broader than long.
Ventral clypeal margin: concave. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: present. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: with some transverse carinae. Submedian carina: absent. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: present as a single carina. Carina connecting occipital carina to hyperoccipital carina: present. Occipital carina: present laterally, absent medially. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: rugose; umbilicate punctate. Major sculpture of gena posteroventrally: rugose; umbilicate punctate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a ruga. Major sculpture of mesoscutellum centrally: umbilicate punctate. Major sculpture of mesoscutellum peripherally: umbilicate punctate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally and ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 5; 6. T1 lateral carina: straight. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression flat, with oblique interrupted carinae and a strong longitudinal carina that extends throughout its length; submedian carina indicated by weak rugae. Hyperoccipital carina indicated by rugae. Occipital carina connected to hyperoccipital carina by a weak longitudinal carina, laterally with strong corners and arched; area between occipital and hyperoccipital carinae densely sculptured and having many short setae. Mesoscutellum smooth, with scattered setiferous puncta. Metascutellum tiny, concave. Postmarginal vein present. Coxa not darker than rest of leg. Metasomal depression smooth. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female: A3 not longer than pedicel. A4 longer than broad, A5 broader than long. T1 midlobe with 5 longitudinal carinae. Fore wing long enough to reach middle of T6 or beyond metasomal apex. T6 broader than long. Male: A4 about as broad as long, A11 longer than broad. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to exceed metasomal apex. T7 tiny, truncate apically.
Latin noun, genitive case, meaning “a division.”
[
Holotype, female: AUSTRALIA: QLD, summit TV station, rainforest, Bellenden Ker Range, 1560m, 17.X–5.XI.1981, yellow pan trap, OSUC 368204 (deposited in QMBA). Paratypes: AUSTRALIA: 10 females, 2 males, OSUC 368195, 368197-368199, 368205-368206 (ANIC); OSUC 462581 (CNCI); OSUC 368196, 368200-368203 (WINC).
Female. Body length 2.3–3.05 mm (n=15).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: shorter than pedicel. A4: longer than broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: with some transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: granulate. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: omicron-shaped, with sharp corners where median portion meets lateral portions. Occiput sculpture: umbilicate foveate. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; absent. Microsculpture of gena anteroventrally: granulate. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: with shallow irregular carinae, posterodorsal corner with dense microsculpture. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; irregularly rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent; punctate. Microsculpture of mesoscutellum peripherally: absent; punctate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: reaching middle of T5. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: with medially truncate emargination, sloping down to apical rim. T6 metasomal flanges: present as tiny apical sharp projections. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep, carinae present above dorsal separator; submedian carina weak or absent medially. Hyperoccipital carina indicated by rugae. Occipital carina complete, medially sinuate. Metascutellum concave dorsally, weakly emarginate apically, projecting dorsally. Coxa darker than rest of leg. Postmarginal vein absent. Female: A3 longer than pedicel. A4, A5 broader than long. Fore wing long enough to reach middle of T5 or to metasomal apex. T4, T5 without metasomal flanges. T6 abruptly narrower than T5. Main body of T6 raised above apical rim, medially sloping down to it. T6 with broadly rounded apical lobes. S6 not exposed to dorsal view.
Latin noun, genitive case, meaning “smallness.”
[
Holotype, female: AUSTRALIA: QLD, 7km S Batavia Downs,
Female. Body length 2.85–4 mm (n=20).
Radicle color and shade: same as scape, both yellowish or reddish. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: with transverse ledge, face sharply receding below it. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: present. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present; present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: present as a single carina. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: umbilicate punctate. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate; umbilicate punctate; absent. Major sculpture of gena posteroventrally: umbilicate punctate; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: smooth anteriorly, densely setose posteriorly. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate; umbilicate punctate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate; umbilicate punctate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, but parallel for a short distance anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T6. Coxae color brightness: same color as femora.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: straight. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: present. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression shallow, transverse carinae absent or interrupted; submedian carina indicated by a weak ruga, flat or only weakly rounded dorsally. Hyperoccipital carina sharp and strong. Occipital carina incomplete, lateral portions nearly reaching hyperoccipital carina. Occiput almost entirely smooth, with a row of setiferous puncta. Mesoscutum without median carina. Metascutellum broad and concave, truncate or slightly convex apically, projecting dorsally. Postmarginal vein present. Coxa not darker than rest of leg. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female: A3 much longer than pedicel. A4 longer than broad, A5 broader than long. Mesoscutellum weakly sculptured, with only setiferous puncta or small foveae. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach T6. T6 broader than long. Male: A4 broader than long, A11 longer than broad; A5 tyloid narrow. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach far beyond metasomal apex. T7 tiny, truncate.
[
Holotype, female, S. flavipes: AUSTRALIA: QLD, Mackay, II-1900, R. E. Turner, B.M. TYPE HYM. 9.510 (deposited in BMNH). Other material: AUSTRALIA: 106 females, 64 males, Australian Museum K245259, K245269 (AMSA); OSUC 368242-368244, 368254, 438857-438858, 438874-438878, 438886, 438901-438906, 438914, 438916-438917, 438920, 438923, 445348, 451292, 451294, 451296-451297 (ANIC); OSUC 227566, 227606, 227609, 227615, 227622, 227629, 462592-462597 (CNCI); OSUC 376737 (MCZC); OSUC 165732, 221760-221762, 221764, 451273, 62719, 62728, 62739-62741, 63076, 63086, 63095, 63097, 63102, 63104-63105, 63190, 63228, 72257-72258 (OSUC); OSUC 368253, 438891, 438910, 451269, QDPC 0-165632, QDPC 0-165635, QDPC 0-165638, QDPC 0-165639, QDPC 0-165642, QDPC 0-165644, QDPC 0-165657, QDPC 0-165665, QDPC 0-165679, QDPC 0-165693, QDPC 0-165706, QDPC 0-165720, QDPC 0-165721, QDPC 0-165730, QDPC 0-165734, QDPC 0-165736, QDPC 0-165747, QDPC 0-165758, QDPC 0-165770, QDPC 0-165772, QDPC 0-165778, QDPC 0-165787 (QDPC); OSUC 148371, 368240-368241, 368256, 438879-438880, 438907-438909, 438915 (QMBA); OSUC 448596 (QPIM); UCRC ENT 100244 (UCRC); OSUC 368238-368239, 368245-368252, 368255, 438855-438856, 438859-438873, 438881-438885, 438887-438890, 438892-438900, 438911-438913, 438918-438919, 438921-438922, 438924-438929, 445347, 451268, 451270-451272, 451293, 451295, 451298 (WINC).
Our concept of this species allows for considerable variation in sculpture and setation. The sculptural variation includes differences in the height and distinction of the submedian carina, strength of a longitudinal ridge along the median axis of the frontal depression, sculpture of the metasomal depression, and strength of the longitudinal elevations on the posteromedian part of the mesoscutal midlobe. Density of setation also may vary on the occiput and mesoscutellum. This variation does correlate with collection locality. It is possible that our concept of
Female. Body length 2.55–3.25 mm (n=17).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: with transverse ledge, face sharply receding below it. Transverse curved rugae extending from frontal depression to eye: present. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: present as a single carina. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: umbilicate punctate. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate punctate; absent. Major sculpture of gena posteroventrally: umbilicate punctate; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: smooth anteriorly, densely setose posteriorly. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate punctate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: obliquely rugose; umbilicate punctate. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent. Major sculpture of mesoscutellum centrally: umbilicate punctate. Major sculpture of mesoscutellum peripherally: umbilicate punctate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: narrowly separated, angled anteriorly to become parallel. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: with median areole or pair of pits. Median propodeal carina: present.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: straight. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: densely foveate, longitudinal sculpture irregular. T4 metasomal flanges: absent. T5 sculpture: densely foveate, longitudinal sculpture irregular. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression shallow; submedian carina indicated by a set of weak rugae, flat or only weakly rounded dorsally. Hyperoccipital carina sharp and strong. Occipital carina incomplete, lateral portions nearly reaching hyperoccipital carina. Occiput almost entirely smooth, with a row of setiferous puncta. Mesoscutum without median carina. Metascutellum broad and concave, truncate or slightly convex apically, projecting dorsally. Postmarginal vein present. Coxa not darker than rest of leg. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female: A4, A5 broader than long. Frontal depression with very weak oblique, interrupted carinae in lower portion. Mesoscutellum weakly sculptured, with only setiferous puncta or small foveae. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach far beyond metasomal apex. T6 broader than long. Male: A4 broader than long, A11 longer than broad. Frontal depression with long, strongly oblique carinae that are continuous with rugae that extend outside frontal depression to approach eye margin. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach far beyond metasomal apex. T7 tiny, truncate. The head of
Latin noun, genitive case, meaning “waves.”
[
Holotype, female: AUSTRALIA: QLD, rainforest, Charmillin Creek,
Female. Body length 4.6 mm (n=1).
Radicle color and shade: same as scape, both yellowish or reddish. Pedicel color: same as scape. A3: longer than pedicel. A4: as long as broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: absent. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate; rugose. Major sculpture of gena posteroventrally: umbilicate foveate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: with a series of arched carinae, posterodorsal corner with weak longitudinal rugae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: transversely rugose; umbilicate punctate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent. Major sculpture of mesoscutellum centrally: absent. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent; punctate. Microsculpture of mesoscutellum peripherally: punctate. Mesoscutellar rim: expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: reaching base of T5. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: present. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: present as slightly protruding sharp corners. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as strongly protruding acuminate flanges. T6: broader than long. Major sculpture of T6: umbilicate punctate; longitudinally striate. Microsculpture of T6: absent. T6 medially: with medially truncate emargination, sloping down to apical rim. T6 metasomal flanges: present, broad and elongate, with slight lateral incisions near midlength. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: present, posteriorly complete. S6 apex in relation to T6: exposed to dorsal view by T6 emargination. S6 apex: truncate.
Both sexes: Frontal depression deep, all carinae complete medially, carinae present above dorsal separator; submedian carina very strong. Hyperoccipital carina absent. Occipital carina complete, medially weakly convex. Mesoscutal midlobe with strong transverse carinae. Mesoscutellar rim expanded and strongly sculptured, without median notch. Metascutellum pentagonal and shovel-like, projecting dorsally. Coxa darker than rest of leg. Postmarginal vein absent. Female: A3 longer than pedicel. A4, A5 broader than long. T4 with small sharp metasomal flanges. T5, T6 with elongate, flat and sharp metasomal flanges. T6 broadly and deeply emarginate, truncate or roundly concave medially. S6 exposed to dorsal view, truncate apically. Male: A3, A11 longer than broad; A4 slightly broader than long. Mesoscutellum and mesoscutal midlobe posteriorly with small foveae, mesoscutellum without smooth area medially. T5, T6 with long sharp metasomal flanges. T7 with flat sharp metasomal flanges, deeply emarginate medially, posterior margin deeply M-shaped.
Latin noun, genitive case, meaning “leaves.”
[
On flower of
Holotype, female: AUSTRALIA: SA, nr. Ikara (Wilpena Pound) Valley, Edeowie Homestead, 29.X.1972, H. E. Evans, OSUC 376702 (deposited in MCZC). Paratype: AUSTRALIA: 1 male, OSUC 376701 (MCZC).
Female. Body length 3.75–5.2 mm (n=17).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: longer than broad. A5: as long as broad.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: with some transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, medially incomplete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate; rugose. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: with shallow irregular carinae, posterodorsal corner with dense microsculpture. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: with large smooth areas. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent; present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: absent; umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch; with notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: not reaching base of T5. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: present as slightly protruding sharp corners. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present, rounded and lobe-like. T6: broader than long. Major sculpture of T6: umbilicate punctate; longitudinally striate. Microsculpture of T6: absent. T6 medially: with deep emargination that is V-shaped medially, separated from apical rim. T6 metasomal flanges: present as slightly expanded lateral rims, rounded posteriorly. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: present, posteriorly complete. S6 apex in relation to T6: exposed to dorsal view by T6 emargination. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep, all carinae complete medially, carinae present above dorsal separator; submedian carina weakly defined or absent medially. Hyperoccipital carina absent. Occipital carina complete, medially weakly convex or sinuate. Mesoscutellar rim not expanded, with or without median notch. Metascutellum scoop-like, projecting dorsally. Coxa darker than rest of leg. Postmarginal vein absent. Female: A3 longer than pedicel. A4 longer than broad. A5 as long or longer than broad. T4 with very weak metasomal flanges. T5 with lobe-like metasomal flanges. T6 with expanded lateral margins. T6 deeply emarginate, angularly emarginate medially. S6 exposed to dorsal view, rounded apically. Male: A3, A4, A11 longer than broad. Mesoscutellum and mesoscutal midlobe posteriorly with small and densely set foveae, mesoscutellum without smooth area medially. T5 with weak and indistinct metasomal flanges, T6 with sharp metasomal flanges. T7 with long postero-lateral lobes, deeply emarginate medially, posterior margin M-shaped.
Latin noun, genitive case, meaning “rope.”
[
On blossom of
Holotype, female: AUSTRALIA: SA, 12km NE Morgan,
Female. Unknown.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: not steep, forming less than a right angle. Median mesoscutal carina: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Coxae color brightness: same color as femora. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Major sculpture of mesoscutellum centrally: umbilicate foveate; absent. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Fore wing apex at rest: unknown.
T1 midlobe longitudinal carinae: unknown. T3 metasomal flanges: absent. T4 metasomal flanges: absent. T5 metasomal flanges: absent. T6 metasomal flanges: absent. T7: with a pair of sharply defined spine-like posterolateral projections.
Male: Median carina of mesoscutum absent. Mesoscutellum without granulate sculpture. T1 lateral carina not expanded. T7 with narrow and elongate posterior spines.
[
Holotype, male: H. grandis: AUSTRALIA: QLD, nr. Cairns, foliage / jungle, Gordonvale (Nelson), 8.V.1913, sweeping, A. P. Dodd, SAMA DB 32-001588 (deposited in SAMA).
Female. Body length 3.55–3.7 mm (n=2).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: shorter than pedicel; as long as pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae; smooth. Dorsal portion of frontal depression: with medially interrupted transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: transversely rugose. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: with large smooth areas. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; irregularly rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a narrow carina. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: deeply emarginate, with the resulting pair of posterior processes subtriangular and directed dorsally. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: reaching middle of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: present. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 5. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: present as slightly protruding sharp corners. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as slightly protruding sharp corners. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: with a median projection set off by an abrupt narrowing posterior to tiny metasomal flanges, not separated from apical rim. T6 metasomal flanges: present subapically. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep; submedian carina distinct, sharp. Hyperoccipital carina indicated by rugae. Occipital carina complete, medially convex. Metascutellum concave dorsally, weakly emarginate apically, projecting dorsally. Coxa same color as most of femur, leg otherwise lighter in color. Postmarginal vein absent. Female: A3 not longer than pedicel. A4, A5 broader than long. Fore wing long enough to reach middle of T5. T1 midlobe with 5 longitudinal carinae. T4 with sharp corners, T5 with small sharp metasomal flanges, not otherwise abruptly narrower than preceding tergum. Main body of T6 not raised above apical rim, not abruptly separated from it; T6 with small and sharp subapical metasomal flanges. S6 not exposed to dorsal view. Male: A4, A11 broader than long. Fore wing long enough to reach middle of T6. T7 deeply emarginate, with broad posterolateral lobes.
Latin 4th declension noun, genitive case, meaning “a step.”
[
Holotype, female: AUSTRALIA: WA, 40km SE Armadale, Banksia-jarrah forest, Mount Cooke, 7.XII-22.XII.1990, malaise trap, A. D. Austin, OSUC 456176 (deposited in WAMP). Paratypes: AUSTRALIA: 1 female, 2 males, OSUC 456177-456179 (WINC).
Female. Body length 3.2–3.45 mm (n=5).
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: shorter than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: transversely rugose. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; umbilicate punctate; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: with large smooth areas. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: absent. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: with slight, indistinct sculpture dorsally, smooth ventrally. Metascutellum shape: deeply emarginate, with the resulting pair of posterior processes subtriangular and directed dorsally. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: same color as femora. Spines along tibiae: present. Lateral propodeal carinae: broadly separated, but parallel for a short distance anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: with median areole or pair of pits. Median propodeal carina: present.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 3. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: densely foveate, longitudinal sculpture irregular. T4 metasomal flanges: absent. T5 sculpture: densely foveate, interspaces not raised. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: with medially truncate emargination, sloping down to apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: sparsely foveolate, with tiny pits in interspaces. S5 sculpture: sparsely foveate, with tiny pits in interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Body entirely dark brown, including antennae and legs. Frontal depression shallow and very broad, transverse carinae weakly indicated above dorsal separator; submedian carina indicated by ruga. Hyperoccipital carina indicated by rugae. Occipital carina complete, sinuate or omicron-shaped medially. Metascutellum deeply concave, emarginate or truncate apically, projecting dorsally. Tibiae with flattened spines. T1 lateral carina expanded laterally. Metasomal sterna with few or no longitudinal rugae. Female: A3 much shorter than pedicel. A4, A5 broader than long. T1 midlobe with 4 longitudinal carinae. T6 without metasomal flanges. Fore wing long enough to reach beyond metasomal apex. Main body of T6 concave apically.
Latin noun, genitive case, meaning “the barb of an arrow.”
[
Collected on
Holotype, female: AUSTRALIA: TAS, 1km NE Herrick,
Female. Body length 2.6–3.85 mm (n=20).
Radicle color and shade: same as scape, both yellowish or reddish. Pedicel color: same as scape. A3: shorter than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: concave. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: with transverse ledge, face sharply receding below it. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: absent. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: uniformly rounded dorsally. Occiput sculpture: umbilicate foveate medially, becoming smooth laterally. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate punctate; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: with shallow irregular carinae, posterodorsal corner with dense microsculpture. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent. Postmarginal vein: present. Fore wing apex at rest: exceeding metasomal apex; reaching middle of T6. Coxae color brightness: same color as femora.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 5; obscured by other raised sculpture. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: strongly convex, tapering and sloping down to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: densely setose, setal pits between very weak longitudinal rugae. S5 sculpture: densely setose, setal pits between very weak longitudinal rugae. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression shallow; transverse carinae present ventrally, interrupted; submedian carina indicated by rounded ruga. Hyperoccipital carina indicated by rugae. Occipital carina complete, convex medially. Metascutellum deeply concave, not or hardly emarginate apically, projecting dorsally. Postmarginal vein present but sometimes very short. Coxa not darker than rest of leg. T1 lateral carina expanded laterally. Metasomal flanges absent. Female: A4, A5 broader than long. Mesoscutellum without granulate sculpture. T1 midlobe with 4 longitudinal carinae. Main body of T6 not abruptly separated from apical rim, T6 not concave apically. Fore wing long enough to reach middle of T6 or beyond metasomal apex. Male: Flagellomeres beyond A3 not or hardly longer than broad. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach to or beyond metasomal apex. T7 tiny, truncate or very weakly emarginate.
[
Inhabits brigalow: [
Holotype, male, S. hyalinipennis: AUSTRALIA: QLD, tree foliage / forest, Gordonvale (Nelson), 30.IV.1913, A. P. Dodd, SAMA DB 32-001584 (deposited in SAMA). Other material: AUSTRALIA: 152 females, 73 males, ANIC DB 32-020077, 32-020078, 32-020119, 32-020120, OSUC 437877, OSUC 437879, OSUC 437882, OSUC 437886, OSUC 437891, OSUC 437971, OSUC 437972, OSUC 437973, OSUC 437974, OSUC 437979, OSUC 437980, OSUC 437981, OSUC 437982, OSUC 437983, OSUC 437986, OSUC 437990, OSUC 437991, OSUC 437994, OSUC 437995, OSUC 437996, OSUC 437997, OSUC 437998, OSUC 437999, OSUC 438002, OSUC 438003, OSUC 438019, OSUC 441298, OSUC 441300, OSUC 441301, OSUC 441308, OSUC 441311 (ANIC); OSUC 462602 (CNCI); OSUC 435938, 437887-437889, 437959, 437975, 437985, 437989, 451341-451345, QDPC 0-165625, QDPC 0-165626, QDPC 0-165629, QDPC 0-165633, QDPC 0-165637, QDPC 0-165640, QDPC 0-165643, QDPC 0-165661, QDPC 0-165664, QDPC 0-165672, QDPC 0-165673, QDPC 0-165759, QDPC 0-165761, QDPC 0-165762 (QDPC); OSUC 148477, 437885, 437978, 437993, 438024, 441309 (QMBA); OSUC 437005, 448605-448606, 448608-448609 (QPIM); UCRC ENT 151374 (UCRC); OSUC 437961-437966, 451361-451366 (UQIC); OSUC 437006-437009, 437880-437881, 437883-437884, 437890, 437892-437958, 437960, 437967-437970, 437976-437977, 437984, 437987-437988, 437992, 438000-438001, 438004-438018, 438020-438023, 438025-438038, 441293-441297, 441299, 441302-441307, 441310, 441312, 451346, 451367 (WINC).
Female. Body length 3.65–4.4 mm (n=11).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with transverse carinae. Dorsal portion of frontal depression: with some transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: absent. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate; rugose. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: irregularly foveate, with smooth area dorsally. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a flattened or rounded elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: expanded. Mesoscutellar rim medially: with notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: reaching base of T5. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: narrowly separated, angled anteriorly to become parallel. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: present. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: present as slightly protruding sharp corners. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present, blade-like with rounded margins. T6: broader than long. Major sculpture of T6: longitudinally striate; umbilicate foveate. Microsculpture of T6: absent. T6 medially: with medially truncate emargination, separated from apical rim. T6 metasomal flanges: present, very broadly rounded but with sharp apices. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: present, posteriorly complete. S6 apex in relation to T6: exposed to dorsal view by T6 emargination. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep, all carinae except dorsal separator interrupted medially, carinae present above dorsal separator; submedian carina distinct but irregular medially. Hyperoccipital carina absent. Occipital carina complete, medially sinuate. Mesoscutal midlobe densely foveate. Mesoscutellar rim not expanded, with median notch. Metascutellum scoop-like, projecting dorsally. Coxa darker than rest of leg. Postmarginal vein absent. Female: A3 longer than pedicel. A4, A5 broader than long. T4 with weak metasomal flanges. T5 with lobe-like metasomal flanges. T6 with broad, wavy lateral metasomal flanges. T6 broadly and deeply emarginate, truncate medially. S6 exposed to dorsal view, rounded apically. Male: A3, A4, A11 longer than broad. Mesoscutellum and mesoscutal midlobe posteriorly with small and densely set foveae, mesoscutellum without smooth area medially. T5 with weak metasomal flanges, T6 with lobe-like metasomal flanges. T7 with broad postero-lateral lobes, deeply emarginate medially, posterior margin broadly M-shaped.
Latin noun, genitive case, meaning “incision.”
[
Holotype, female: AUSTRALIA: SA, 14km WNW Renmark, Calperum Station, Bookmark Biosphere Reserve,
Female. Body length 4 mm (n=1).
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: longer than pedicel. A4: longer than broad. A5: broader than long.
Ventral clypeal margin: uniformly convex. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: present as a single carina. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: umbilicate punctate. Extra carina ventral to occipital carina: present, medially incomplete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate; absent. Major sculpture of gena posteroventrally: umbilicate foveate; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate punctate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate punctate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: absent. Major sculpture of mesoscutellum peripherally: umbilicate punctate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: with slight, indistinct sculpture dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T6. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: narrowly separated, angled anteriorly to become parallel. Setae in metasomal depression: present. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: unknown. T1 lateral carina: straight. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: truncate, separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression nearly flat, transverse carinae absent; submedian carina indicated by a weak ruga. Hyperoccipital carina indicated by rugae. Occipital carina absent medially, lateral corners not protruding. Metascutellum broad and short, concave. Postmarginal vein present. Coxa not darker than rest of leg. Metasomal depression setose. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female: A3 much longer than pedicel. A4 about as long as broad, A5 broader than long. Mesoscutellum without granulate sculpture. Fore wing long enough to reach middle of T6. T6 broader than long, blunt apically with main body of tergum raised above apical rim.
Latin noun, genitive case, meaning “smoothness.”
[
Holotype, female: AUSTRALIA: QLD, 3km ENE Mount Tozer,
Female. Body length 3.5–4.25 mm (n=5).
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: longer than pedicel. A4: longer than broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: with medially interrupted transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: granulate. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: transversely rugose. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: rugose; umbilicate punctate. Major sculpture of gena posteroventrally: rugose; umbilicate punctate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate punctate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: obliquely rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: granulate. Median mesoscutal carina: absent. Major sculpture of mesoscutellum centrally: absent; umbilicate punctate. Major sculpture of mesoscutellum peripherally: umbilicate punctate. Microsculpture of mesoscutellum centrally: absent; granulate. Microsculpture of mesoscutellum peripherally: absent; granulate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: smooth. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: exceeding metasomal apex; reaching middle of T6. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, but parallel for a short distance anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: more than 6. T1 lateral carina: straight. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: mostly smooth, with tiny umbilicate foveae. T4 metasomal flanges: absent. T5 sculpture: mostly smooth, with tiny umbilicate foveae. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: granulate. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: smooth, with tiny umbilicate foveae. S5 sculpture: smooth, with tiny umbilicate foveae. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Body and limbs entirely dark brown. Frontal depression nearly flat, not crossed by carinae; submedian carina weakly indicated by some rugae. Clypeus not elongate. Hyperoccipital carina indicated by rugae. Occipital carina complete, irregular or weakly convex medially. Notauli absent. Mesoscutellum smooth, with only some scattered setae. Metascutellum broad and short, concave dorsally, rugose. Postmarginal vein present. Metasomal depression sculptured antero-medially. T1 lateral carina not expanded. Metasomal sterna without longitudinal rugae or carinae. Metasomal flanges absent. Female: A3 longer than pedicel. A5 broader than long. T1 midlobe with 5 or more longitudinal carinae. T6 broader than long, apical rim not separated from main body of tergum. Male: All flagellomeres longer than broad. T1 midlobe with 3 longitudinal carinae. T7 very broad and short, without posterior lobes.
Latin noun, genitive case, meaning “smooth-abdomen.”
[
Holotype, female: AUSTRALIA: TAS, Bathurst Harbour, margin of copse / heathy sedgeland, Melaleuca,
Female. Body length 9.1–11 mm (n=20).
Radicle color and shade: same as scape, both yellowish or reddish. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: uniformly convex. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: hood-like, dorsally protruding. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: absent. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: uniformly rounded dorsally. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: with a series of arched carinae, posterodorsal corner with weak longitudinal rugae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate; transversely rugose. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: absent; umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent; punctate. Microsculpture of mesoscutellum peripherally: punctate. Mesoscutellar rim: expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: not reaching base of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: present. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: present and strongly protruding. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as strongly protruding acuminate flanges. T6: broader than long. Major sculpture of T6: umbilicate punctate; longitudinally striate; umbilicate foveate. Microsculpture of T6: absent. T6 medially: with deep emargination that is V-shaped medially, separated from apical rim. T6 metasomal flanges: present, very broadly rounded, with rounded apices. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: present, posteriorly complete. S6 apex in relation to T6: exposed to dorsal view by T6 emargination. S6 apex: truncate.
Both sexes: Frontal depression deep, all carinae complete medially; no carinae present above dorsal separator, submedian carina sharp. Hyperoccipital carina absent. Occipital carina complete, medially weakly convex. Mesoscutellar rim expanded and strongly sculptured, without median notch. Metascutellum scoop-like, projecting dorsally. Coxa same color as rest of leg. Postmarginal vein absent. Female: A3 longer than pedicel. A5, A6 broader than long. T4, T5, T6 with broad, flat metasomal flanges. T6 apically broadly and deeply emarginate, angularly emarginate medially. S6 exposed to dorsal view, truncate apically. Male: A3 longer than broad. A4 broader than long, A11 as long or longer than broad. Mesoscutellum and mesoscutal midlobe posteriorly with small foveae, mesoscutellum with smooth area medially. T5, T6 with metasomal flanges. T7 with long postero-lateral lobes, deeply emarginate medially, posterior margin M-shaped.
Latin noun, genitive case, meaning “rim.”
[
Holotype, female: AUSTRALIA: NSW, Fowlers Gap Research Station,
Female. Body length 3.75–5.65 mm (n=20).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel; as long as pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: uniformly convex. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: absent. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: with a series of arched carinae, posterodorsal corner with weak longitudinal rugae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: with large smooth areas. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: absent; granulate. Microsculpture of mesoscutal midlobe posteriorly: absent; punctate. Median mesoscutal carina: present as a ruga. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: punctate. Microsculpture of mesoscutellum peripherally: punctate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally and ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: not reaching base of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 6. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: present. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: present and strongly protruding. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as strongly protruding acuminate flanges. T6: broader than long. Major sculpture of T6: umbilicate punctate; longitudinally striate. Microsculpture of T6: absent. T6 medially: with deep emargination that is V-shaped medially, separated from apical rim. T6 metasomal flanges: present, very broadly rounded, with rounded apices. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: present, posteriorly complete. S6 apex in relation to T6: exposed to dorsal view by T6 emargination. S6 apex: truncate.
Both sexes: Frontal depression deep, all carinae complete medially; no carinae present above dorsal separator, submedian carina sharp. Hyperoccipital carina absent. Occipital carina complete, medially weakly convex. Mesoscutellar rim not expanded, but sometimes rugose, without median notch. Metascutellum scoop-like, projecting dorsally. Coxa same color as rest of leg. Postmarginal vein absent. Female: A3 longer than pedicel. A4, A5, A6 broader than long. T4, T5, with sharp metasomal flanges. T6 with flat, expanded lateral margins. T6 apically shallowly emarginate, with a short incision medially. S6 exposed to dorsal view, truncate apically.
Latin noun, genitive case, meaning “apex.”
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Holotype, female: AUSTRALIA: WA, Carnarvon Shire, Gascoyne Research Station, 3.X–7.X.1969, H. E. Evans & R. W. Matthews, OSUC 376712 (deposited in MCZC). Paratypes: AUSTRALIA: 66 females, OSUC 436006, 436010, 436041-436048, 436050, 436053, 437855, 437858-437859 (ANIC); OSUC 462738-462745 (CNCI); OSUC 376684, 376704-376705, 376707, 376710-376711, 376713-376715, 376722, 376733-376734 (MCZC); OSUC 436040 (MVMA); OSUC 148486 (QMBA); OSUC 448603 (QPIM); OSUC 148613-148614, 148616 (SAMA); OSUC 436039 (UQIC); OSUC 436038, 436049, 436051-436052, 437856, 449063-449080, 453990 (WINC).
Female. Body length 4.1–4.2 mm (n=2).
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: with oblique carina extending towards mouth corner. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: umbilicate foveate. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate. Microsculpture of gena anteroventrally: punctate. Microsculpture of gena posteroventrally: punctate.
Lateral pronotal area sculpture: mostly granulate, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a flattened or rounded elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: granulate. Microsculpture of mesoscutellum peripherally: granulate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: with slight, indistinct sculpture dorsally, smooth ventrally. Metascutellum shape: not emarginate, convex dorsally. Metascutellar setae: absent. Metascutellum sculpture: with many longitudinal rugae. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: unknown. Median propodeal carina: unknown.
T1 horn: present. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: straight. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: longer than broad. Major sculpture of T6: umbilicate punctate; longitudinally striate. Microsculpture of T6: granulate. T6 medially: tapering to a sharp point, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression concave, transverse carinae present ventrally; submedian carina weakly indicated by a single carina. Weak oblique carina extending from bottom of frontal depression towards lower margin of eye. Head directed downward. Hyperoccipital carina indicated by rugae. Occipital carina absent medially, with protruding lateral corners. Metascutellum tongue-like, very narrow, resting in long groove presented by propodeum medially. Postmarginal vein present. Coxa not darker than rest of leg. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female: A3 longer than pedicel. A4 longer than broad, A5 broader than long. Mesoscutellum with granulate sculpture. T1 midlobe with strong anterior horn, longitudinal carinae obscured by raised area. Fore wing long enough to reach middle of T5. T6 longer than broad.
Latin noun, genitive case, meaning “tongue.”
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Holotype, female: PAPUA NEW GUINEA: Chimbu Prov., New Guinea Isl., Kerowagi, 25.III.1964, D. H. Colless, OSUC 448567 (deposited in ANIC). Paratype: PAPUA NEW GUINEA: 1 female, OSUC 448568 (ANIC).
Female. unknown.
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A5 tyloid shape: narrow, linear. A6: broader than long. A11: longer than broad.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: granulate. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: broadly angular, with rounded median peak. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a ruga. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with many longitudinal rugae. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T6. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 midlobe longitudinal carinae: 3. T3 metasomal flanges: absent. T4 metasomal flanges: absent. T5 metasomal flanges: absent. T6 metasomal flanges: absent. T7: broadly emarginate, with sharply pointed posterolateral lobes.
Male: A4, A11 longer than broad. Frontal depression moderately deep and mostly smooth, with interrupted transverse carinae ventrally and no carinae above dorsal separator; submedian carina sharp. Hyperoccipital carina represented by a set of rugae. Metascutellum subrectangular, with strong rugae. T1 midlobe with 3 complete longitudinal carinae. Fore wing long enough to reach middle of T6. T7 emarginate with tiny sharp posterolateral apices.
Latin noun, genitive case, meaning “trough.”
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Holotype, male: AUSTRALIA: NSW, Lord Howe Island, no date, A. M. Lea, OSUC 453949 (deposited in AMSA). Paratypes: AUSTRALIA: 2 males, OSUC 453950 (ANIC); OSUC 453951 (WINC).
Female. Body length 2.6–3.2 mm (n=20).
Radicle color and shade: same as scape, both yellowish or reddish. Pedicel color: same as scape. A3: shorter than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: absent; present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: omicron-shaped, with sharp corners where median portion meets lateral portions. Occiput sculpture: transversely rugose. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: densely covered with setiferous puncta. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: punctate. Microsculpture of mesoscutellum peripherally: punctate. Mesoscutellar rim: expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, but parallel for a short distance anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: present as slightly protruding sharp corners. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as slightly protruding sharp corners. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: with broad emargination between protruding posterolateral corners, separated from apical rim. T6 metasomal flanges: present as spine-like structures posterolaterally. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Mesoscutum and mesoscutellum blue or green. Frontal depression shallow, transverse carinae absent; submedian carina absent or incomplete. Hyperoccipital carina indicated by strong rugae. Occipital carina complete, sinuate medially. Metascutellum deeply concave, emarginate apically, projecting dorsally. T1 lateral carina expanded laterally. Female: A3 not longer than pedicel. A4, A5 broader than long. T1 midlobe with 4 longitudinal carinae. T6 with narrow, sharp, subapical cornicle-like metasomal flanges. Fore wing long enough to reach middle of T6 or beyond metasomal apex. Main body of T6 not abruptly separated from apical rim. Male: A3 longer than pedicel. Most flagellomeres between A4 and A12 broader than long. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to exceed metasomal apex. T7 with elongate spine-like posterior projections.
Latin participle, not changing with gender, meaning “envying.” Derived from the Latin word for blueness.
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Holotype, female: AUSTRALIA: N QLD, Red Road turnoff, rainforest, Tully Falls Road, 750m, 8.XII–5.I.1990, pitfall trap, Monteith, Thompson & Janetzki, OSUC 148485 (deposited in QMBA). Paratypes: AUSTRALIA: 24 females, 65 males, ANIC DB 32-020086, 32-020087, OSUC 439592, OSUC 439600, OSUC 439601, OSUC 439602, OSUC 439603, OSUC 439604, OSUC 439605, OSUC 439606, OSUC 439607, OSUC 439608, OSUC 439609, OSUC 439610, OSUC 439612, OSUC 439614, OSUC 439652, OSUC 439654 (ANIC); OSUC 451351-451352 (BMNH); OSUC 227565, 227621, 227626, 227635, 462755-462759 (CNCI); OSUC 439591, 439613, QDPC 0-165648, QDPC 0-165676, QDPC 0-165684, QDPC 0-165699, QDPC 0-165705, QDPC 0-165760, QDPC 0-165774 (QDPC); OSUC 148372, 148376, 439596-439599, 439611 (QMBA); OSUC 451348, 451350 (UQIC); OSUC 439593-439595, 439615-439638, 439640-439651, 439653, 451347, 451349 (WINC).
Female. Body length 3.15–3.4 mm (n=3).
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: shorter than pedicel; as long as pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; irregularly rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a flattened or rounded elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, but parallel for a short distance anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: with median areole or pair of pits. Median propodeal carina: absent. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T6. Coxae color brightness: darker than femora.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression shallow and broad, not crossed by carinae; submedian carina very weak. Hyperoccipital carina indicated by rugae. Occipital carina complete, convex medially. Metascutellum deeply concave, truncate or slightly emarginate apically, projecting dorsally. Coxa darker than rest of leg. Postmarginal vein absent. T1 lateral carina expanded laterally. Metasomal flanges absent. Female: A3 not longer than pedicel. A4, A5, A6 broader than long. Mesoscutellum sparsely foveate, with broad smooth areas between foveae. Fore wing long enough to reach or exceed metasomal apex. T1 midlobe with 4 longitudinal carinae anteriorly. T6 broader than long, blunt or slightly concave apically.
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Holotype, female, S. magniclava: AUSTRALIA: NSW, no date, ANIC DB 32-012479 (deposited in ANIC). Other material: AUSTRALIA: 3 females, 1 male, ANIC DB 32-020118, OSUC 439674, OSUC 441387, OSUC 441388 (ANIC).
Female. Body length 4.2–4.55 mm (n=6).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: hood-like, dorsally protruding. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: broadly angular, with rounded median peak. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: with shallow irregular carinae, without microsculpture. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate; umbilicate punctate. Microsculpture of mesoscutellum centrally: punctate. Microsculpture of mesoscutellum peripherally: punctate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally and ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent. Postmarginal vein: absent, but marginal vein curving slightly at apex. Fore wing apex at rest: not reaching base of T5. Coxae color brightness: darker than femora.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: present. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: present as slightly protruding sharp corners. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as slightly protruding sharp corners. T6: broader than long. Major sculpture of T6: umbilicate punctate; longitudinally striate. Microsculpture of T6: absent. T6 medially: with medially truncate emargination, separated from apical rim. T6 metasomal flanges: present, very broadly rounded but with sharp apices. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: present, posteriorly complete. S6 apex in relation to T6: exposed to dorsal view by T6 emargination. S6 apex: truncate.
Both sexes: Frontal depression deep, all carinae except dorsal separator interrupted medially, no carinae present above dorsal separator; submedian carina sharp. Hyperoccipital carina absent. Occipital carina complete, medially weakly convex or forming a vague median peak. Mesoscutellar rim not expanded, without median notch. Metascutellum scoop-like, projecting dorsally. Coxa darker than rest of leg. Postmarginal vein absent. Female: A3 not longer than pedicel. A4, A5, A6 broader than long. T4 with tiny metasomal flanges; T5, T6 with broad, flat metasomal flanges. T6 apically broadly and deeply emarginate, truncate medially. S6 exposed to dorsal view, truncate apically. Male: A3 longer than broad. A4, A11 broader than long. Mesoscutellum and mesoscutal midlobe posteriorly with broad foveae, not smooth medially. T5, T6 with sharp metasomal flanges. T7 with rounded postero-lateral lobes, broadly emarginate medially.
[
On flower of mallee: [
Holotype, female, S. mirella: AUSTRALIA: WA, Yallingup, 1.XII-12.XII.1913, R. E. Turner, B.M. TYPE HYM. 9.511 (deposited in BMNH). Paratypes: AUSTRALIA: 1 female, 4 males, OSUC 435935 (ANIC); OSUC 462562, 462750-462752 (BMNH). Other material: AUSTRALIA: 6 females, 33 males, OSUC 462748 (ANIC); OSUC 449030-449031, 449037-449044, 449049-449055, 449058-449062 (BMNH); OSUC 462746-462747 (CNCI); OSUC 435936 (QDPC); OSUC 449045-449048, 449057 (UQIC); WAM Ent. Reg No. 70285 (WAMP); OSUC 449032-449036, 449056, 462749 (WINC).
Female. Body length 3.1–4.55 mm (n=20).
Radicle color and shade: same as scape, both yellowish or reddish; darker than scape. Pedicel color: same as scape. A3: shorter than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: with medially interrupted transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; umbilicate punctate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate; longitudinally rugose. Major sculpture of mesoscutellum peripherally: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, but parallel for a short distance anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent. Postmarginal vein: absent. Fore wing apex at rest: reaching middle of T5. Coxae color brightness: same color as femora.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as slightly protruding sharp corners. T6: broader than long. Major sculpture of T6: longitudinally striate; umbilicate foveate. Microsculpture of T6: absent. T6 medially: strongly convex, tapering and sloping down to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep and broad but not parallel-sided, interrupted transverse carinae present dorsally; submedian carina sharp and strong. Hyperoccipital carina indicated by rugae. Occipital carina complete, convex medially. Metascutellum deeply concave, not or hardly emarginate apically, projecting dorsally. Postmarginal vein absent. Coxa not darker than rest of leg. T1 lateral carina expanded laterally. Metasomal flanges absent or with only very tiny sharp posterior corners. Female: A3 longer than pedicel. A4, A5 much broader than long. Mesoscutellum without granulate sculpture. T1 midlobe with 4 longitudinal carinae or with these obscured by raised sculpture. T6 without metasomal flanges, main body of tergum raised and vaguely separated from apical rim. Fore wing long enough to reach middle of T4 or base of T5. Male: Flagellomeres beyond A3 broader than long or only slightly longer than broad. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach to or beyond T7. T7 truncate or very weakly emarginate.
[
Holotype, male, S. montana: AUSTRALIA: QLD, nr. Cairns, slopes of Walshs Pyramid, forest, Gordonvale (Nelson), 2500ft, 2.VI.1913, sweeping, A. P. Dodd, SAMA DB 32-001586 (deposited in SAMA). Other material: AUSTRALIA: 112 females, 16 males, ANIC DB 32-020080, 32-020121, 32-020122, 32-020123, 32-020149, 32-020152, OSUC 359702, OSUC 359703, OSUC 429983, OSUC 429985, OSUC 429986, OSUC 441262, OSUC 441263, OSUC 441264, OSUC 441265, OSUC 441266, OSUC 441267, OSUC 441268, OSUC 441269, OSUC 441270, OSUC 441272, OSUC 441274, OSUC 441275, OSUC 441276, OSUC 441277, OSUC 441278, OSUC 441279, OSUC 441280, OSUC 441281, OSUC 441282, OSUC 441283, OSUC 441284, OSUC 441285, OSUC 441286, OSUC 441287, OSUC 441288, OSUC 441290, OSUC 441291, OSUC 441292, OSUC 441313, OSUC 441314, OSUC 441315, OSUC 441316, OSUC 441317, OSUC 441318, OSUC 441319, OSUC 441320, OSUC 441321, OSUC 441322, OSUC 441323, OSUC 441324, OSUC 441325, OSUC 441326, OSUC 441327, OSUC 441328, OSUC 441329, OSUC 441330, OSUC 441333, OSUC 441334, OSUC 441335, OSUC 441336, OSUC 441337, OSUC 441338, OSUC 441341, OSUC 441342, OSUC 441343, OSUC 441354, OSUC 441357, OSUC 441358, OSUC 441359, OSUC 441360, OSUC 441361, OSUC 441362, OSUC 441363, OSUC 441364 (ANIC); OSUC 445342 (BMNH); OSUC 441339-441340, 441349-441350 (MVMA); OSUC 148364, 429987-429988, 441271, 441351, QM Reg. No. T35148, QM Reg. No. T35150, QM Reg. No. T35152 (QMBA); OSUC 429984, 441353, 441355, 445341, 445343 (UQIC); OSUC 268189, 268191-268193, 268195, 268200-268201, 268203, 268207, 268214-268215 (USNM); OSUC 359696-359701, 441261, 441273, 441289, 441331-441332, 441344-441348, 441352, 441356, 445338-445339, 445344-445345, 453935-453936 (WINC).
Female. Body length 4.35–4.6 mm (n=3).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: longer than broad. A5: longer than broad.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: with oblique carina extending towards mouth corner. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: granulate. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: umbilicate foveate. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: mostly granulate, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a flattened or rounded elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally and ventrally. Metascutellum shape: not emarginate, convex dorsally. Metascutellar setae: absent. Metascutellum sculpture: with a median carina, otherwise weakly sculptured. Postmarginal vein: present. Fore wing apex at rest: not reaching base of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: present. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: straight. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: longer than broad. Major sculpture of T6: umbilicate punctate; longitudinally striate. Microsculpture of T6: granulate. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression nearly flat, transverse carinae absent; submedian carina indicated by a few weak rugae. Oblique carina extending from bottom of frontal depression towards lower margin of eye. Head directed downward. Hyperoccipital carina indicated by rugae. Occipital carina absent medially, with protruding lateral corners. Metascutellum narrow, slightly convex. Postmarginal vein present. Coxa not darker than rest of leg. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female: A3 longer than pedicel. A4, A5 about as long as broad. Mesoscutellum without granulate sculpture. T1 midlobe carinae obscured by raised smooth area. Fore wing long enough to reach middle of T4. T6 longer than broad.
New Latin noun, genitive case, meaning “moustache.”
[
Holotype, female: AUSTRALIA: QLD, light trap 18, 15km WNW South Johnstone, 1986, light trap, Fay & Halfpapp, OSUC 368224 (deposited in QMBA). Paratypes: AUSTRALIA: 2 females, OSUC 227614, 227623 (CNCI).
Female. unknown.
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: longer than pedicel. A5 tyloid shape: narrow, linear. A6: longer than broad. A11: longer than broad.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: smooth. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate punctate. Major sculpture of mesoscutal midlobe posteriorly: umbilicate punctate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: granulate. Median mesoscutal carina: present as a flattened or rounded elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: granulate. Microsculpture of mesoscutellum peripherally: granulate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: with median areole or pair of pits. Median propodeal carina: absent.
T1 midlobe longitudinal carinae: 4. T3 metasomal flanges: absent. T4 metasomal flanges: absent. T5 metasomal flanges: absent. T6 metasomal flanges: absent. T7: broadly and deeply emarginate, with rounded posterolateral margins; weakly emarginate.
Male: Interantennal process elongate, extending far beyond antennal insertions. Frontal depression low and flat, with some rounded transverse carinae. Hyperoccipital carina indicated by a strong ruga; occipital carina incomplete medially; occiput almost entirely smooth. Mesoscutum and mesoscutellum granulate, without strong sculpture.
Latin noun, genitive case, meaning “nose.”
[
Holotype, male: PAPUA NEW GUINEA: West New Britain Prov., New Britain Isl., primary forest edge, Cape Hoskins, 21.VI-27.VI.1973, malaise trap, Stibick, OSUC 368915 (deposited in CNCI). Paratype: PAPUA NEW GUINEA: 1 male, OSUC 369128 (CNCI).
Female. Body length 3.95–4.3 mm (n=6).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse foveae. Dorsal portion of frontal depression: with medially interrupted transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: granulate. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a flattened or rounded elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: punctate. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent. Postmarginal vein: present. Fore wing apex at rest: not reaching base of T5. Coxae color brightness: same color as femora.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as slightly protruding sharp corners. T6: broader than long. Major sculpture of T6: longitudinally striate; umbilicate foveate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: present. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep and broad, without transverse carinae; submedian carina sharp but irregular. Hyperoccipital carina indicated by weak rugae. Occipital carina complete, weakly convex or sinuate medially. Metascutellum deeply concave, truncate apically, projecting dorsally. Postmarginal vein absent. Coxa same color as rest of leg. T1 lateral carina expanded laterally. Female: A3 longer than pedicel. A4, A5 broader than long. Mesoscutellum without granulate sculpture. T1 midlobe carinae obscured by raised sculpture. T5 with tiny sharp posterior metasomal flanges, T6 with a narrow peripheral carina, main body of T6 raised and abruptly separated from apical rim. Fore wing long enough to reach middle of T4 or base of T5. Male: All flagellomeres past A3 as broad or broader than long. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach T7. T6 with tiny sharp metasomal flanges. T7 truncate, arched.
[
Holotype, female, S. nigriclava: AUSTRALIA: NSW, no date, ANIC DB 32-012569 (deposited in ANIC). Other material: AUSTRALIA: 7 females, 2 males, OSUC 441379-441382, 441385-441386 (ANIC); OSUC 148370, 441384 (QMBA); OSUC 441383 (WINC).
Female. Body length 2.35–3.45 mm (n=20).
Radicle color and shade: darker than scape. Pedicel color: same as scape; at least partially darker than scape. A3: shorter than pedicel. A4: longer than broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: absent. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: absent. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: uniformly rounded dorsally. Occiput sculpture: umbilicate foveate medially, becoming smooth laterally. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: with shallow irregular carinae, posterodorsal corner with dense microsculpture. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate; umbilicate punctate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: absent; umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent; punctate. Microsculpture of mesoscutellum peripherally: absent; punctate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally; smooth. Metascutellum shape: deeply emarginate, with the resulting pair of posterior processes subtriangular and directed dorsally. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent. Postmarginal vein: absent. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: darker than femora.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: slightly emarginate, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: densely setose, setal pits between very weak longitudinal rugae. S5 sculpture: densely setose, setal pits between very weak longitudinal rugae. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression shallow, transverse carinae absent or broadly interrupted ventrally, no carinae present above dorsal separator; submedian carina absent medially. Hyperoccipital carina indicated by weak rugae. Occipital carina complete, weakly convex medially. Metascutellum deeply concave, emarginate apically, projecting dorsally. Postmarginal vein absent. Coxa darker than rest of leg. T1 lateral carina expanded laterally. Metasomal flanges absent. Female: A3 longer than pedicel. A4 about as long as broad, A5 broader than long. Mesoscutellum without granulate sculpture. T1 midlobe with 4 longitudinal carinae. T6 without metasomal flanges, emarginate apically. Fore wing long enough to exceed metasomal apex. Male: A4 broader than long, A11 longer than broad. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach beyond T7. T7 emarginate apically. The smaller-bodied
[
On sticky seed of
Holotype, male, S. nigricoxa: AUSTRALIA: QLD, summit of mountain range, grass / forest, Gordonvale (Nelson), 1500ft, 30.V.1913, sweeping, A. A. Girault, SAMA DB 32-001587 (deposited in SAMA). Other material: AUSTRALIA: 62 females, 152 males, ANIC DB 32-020124, 32-020125, 32-020144, OSUC 359709, OSUC 359710, OSUC 359711, OSUC 438058, OSUC 438059, OSUC 438060, OSUC 438061, OSUC 438062, OSUC 438063, OSUC 438064, OSUC 438067, OSUC 438069, OSUC 438072, OSUC 438093, OSUC 438094, OSUC 438095, OSUC 438096, OSUC 438097, OSUC 438098, OSUC 438099, OSUC 438100, OSUC 438102, OSUC 438103, OSUC 438110, OSUC 438111, OSUC 438112, OSUC 438113, OSUC 438114, OSUC 438115, OSUC 438116, OSUC 438117, OSUC 438118, OSUC 438119, OSUC 438120, OSUC 438121, OSUC 438122, OSUC 438123, OSUC 438133, OSUC 438134, OSUC 438135, OSUC 438136, OSUC 438137, OSUC 438138, OSUC 438139, OSUC 438140, OSUC 438141, OSUC 438142, OSUC 438143, OSUC 438152 (ANIC); OSUC 449022, 451335, 451359 (BMNH); OSUC 462598-462601 (CNCI); OSUC 438068, 438104, 438108, 438182, QDPC 0-165634, QDPC 0-165641, QDPC 0-165645, QDPC 0-165651, QDPC 0-165662, QDPC 0-165668, QDPC 0-165670, QDPC 0-165680, QDPC 0-165681, QDPC 0-165683, QDPC 0-165685, QDPC 0-165689, QDPC 0-165697, QDPC 0-165700, QDPC 0-165710, QDPC 0-165712, QDPC 0-165716, QDPC 0-165717, QDPC 0-165722, QDPC 0-165733, QDPC 0-165742, QDPC 0-165743, QDPC 0-165744, QDPC 0-165748, QDPC 0-165755, QDPC 0-165763, QDPC 0-165764, QDPC 0-165766, QDPC 0-165767, QDPC 0-165783 (QDPC); OSUC 438073, 438092, 438101 (QMBA); OSUC 448607 (QPIM); OSUC 438070, 449028, 451329-451334, 451339-451340 (UQIC); OSUC 359704-359708, 359712, 436936, 438039-438057, 438065-438066, 438071, 438074-438091, 438105-438107, 438109, 438124-438132, 438144-438147, 438149-438151, 438153-438181, 438183-438186, 449029, 451336-451338, 451357-451358, 451360 (WINC).
Female. Body length 2.9–3.55 mm (n=20).
Radicle color and shade: darker than scape. Pedicel color: at least partially darker than scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: omicron-shaped, with sharp corners where median portion meets lateral portions. Occiput sculpture: transversely rugose. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: absent; umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: deeply emarginate, with the resulting pair of posterior processes subtriangular and directed dorsally. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, but parallel for a short distance anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: with median areole or pair of pits. Median propodeal carina: present.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: slightly emarginate, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Mesoscutum and mesoscutellum black. Frontal depression shallow, transverse carinae absent or interrupted; submedian carina absent. Hyperoccipital carina indicated by rugae. Occipital carina complete, sinuate medially. Metascutellum deeply concave, emarginate apically, projecting dorsally. Coxa same color as rest of leg. T1 lateral carina expanded laterally. Female: A3 not shorter than pedicel. A4, A5 broader than long. T1 midlobe with 4 longitudinal carinae. T6 without metasomal flanges. Fore wing long enough to reach middle of T6 or beyond metasomal apex. Main body of T6 not abruptly separated from apical rim. Male: A3 longer than pedicel. A4, A11 longer than broad. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to exceed metasomal apex. T7 with elongate spine-like posterior projections.
Latin noun, genitive case, meaning “brightness.”
[
Holotype, female: AUSTRALIA: QLD, Maroochy Horticultural Research Station, Nambour, 8.III-15.III.1985, malaise trap, OSUC 359530 (deposited in QMBA). Paratypes: AUSTRALIA: 55 females, 24 males, ANIC DB 32-020089, 32-020091, 32-020092, 32-020093, 32-020101, 32-020102, 32-020143, OSUC 359540, OSUC 359541, OSUC 359543, OSUC 359544, OSUC 359545, OSUC 359547, OSUC 359548, OSUC 359552, OSUC 359553, OSUC 359555, OSUC 359558, OSUC 359559, OSUC 359561 (ANIC); OSUC 449087 (BMNH); OSUC 227550, 227632-227633 (CNCI); OSUC 359549-359550, QDPC 0-165631, QDPC 0-165677 (QDPC); OSUC 148382, 359551, 435937, QM Reg. No. T35135, QM Reg. No. T35137, QM Reg. No. T35138, QM Reg. No. T35141, QM Reg. No. T35143, QM Reg. No. T35144, QM Reg. No. T35146 (QMBA); OSUC 359527-359529, 359531-359539, 359542, 359546, 359554, 359556-359557, 359560, 359562-359574, 436937-436938, 449081-449086, 449088-449089 (WINC).
Female. Body length 2.65–3.7 mm (n=20).
Radicle color and shade: same as scape, both yellowish or reddish. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: present. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: absent. Hyperoccipital carina: present as a single carina. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: smooth. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: rugose; umbilicate punctate; absent. Major sculpture of gena posteroventrally: umbilicate punctate; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate punctate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: longitudinally rugose; umbilicate punctate. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent. Major sculpture of mesoscutellum centrally: longitudinally rugose; obliquely rugose; umbilicate punctate. Major sculpture of mesoscutellum peripherally: obliquely rugose; umbilicate punctate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: deeply emarginate, with the resulting pair of posterior processes subtriangular and directed dorsally. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: with median areole or pair of pits. Median propodeal carina: present.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: straight; protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: densely foveate, longitudinal sculpture irregular. T4 metasomal flanges: absent. T5 sculpture: densely foveate, longitudinal sculpture irregular. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Latin noun, genitive case, meaning “obliqueness.”
[
Holotype, female: AUSTRALIA: QLD, GS1, Mount Haig,
Female. Body length 4–4.15 mm (n=5).
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with elongate median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: with medially interrupted transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: granulate. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: uniformly rounded dorsally. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: rugose; umbilicate punctate. Major sculpture of gena posteroventrally: rugose; umbilicate punctate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate punctate; irregularly rugose. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: longitudinally rugose; umbilicate punctate. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent. Major sculpture of mesoscutellum centrally: umbilicate punctate. Major sculpture of mesoscutellum peripherally: umbilicate punctate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: smooth. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: present. Metascutellum sculpture: dense and irregular. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, but parallel for a short distance anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: with median areole or pair of pits. Median propodeal carina: present.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: more than 6. T1 lateral carina: straight. T2 sculpture: mostly smooth with setiferous puncta medially, laterally with some longitudinal carinae. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: mostly smooth, with tiny umbilicate foveae. T4 metasomal flanges: absent. T5 sculpture: mostly smooth, with tiny umbilicate foveae. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: granulate. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: smooth, with tiny umbilicate foveae. S5 sculpture: smooth, with tiny umbilicate foveae. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Body and limbs entirely dark brown. Frontal depression nearly flat, not crossed by carinae; submedian carina absent or weakly indicated by some rugae. Clypeus elongate. Hyperoccipital carina indicated by rugae. Occipital carina complete convex medially. Mesoscutellum smooth, with only some scattered setae. Metascutellum dorsally setose, broad and short, rugose. Postmarginal vein present. Metasomal depression sculptured antero-medially. T1 lateral carina not expanded. Metasomal sterna without longitudinal rugae or carinae. Metasomal flanges absent. Female: A3 longer than pedicel. A5 broader than long. T1 midlobe with 6 or more longitudinal carinae. T6 broader than long, apical rim not separated from main body of tergum.
Latin noun, genitive case, meaning “oblong clypeus.”
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Holotype, female: AUSTRALIA: TAS, 14km S Bronte Park,
Female. Body length 2.8–3.8 mm (n=20).
Radicle color and shade: same as scape, both yellowish or reddish. Pedicel color: same as scape. A3: shorter than pedicel; as long as pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: present as multiple regular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: uniformly rounded dorsally. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a ruga. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T6. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, but parallel for a short distance anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Mesoscutum and mesoscutellum black. Frontal depression shallow, transverse carinae absent or interrupted; submedian carina indicated by a set of irregular elevations or weak carinae. Hyperoccipital carina indicated by sharp carina. Occipital carina complete, convex medially. Mesoscutal midlobe with many fine longitudinal rugae posteriorly. Metascutellum deeply concave, broad and short, projecting dorsally. Coxa darker than rest of leg. T1 lateral carina expanded laterally. Metasomal flanges absent. Female: A3 longer than pedicel. A4, A5 broader than long. T1 midlobe with 4 longitudinal carinae. T6 without metasomal flanges. Fore wing long enough to reach middle of T6 or exceed metasomal apex. Male: A4, A11 longer than broad. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach metasomal apex. T7 much broader than long, slightly concave apically, without posterior projections.
Latin noun, genitive case, meaning “a plug.” Refers to the compact, stout body shape.
[
Holotype, female: AUSTRALIA: QLD, via Samford, open forest, Stony Creek,
Female. Body length 2.7–3.2 mm (n=20).
Radicle color and shade: same as scape, both yellowish or reddish. Pedicel color: same as scape. A3: shorter than pedicel; as long as pedicel. A4: longer than broad. A5: longer than broad.
Ventral clypeal margin: concave. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: granulate. Hyperoccipital carina: present as a single carina. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: smooth. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: rugose; umbilicate punctate. Major sculpture of gena posteroventrally: rugose; umbilicate punctate. Microsculpture of gena anteroventrally: granulate. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: irregularly sculptured. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate punctate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent. Major sculpture of mesoscutellum centrally: umbilicate foveate; umbilicate punctate. Major sculpture of mesoscutellum peripherally: umbilicate punctate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent; granulate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T6. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: meeting near propodeal midlength. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: with series of longitudinal carinae extending to lateral propodeal carinae. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: straight. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: densely setose, setal pits between very weak longitudinal rugae. S5 sculpture: densely setose, setal pits between very weak longitudinal rugae. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression nearly parallel-sided, transverse carinae absent; submedian carina sharp. Hyperoccipital carina sharp and strong. Occipital carina incomplete, lateral portions almost reaching hyperoccipital carina; occiput smooth, with only a few setiferous puncta and fine rugae. Metascutellum with a small concave postero-medial area, laterally with broad longitudinally striate area, weakly emarginate, projecting dorsally. Postmarginal vein present. Coxa not darker than rest of leg. T1 lateral carina not expanded laterally. Female: A3 longer than pedicel. A4 longer than broad, A5 nearly as long as broad. Mesoscutellum nearly smooth, with a few setiferous puncta. T1 midlobe with 5 longitudinal carinae. Fore wing long enough to reach or exceed T6. Male: All flagellomeres longer than broad. T1 midlobe with 3 longitudinal carinae, hardly raised above sidelobes. Mesoscutellum with strong longitudinal rugae and no granulate sculpture. Fore wing long enough to reach to or beyond T7. T7 truncate, steeply sloping.
Latin noun, genitive case, meaning “eye.”
[
Holotype, female: AUSTRALIA: QLD, Mount Glorious, 1977, malaise trap, OSUC 442277 (deposited in QMBA). Paratypes: AUSTRALIA: 35 females, 36 males, Australian Museum K245262, K245267, K245268, K245270, K245274 (AMSA); ANIC DB 32-020075, 32-020076 (ANIC); OSUC 227548, 227608, 462563-462566, 462727-462728 (CNCI); OSUC 442273, 442278, 442301-442302, 451276-451277, QDPC 0-165656, QDPC 0-165658 (QDPC); OSUC 442298 (QMBA); OSUC 359782-359787, 442271-442272, 442274-442276, 442279-442297, 442299-442300, 442303, 442305-442310, 451274-451275, 451278-451283 (WINC).
Female. Body length 2.5–3.4 mm (n=18).
Radicle color and shade: same as scape, both yellowish or reddish. Pedicel color: same as scape. A3: longer than pedicel. A4: longer than broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with sharp lateral corners; omicron-shaped, with sharp corners where median portion meets lateral portions. Occiput sculpture: transversely rugose. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: densely covered with setiferous puncta. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate; irregularly rugose. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; irregularly rugose. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate; irregularly rugose. Major sculpture of mesoscutellum peripherally: umbilicate foveate; irregularly rugose. Microsculpture of mesoscutellum centrally: absent; punctate. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: with slight, indistinct sculpture dorsally, smooth ventrally. Metascutellum shape: deeply emarginate, with the resulting pair of posterior processes subtriangular and directed dorsally. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 5. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: slightly emarginate, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression shallow; transverse carinae present ventrally, interrupted; submedian carina very weak or absent medially. Hyperoccipital carina indicated by rugae. Occipital carina complete, broadly omicron-shaped medially. Metascutellum deeply concave, emarginate apically, projecting dorsally. Postmarginal vein absent. Coxa not darker than rest of leg. T1 lateral carina expanded laterally. Metasomal flanges absent. Female: A4 longer than broad, A5 almost as long as broad. Mesoscutellum without granulate sculpture. T1 midlobe with 4 longitudinal carinae. Main body of T6 not abruptly separated from apical rim, T6 slightly concave apically. Fore wing long enough to reach beyond metasomal apex. Male: All flagellomeres longer than broad. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach beyond metasomal apex. T7 tiny, truncate.
Latin noun, genitive case, meaning “vault.”
[
Holotype, female: AUSTRALIA: QLD, cableway base station, Bellenden Ker Range, 100m, 17.X–9.XI.1981, window trap, OSUC 359578 (deposited in QMBA). Paratypes: AUSTRALIA: 17 females, 19 males, ANIC DB 32-020100, 32-020145, OSUC 359583, OSUC 359584, OSUC 359585, OSUC 359586, OSUC 359587, OSUC 359602, OSUC 359603, OSUC 359604, OSUC 436939 (ANIC); QDPC 0-165695 (QDPC); OSUC 148367, 359582, 359588, 359600-359601 (QMBA); OSUC 359605 (UQIC); OSUC 359575-359577, 359579-359581, 359589-359599, 451356 (WINC).
Female. Body length 3.2–3.3 mm (n=2).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: shorter than pedicel. A4: as long as broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse foveae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: present as a single carina. Carina connecting occipital carina to hyperoccipital carina: present. Occipital carina: present laterally, absent medially. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: rugose; umbilicate punctate. Major sculpture of gena posteroventrally: rugose; umbilicate punctate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a ruga. Major sculpture of mesoscutellum centrally: longitudinally rugose; umbilicate punctate. Major sculpture of mesoscutellum peripherally: longitudinally rugose; umbilicate punctate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally and ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: reaching near apex of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: present. Number of longitudinal carinae of T1 midlobe: 5. T1 lateral carina: straight. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Frontal depression shallow, transverse carinae absent; submedian carina weak. Hyperoccipital carina indicated by rugae. Occipital carina connected to hyperoccipital carina by a weak longitudinal carina, laterally with strong corners and medially absent; area between occipital and hyperoccipital carinae densely sculptured and having many short setae. Mesoscutellum and mesoscutal midlobe posteriorly with strong longitudinal rugae. Metascutellum exceedingly tiny, concave. Postmarginal vein present. Coxa not darker than rest of leg. Metasomal depression smooth. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female only: A3 shorter than pedicel. A4 about as long as broad, A5 broader than long. T1 midlobe slight smooth area obscuring its longitudinal carinae. Fore wing long enough to reach middle of T6. T6 slightly broader than long.
Latin noun, genitive case, meaning “comb.”
[
Holotype, female: AUSTRALIA: QLD, rainforest, Mount Glorious, 28.II.1984, sweeping, I. D. Galloway, OSUC 368207 (deposited in QMBA). Paratypes: AUSTRALIA: 2 females, QDPC 0-165773 (QDPC); OSUC 449001 (WINC).
Female. Body length 3.25–3.9 mm (n=14).
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: longer than pedicel. A4: longer than broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: uniformly rounded dorsally. Occiput sculpture: umbilicate foveate. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate punctate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent; present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: absent. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: deeply emarginate, with the resulting pair of posterior processes subtriangular and directed dorsally. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate foveate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Body and limbs entirely dark brown. Frontal depression shallow and broad, transverse carinae absent or interrupted; submedian carina weak. Hyperoccipital carina indicated by rugae. Occipital carina complete, convex medially. Metascutellum subrectangular, deeply concave, emarginate apically, projecting dorsally. T1 lateral carina expanded laterally. Metasomal flanges absent. Female: A3 longer than pedicel. A4 longer than broad; A5, A6 broader than long. T1 midlobe with a slight smooth bump obscuring anterior carinae. Fore wing long enough to reach middle of T5. T6 broader than long, rounded apically. Male: A3 longer than pedicel. most flagellomeres between A4 and A12 broader than long. T1 midlobe with 4 longitudinal midlobe carinae. Fore wing long enough to exceed metasomal apex. T7 much broader than long, blunt apically, without posterior projections.
Latin noun, genitive case, meaning “thumb.” Refers to the shape of the fore wing venation.
[
Under bark of
Holotype, female: AUSTRALIA: ACT, Piccadilly Circus,
Female. Body length 4.5–4.7 mm (n=4).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: longer than broad. A5: longer than broad.
Ventral clypeal margin: concave. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: uniformly rounded dorsally. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate; absent. Major sculpture of gena posteroventrally: umbilicate foveate; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: irregularly sculptured. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a ruga. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally and ventrally. Metascutellum shape: not emarginate, forming a flat, concave shelf. Metascutellar setae: absent. Metascutellum sculpture: dense and irregular. Postmarginal vein: present. Fore wing apex at rest: reaching base of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: present. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: straight. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: longer than broad. Major sculpture of T6: umbilicate punctate; longitudinally striate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression with broadly interrupted transverse carinae; submedian carina sharp. Hyperoccipital carina indicated by rugae. Occipital carina convex medially. Postmarginal vein present. Coxa not darker than rest of leg. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female only: A3 longer than pedicel. A4 longer than broad, A5 slightly longer than broad. Mesoscutellum without granulate sculpture. Metascutellum with irregular rugose sculpture. T1 midlobe carinae obscured by raised area. Fore wing long enough to reach base of T5.
Latin noun, genitive case, meaning “length.”
[
Holotype, female: AUSTRALIA: QLD, Mount Glorious, VII-1977 - XII-1977, malaise trap, A. Hiller, OSUC 368221 (deposited in QMBA). Paratypes: AUSTRALIA: 3 females, OSUC 227567, 462572 (CNCI); OSUC 437002 (WINC).
Female. Body length 3.95 mm (n=1).
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: longer than pedicel. A4: longer than broad. A5: longer than broad.
Ventral clypeal margin: concave. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: umbilicate foveate. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose; umbilicate punctate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; obliquely rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent; granulate. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally and ventrally. Metascutellum shape: not emarginate, forming a flat, concave shelf. Metascutellar setae: absent. Metascutellum sculpture: with three parallel and evenly spaced longitudinal carinae. Postmarginal vein: present. Fore wing apex at rest: reaching near apex of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: present. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: straight. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: longer than broad. Major sculpture of T6: umbilicate punctate; longitudinally striate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression shallow, transverse carinae weak and interrupted; submedian carina strong but irregular. Hyperoccipital carina indicated by rugae. Occipital carina becoming weak or absent medially. Postmarginal vein present. Coxa not darker than rest of leg. Metasomal depression smooth. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female only: A3 longer than pedicel. A4 longer than broad, A5 as long or longer than broad. Mesoscutellum without granulate sculpture. Metascutellum with 3 longitudinal carinae and some weak sculpture underlying them. T1 midlobe carinae obscured by raised smooth area. Fore wing long enough to reach middle of T5.
Latin noun, genitive case, meaning “an extension.”
[
Holotype, female: AUSTRALIA: QLD, Mount Tamborine, XII-1925, A. P. Dodd, ANIC DB 32-020073 (deposited in ANIC).
Female. Body length 4–4.65 mm (n=19).
Radicle color and shade: darker than scape. Pedicel color: at least partially darker than scape. A3: longer than pedicel. A4: broader than long; as long as broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: present. Occipital carina: present laterally, absent medially. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate; rugose; umbilicate punctate. Major sculpture of gena posteroventrally: rugose; umbilicate punctate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a ruga. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally and ventrally. Metascutellum shape: not emarginate, forming a flat, concave shelf. Metascutellar setae: absent. Metascutellum sculpture: with a longitudinal median ruga and some weak transverse carinae. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: present. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: straight. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: longer than broad. Major sculpture of T6: umbilicate punctate; longitudinally striate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression flat, with oblique interrupted carinae and sometimes with an incomplete longitudinal carina; submedian carina weak. Hyperoccipital carina indicated by sharp rugae. Occipital carina connected to hyperoccipital carina by a weak longitudinal carina or irregular ruga, laterally with strong corners and medially sinuate; area between occipital and hyperoccipital carinae densely sculptured. Mesoscutellum densely foveolate. Metascutellum flat, rugose. Postmarginal vein present. Coxa not darker than rest of leg. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female: A3 not longer than pedicel. A4 slightly longer than broad, A5 broader than long. T1 with raised horn obscuring longitudinal carinae. Fore wing long enough to reach middle of T5 or base of T6. T6 slightly longer than broad. Male: A4 slightly longer than broad, A11 longer than broad. T1 midlobe with 3 longitudinal carinae. Fore wing not long enough to exceed metasomal apex. T7 truncate or slightly emarginate apically.
Latin noun, genitive case, meaning “rod.”
[
Holotype, female: AUSTRALIA: QLD, via Mount Carbine, Mount Windsor Tableland, 28.I–5.III.1981, malaise trap, OSUC 368185 (deposited in QMBA). Paratypes: AUSTRALIA: 20 females, 14 males, Australian Museum K256238 (AMSA); OSUC 368176, 368178-368184, 368186-368189 (ANIC); OSUC 448966 (BMNH); OSUC 462582, 462585-462586 (CNCI); OSUC 368177, QDPC 0-165630, QDPC 0-165750, QDPC 0-165784, QDPC 0-165790 (QDPC); OSUC 148478, 368191 (QMBA); OSUC 448601 (QPIM); OSUC 448955, 448957 (UQIC); OSUC 368175, 368190, 368192-368194, 448949-448950 (WINC).
Distinction between
Female. Body length 2.8–3.85 mm (n=12).
Radicle color and shade: darker than scape. Pedicel color: same as scape; at least partially darker than scape. A3: as long as pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse foveae. Dorsal portion of frontal depression: with some transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: broadly angular, with rounded median peak. Occiput sculpture: umbilicate foveate. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: with shallow irregular carinae, posterodorsal corner with dense microsculpture. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent; punctate. Microsculpture of mesoscutellum peripherally: absent; punctate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: reaching base of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: present as slightly protruding sharp corners. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as slightly protruding sharp corners. T6: broader than long. Major sculpture of T6: umbilicate foveate. Microsculpture of T6: absent. T6 medially: with medially truncate emargination, sloping down to apical rim. T6 metasomal flanges: present as spine-like structures posterolaterally. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: exposed to dorsal view by T6 emargination. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep, with interrupted carinae ventrally, dorsal separator complete medially, carinae present above dorsal separator; submedian carina distinct but only weakly protruding. Hyperoccipital carina indicated by vague rugae. Occipital carina complete, medially sinuate. Mesoscutellar rim not expanded, without median notch. Metascutellum small, weakly emarginate apically, projecting dorsally. Coxa same color as rest of leg. Postmarginal vein absent. Female: A4, A5 broader than long. T4, T5 with sharp posterior corners. T6 abruptly narrower than T5, without expanded lateral margins but with narrow and sharp elongate posterior metasomal flanges. Main surface of T6 strongly emarginate medially, sharply raised above apical rim. S6 not exposed to dorsal view. Male: A4, A11 broader than long. Mesoscutellum and mesoscutal midlobe posteriorly with small and densely set foveae, mesoscutellum without smooth area medially. T5, T6 with sharp posterior corners; T6, T7 abruptly narrower than preceding tergum. T7 with narrow and sharp postero-lateral lobes, deeply emarginate, truncate medially.
Latin noun, genitive case, meaning “branch” or “prong.”
[
Collected on
Holotype, female: AUSTRALIA: NE QLD, Tea Tree Cave, 4km SE Chillagoe,
Female. Body length 2.55–3 mm (n=20).
Radicle color and shade: same as scape, both yellowish or reddish. Pedicel color: same as scape. A3: shorter than pedicel; as long as pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: concave. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: with transverse ledge, face sharply receding below it. Transverse curved rugae extending from frontal depression to eye: present. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: present as a single carina. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: smooth. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate; umbilicate punctate. Major sculpture of gena posteroventrally: umbilicate foveate; umbilicate punctate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: smooth anteriorly, densely setose posteriorly. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate; transversely rugose. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate punctate. Major sculpture of mesoscutellum peripherally: umbilicate foveate; umbilicate punctate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Spines along tibiae: absent. Lateral propodeal carinae: narrowly separated, angled anteriorly to become parallel. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: with median areole or pair of pits. Median propodeal carina: present. Postmarginal vein: absent. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: same color as femora.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: straight. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: slightly emarginate, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: densely setose, setal pits between very weak longitudinal rugae. S5 sculpture: densely setose, setal pits between very weak longitudinal rugae. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression shallow, weak transverse carinae present; submedian carina indicated by a set of weak rugae, flat or only weakly rounded dorsally. Hyperoccipital carina sharp and strong. Occipital carina incomplete, lateral portions nearly reaching hyperoccipital carina. Occiput almost entirely smooth, with a row of setiferous puncta. Mesoscutum with median carina. Metascutellum broad and concave, truncate or slightly concave apically, projecting dorsally. Postmarginal vein present. Coxa not darker than rest of leg. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female: A3 as long or longer than pedicel. A4, A5 broader than long. Mesoscutellum weakly sculptured, with only setiferous puncta. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach beyond T6. T6 broader than long. Male: All flagellomeres past A3 about as long as broad. Frontal depression without complete transverse carinae. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach beyond metasomal apex. T7 tiny, truncate.
[
Syntype, male, S. rugulosa: AUSTRALIA: QLD, nr. Cairns, jungle edge, Gordonvale (Nelson), no date, sweeping, A. P. Dodd, SAMA DB 32-001589 (deposited in SAMA). Syntype, female, S. rugulosa: Female holotype of, D. glabriscutellum: AUSTRALIA: QLD, nr. Cairns, plant foliage / jungle country, Harvey Creek, 12.X.1914, A. P. Dodd, SAMA DB 32-001585 (deposited in SAMA). AUSTRALIA: QLD, jungle edge, nr. Gordonvale (Nelson), no date, sweeping, A. P. Dodd, SAMA DB 32-001590 (deposited in SAMA). Paratype: AUSTRALIA: 1 female, ANIC DB 32-020109 (ANIC). Other material: AUSTRALIA: 49 females, 68 males, 442441b, ANIC DB 32-020104, ANIC DB 32-020105, ANIC DB 32-020106, ANIC DB 32-020107, ANIC DB 32-020108, OSUC 442402, OSUC 442403, OSUC 442404, OSUC 442405, OSUC 442406, OSUC 442407, OSUC 442408, OSUC 442409, OSUC 442413, OSUC 442414, OSUC 442415, OSUC 442416, OSUC 442417, OSUC 442418, OSUC 442419, OSUC 442420, OSUC 442421, OSUC 442422, OSUC 442423, OSUC 442424, OSUC 442425, OSUC 442426, OSUC 442427, OSUC 442428, OSUC 442429, OSUC 442430, OSUC 442431, OSUC 442432, OSUC 442433, OSUC 442434, OSUC 442435, OSUC 442436, OSUC 442438, OSUC 442439, OSUC 442440, OSUC 442442, OSUC 442443, OSUC 442444, OSUC 442445, OSUC 442446, OSUC 442448, OSUC 442449, OSUC 442450, OSUC 442451, OSUC 442452, OSUC 442453, OSUC 442454, OSUC 442455, OSUC 442456, OSUC 442457, OSUC 442458, OSUC 442459, OSUC 442460, OSUC 442461, OSUC 442462, OSUC 442463, OSUC 442464, OSUC 442465, OSUC 442466, OSUC 442467, OSUC 442468, OSUC 442469, OSUC 442470, OSUC 442471, OSUC 442473, OSUC 442474, OSUC 442475, OSUC 442477, OSUC 442478, OSUC 442479, OSUC 442480, OSUC 442481, OSUC 442482, OSUC 442484, OSUC 442488, OSUC 451299, OSUC 451300, OSUC 451302 (ANIC); OSUC 451301, 451303, 451305 (BMNH); OSUC 324331, 58660 (OSUC); OSUC 451304, QDPC 0-165688, QDPC 0-165691, QDPC 0-165694, QDPC 0-165701, QDPC 0-165703, QDPC 0-165713, QDPC 0-165724, QDPC 0-165731, QDPC 0-165735, QDPC 0-165737, QDPC 0-165745, QDPC 0-165749 (QDPC); OSUC 442437, 442485-442486 (QMBA); OSUC 448597, 448600 (QPIM); UCRC ENT 151376-151377 (UCRC); OSUC 442410-442412, 442447, 442472, 442476, 442483, 442487 (WINC).
Female. Body length 3.25–3.65 mm (n=4).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: longer than broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: with some transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: omicron-shaped, with sharp corners where median portion meets lateral portions. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, medially incomplete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate; longitudinally rugose. Major sculpture of mesoscutellum peripherally: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T5. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as strong posterior corners. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: with medially truncate emargination, sloping down to apical rim. T6 metasomal flanges: present as slightly expanded lateral rims, truncate posteriorly. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep, carinae interrupted medially, carinae present above dorsal separator; submedian carina absent or weak and irregular. Hyperoccipital carina indicated by rugae. Occipital carina complete, sinuate medially. Mesoscutellar rim not expanded. Metascutellum scoop-shaped, deeply concave, slightly emarginate apically, projecting dorsally. Coxa darker than rest of leg. T1 lateral carina expanded laterally. Female: A3 longer than pedicel. A4 longer than broad, A5, A6 broader than long. T1 midlobe with 4 longitudinal midlobe carinae anteriorly. Fore wing long enough to reach middle of T5. T4, T5 without metasomal flanges. Main body of T6 raised above apical rim and laterally abruptly separated from it, but medially not abruptly separated from it; lateral margins of T6 slightly expanded, forming a pair of lateral lobe-like shelves. Male: A3 longer than pedicel. A4 as broad or broader than long. A11 longer than broad. Mesoscutellum and mesoscutal midlobe posteriorly with small and densely set foveae, mesoscutellum without smooth area medially. T5, T6 without metasomal flanges. T7 with broad postero-lateral lobes, deeply emarginate medially, posterior margin roughly sculptured.
Latin noun, genitive case, meaning “breach.”
[
Holotype, female: AUSTRALIA: QLD, Townsville, 10.XII–17.XII.1987, malaise trap, T. Goertemiller, OSUC 268197 (deposited in USNM). Paratypes: AUSTRALIA: 7 females, 2 males, OSUC 268190, 268196, 268198-268199, 268202, 268204-268206, 268216 (USNM).
Female. Body length 3.25–3.5 mm (n=8).
Radicle color and shade: darker than scape; same as scape, both dark brown. Pedicel color: same as scape. A3: shorter than pedicel; as long as pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: absent. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: densely covered with setiferous puncta. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent; granulate. Median mesoscutal carina: present as a ruga. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: punctate. Microsculpture of mesoscutellum peripherally: punctate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, but parallel for a short distance anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4; obscured by other raised sculpture. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate foveate. Microsculpture of T6: absent. T6 medially: strongly convex, tapering and sloping down to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: truncate.
Both sexes: Frontal depression shallow and broad, crossed by some carinae above dorsal separator; submedian carina weak and irregular. Hyperoccipital carina indicated by rugae. Occipital carina complete, weakly convex medially. Metascutellum subrectangular, deeply concave, slightly emarginate apically, projecting dorsally. Coxa darker than rest of leg. T1 lateral carina expanded laterally. Metasomal flanges absent. Female: A3 not longer than pedicel. A4, A5, A6 broader than long. T1 midlobe with 4 longitudinal carinae anteriorly or with a slight smooth bump obscuring them. Fore wing long enough to reach or exceed metasomal apex. T6 broader than long, blunt apically. Male: A3 not longer than pedicel. A4, A11 broader than long. Fore wing long enough to reach T1 midlobe with 4 longitudinal midlobe carinae. T6 or to metasomal apex. T7 much broader than long, blunt apically, without posterior projections.
Latin noun, genitive case, meaning “a pack.”
[
Holotype, female: AUSTRALIA: WA, Mount Cooke, 13.I-28.I.1991, malaise trap, M. S. Harvey & J. M. Waldock, OSUC 439555 (deposited in WAMP). Paratypes: AUSTRALIA: 7 females, OSUC 439556, 439558-439563 (WINC).
Female. Body length 4.4–4.65 mm (n=3).
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: uniformly convex. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: hood-like, dorsally protruding. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: absent. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: with narrow curved smooth elevations. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: punctate. Median mesoscutal carina: present as a ruga. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent; punctate. Microsculpture of mesoscutellum peripherally: absent; punctate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: reaching base of T5. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: present as slightly protruding sharp corners. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as strongly protruding acuminate flanges. T6: broader than long. Major sculpture of T6: umbilicate punctate; longitudinally striate. Microsculpture of T6: absent. T6 medially: truncate, separated from apical rim. T6 metasomal flanges: present as slightly expanded lateral rims, truncate posteriorly. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: present, posteriorly complete. S6 apex in relation to T6: exposed to dorsal view by T6 emargination. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep, with interrupted carinae ventrally, dorsal separator complete medially, no carinae present above dorsal separator; submedian carina strongly defined. Hyperoccipital carina absent. Occipital carina complete, medially weakly convex. Mesoscutellar rim not expanded, without median notch. Metascutellum broad and concave, projecting dorsally. Coxa darker than rest of leg. Postmarginal vein absent. Female: A3 longer than pedicel. A4, A5 broader than long. T4 without metasomal flanges, T5 with lobe-like metasomal flanges. Main surface of T6 with slightly expanded lateral margins with sharp posterior corners, truncate or very slightly emarginate apically, but sharply separated from apical rim. S6 exposed to dorsal view, rounded apically.
Latin noun, genitive case, meaning “a separation.”
[
Holotype, female: AUSTRALIA: ACT, Canberra, 2.II.1950, E. F. Riek, OSUC 436037 (deposited in ANIC). Paratypes: AUSTRALIA: 3 females, 2 males, OSUC 436035-436036 (ANIC); OSUC 456180-456182 (WINC).
Female. Body length 4.05–4.5 mm (n=6).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: shorter than pedicel; as long as pedicel. A4: longer than broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: broadly angular, with rounded median peak. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: absent. Gena length: as long as eye. Major sculpture of gena anteroventrally: rugose; umbilicate punctate. Major sculpture of gena posteroventrally: rugose; umbilicate punctate. Microsculpture of gena anteroventrally: absent; granulate. Microsculpture of gena posteroventrally: absent; granulate.
Lateral pronotal area sculpture: with shallow irregular carinae, posterodorsal corner with dense microsculpture. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a ruga. Major sculpture of mesoscutellum centrally: umbilicate foveate; irregularly rugose. Major sculpture of mesoscutellum peripherally: umbilicate foveate; irregularly rugose. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: present. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: straight. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate; longitudinally striate. Microsculpture of T6: granulate. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression broad, transverse carinae absent; submedian carina strong. Gena greatly enlarged, about as long as the eye, strongly rugose. Hyperoccipital carina indicated by rugae. Occipital carina convex, forming a rounded peak medially. Metascutellum tiny, concave. Postmarginal vein present. Coxa not darker than rest of leg. Metasomal depression smooth. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female: A3 not longer than pedicel. A4 about as long as broad, A5 broader than long. Mesoscutellum without granulate sculpture. T1 midlobe carinae obscured by raised smooth area. Fore wing long enough to reach middle of T5. Male: A4 broader than long, A11 slightly longer than broad. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach middle of T6. T7 emarginate apically, with sharp posterior corners angled medially.
Latin noun, genitive case, meaning “squirrel.”
[
Holotype, female: AUSTRALIA: QLD, Mt. William, rainforest edge, Eungella National Park, 18.IV.1979, E Dahms, OSUC 368212 (deposited in QMBA). Paratypes: AUSTRALIA: 5 females, 3 males, OSUC 368208, 368213 (ANIC); OSUC 227564, 462576-462577 (CNCI); OSUC 448572, 451355 (QDPC); OSUC 368214 (WINC).
Female. Body length 2.45–3.4 mm (n=20).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: shorter than pedicel; as long as pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: present as a single carina. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: transversely rugose. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a ruga. Major sculpture of mesoscutellum centrally: umbilicate foveate; obliquely rugose. Major sculpture of mesoscutellum peripherally: umbilicate foveate; obliquely rugose. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: deeply emarginate, with the resulting pair of posterior processes subtriangular and directed dorsally. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Mesoscutum and mesoscutellum black. Frontal depression shallow, transverse carinae absent; submedian carina indicated by a set of irregular elevations or weak carinae. Hyperoccipital carina indicated by sharp carina. Occipital carina with a faintly indicated convex medial portion. Metascutellum concave, broad and short, projecting dorsally. Coxa not darker than rest of leg. T1 lateral carina expanded laterally. Metasomal flanges absent. Female: A3 shorter than pedicel. A4, A5 much broader than long. Mesoscutellum with obliquely raised sculpture. T1 midlobe with 4 longitudinal carinae. T6 without metasomal flanges. Fore wing long enough to reach or exceed metasomal apex. Male: A4 broader than long, A11 longer than broad. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to exceed metasomal apex. T7 much broader than long, blunt apically, without posterior projections.
Latin noun, genitive case, meaning “shields.”
[
Holotype, female: AUSTRALIA: QLD, 3.5km SSW Mount Baird,
Female. Body length 3.75–4.3 mm (n=21).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: shorter than pedicel. A4: longer than broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: with transverse ledge, face sharply receding below it. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: present as a single carina. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: transversely rugose. Extra carina ventral to occipital carina: present, medially incomplete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate punctate; absent. Major sculpture of gena posteroventrally: umbilicate foveate; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: absent; granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent; present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate; umbilicate punctate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T6. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: straight. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: longitudinally striate; umbilicate foveate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression shallow, transverse carinae absent; submedian carina indicated by a set of weak rugae. Hyperoccipital carina sharp and strong. Occipital carina incomplete (difficult to discern due to rough sculpture). Metascutellum broad and concave, truncate or slightly emarginate apically, projecting dorsally. Postmarginal vein present. Coxa not darker than rest of leg. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female: A3 much longer than pedicel. A4 about as long as broad, A5 broader than long. Mesoscutellum without granulate sculpture. T1 midlobe carinae obscured by raised smooth area. T6 without metasomal flanges, main body of tergum sloping down to apical rim. Fore wing long enough to reach base of T6. Male: Flagellomeres beyond A3 broader than long or only slightly longer than broad. T1 midlobe with 3 longitudinal carinae. Fore wing long enough to reach to or beyond T7. T7 truncate or very weakly emarginate.
Latin noun in apposition, meaning “the fence-sitter.”
[
Holotype, female: AUSTRALIA: QLD, Brisbane, DPI Indooroopilly site, 8.II.1971, malaise trap, OSUC 438936 (deposited in QMBA). Paratypes: AUSTRALIA: 20 females, 5 males, OSUC 438930, 438946 (ANIC); OSUC 227542, 462561 (CNCI); QDPC 0-165646 (QDPC); OSUC 438931-438935, 438937-438945, 438947-438949, 438951, 451325-451326 (WINC).
Female. Body length 2.25–3.15 mm (n=20).
Radicle color and shade: same as scape, both yellowish or reddish. Pedicel color: same as scape. A3: shorter than pedicel. A4: as long as broad. A5: broader than long.
Ventral clypeal margin: concave. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: granulate. Hyperoccipital carina: present as a single carina. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: smooth. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: granulate. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: smooth anteriorly, densely setose posteriorly. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate; longitudinally rugose. Major sculpture of mesoscutellum peripherally: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent. Postmarginal vein: absent. Fore wing apex at rest: reaching middle of T6. Coxae color brightness: same color as femora.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: strongly convex, tapering and sloping down to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression shallow; submedian carina absent or weakly indicated by a ruga. Hyperoccipital carina indicated by set of strong rugae. Occipital carina incomplete, nearly reaching hyperoccipital carina laterally. Metascutellum concave, emarginate apically, projecting dorsally. Postmarginal vein absent. Coxa not darker than rest of leg. T1 lateral carina expanded laterally. Metasomal flanges absent. Female: A4, A5 broader than long. Mesoscutellum with strong longitudinal rugae. T1 midlobe carinae obscured by raised smooth area. T6 without metasomal flanges, main body of tergum not abruptly separated from apical rim. Fore wing long enough to reach beyond metasomal apex. Male: A4, A11 as long or slightly longer than broad. Mesoscutellum with very strong rugose sculpture. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach beyond metasomal apex. T7 tiny, truncate.
[
Holotype, female, S. Shakespearei: AUSTRALIA: QLD, Gordonvale (Nelson), no date, QMBA HY3118 (deposited in QMBA). Other material: AUSTRALIA: 23 females, 29 males, OSUC 442355-442357, 442365, 442367-442372, 442375-442398 (ANIC); OSUC 227545-227547 (CNCI); OSUC 451327-451328 (QDPC); UCRC ENT 76364 (UCRC); OSUC 442373 (UQIC); OSUC 442358-442364, 442366, 442399-442401 (WINC).
Female. Body length 2.75–3.65 mm (n=6).
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: shorter than pedicel. A4: longer than broad. A5: broader than long.
Ventral clypeal margin: concave. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: present. Occipital carina: omicron-shaped, with sharp corners where median portion meets lateral portions. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally and ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T6. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: narrowly separated, angled anteriorly to become parallel. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: with median areole or pair of pits. Median propodeal carina: present.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 6. T1 lateral carina: straight. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep, transverse carinae very weak and interrupted; submedian carina strong. Hyperoccipital carina indicated by rugae. Occipital carina complete, laterally with strong corners and medially sinuate. Metascutellum exceedingly tiny, concave. Postmarginal vein present. Coxa not darker than rest of leg. Metasomal depression roughly sculptured. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female only: A4 longer than broad, A5 slightly broader than long. Mesoscutellum without granulate sculpture. T1 midlobe with 5 longitudinal carinae. Fore wing long enough to reach middle of middle or apex of T6. T6 broader than long.
Latin noun, genitive case, meaning “a curve.”
[
Holotype, female: AUSTRALIA: QLD, GS2, Mount Edith,
Female. Body length 4.1–4.95 mm (n=20).
Radicle color and shade: same as scape, both yellowish or reddish. Pedicel color: at least partially darker than scape. A3: longer than pedicel. A4: longer than broad. A5: longer than broad.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: with some transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: present. Occipital carina: present laterally, absent medially. Occiput sculpture: umbilicate foveate. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate. Microsculpture of gena anteroventrally: punctate. Microsculpture of gena posteroventrally: punctate.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate; transversely rugose. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a flattened or rounded elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate; umbilicate punctate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally and ventrally. Metascutellum shape: not emarginate, forming a flat, concave shelf. Metascutellar setae: absent. Metascutellum sculpture: with a median carina ending in a broad granulate patch posteriorly. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T5. Coxae color brightness: same color as femora.
T1 horn: present. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: straight. T2 sculpture: densely foveolate with granulate microsculpture, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: present. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: longer than broad. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: granulate. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression nearly flat, dorsally crossed by some weak arched carinae. Mesoscutum and mesoscutellum without granulate sculpture. Metascutellum flat with concave areas anteriorly and granulate sculpture posteriorly. Postmarginal vein present. Female: A4 longer than broad. T1 midlobe with broad horn, without longitudinal carinae. T2, T3 with sublateral longitudinal depressions. T6 not sharply pointed apically. Male: All flagellomeres longer than broad. Metasomal depression with antero-medial sculpture. T1 midlobe with 5 longitudinal carinae. T2 with slight longitudinal sublateral depressions. T7 with rounded posterior lobes.
[
Holotype, female, D. solitaria: AUSTRALIA: QLD, nr. Cairns, forest, Gordonvale (Nelson), 20.VI.1914, sweeping, A. A. Girault, SAMA DB 32-001591 (deposited in SAMA). Other material: AUSTRALIA: 30 females, 12 males, ANIC DB 32-020103, 32-020155, OSUC 367990, OSUC 367992, OSUC 368088, OSUC 368089, OSUC 368092, OSUC 368093, OSUC 368094, OSUC 368095, OSUC 368096, OSUC 368099, OSUC 368100, OSUC 368101, OSUC 368102, OSUC 368103, OSUC 368105 (ANIC); OSUC 227618-227619 (CNCI); OSUC 368106 (MVMA); OSUC 367991 (QDPC); OSUC 148481, 367989, 368104 (QMBA); UCRC ENT 121056 (UCRC); OSUC 368084-368087, 368097-368098, 451307-451317 (WINC).
Female. Body length 2.5–3.5 mm (n=20).
Radicle color and shade: same as scape, both yellowish or reddish. Pedicel color: same as scape. A3: shorter than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: concave. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: hood-like, dorsally protruding. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: omicron-shaped, with sharp corners where median portion meets lateral portions. Occiput sculpture: transversely rugose. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: absent; umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent; punctate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: deeply emarginate, with the resulting pair of posterior processes subtriangular and directed dorsally. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: same color as femora. Spines along tibiae: present. Lateral propodeal carinae: broadly separated, but parallel for a short distance anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as slightly protruding sharp corners. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: with medially truncate emargination, sloping down to apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Mesoscutum and mesoscutellum black. Frontal depression deep, transverse carinae absent; submedian carina strong and sharp. Hyperoccipital carina indicated by strong rugae. Occipital carina complete, sinuate medially. Metascutellum concave, nearly square, slightly emarginate apically, projecting dorsally. Coxa not darker than rest of leg. Tibiae with spines. T1 lateral carina expanded laterally. Metasomal flanges absent. Female: A3 shorter than pedicel. A4, A5 much broader than long. T1 midlobe with 4 longitudinal carinae. T6 without metasomal flanges, blunt or slightly concave apically. Fore wing long enough to reach middle of T6 or exceed metasomal apex. Male: A4, A11 broader than long. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to exceed metasomal apex. T7 with sharp, broad, flat posterior projections.
Latin noun in apposition, meaning “the dirty.”
[
Holotype, female: AUSTRALIA: QLD, via Taroom, FIT 035, Boggomoss No. 8,
Female. Body length 4.2 mm (n=1).
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: longer than pedicel. A4: longer than broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: with medially interrupted transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: umbilicate foveate. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: with shallow irregular carinae, posterodorsal corner with dense microsculpture. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate; transversely rugose. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a flattened or rounded elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate; irregularly rugose. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: granulate. Microsculpture of mesoscutellum peripherally: granulate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: not emarginate, forming a flat, concave shelf. Metascutellar setae: absent. Metascutellum sculpture: with a median carina ending in a broad granulate patch posteriorly. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: unknown. Anterior sculpture of metasomal depression: unknown. Median propodeal carina: unknown.
T1 horn: present. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: straight. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: longer than broad. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: granulate. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Female: A3 longer than pedicel. Submedian carina weakly indicated by a rounded carina. Hyperoccipital carina indicated by rugae; occipital carina incomplete medially. Mesoscutellum with weak granulate sculpture. Metascutellum spatulate, with a median carina that ends in a broad granulate patch. T1 with moderate anterior horn. T2 without longitudinal foveae or dense striae. T6 longer than broad.
Latin noun, genitive case, meaning “broad flat structure.”
[
Holotype, female: PAPUA NEW GUINEA: West New Britain Prov., New Britain Isl., primary forest edge, Cape Hoskins, 21.VI-27.VI.1973, malaise trap, Stibick, OSUC 368914 (deposited in CNCI).
Female. Body length 4.05–4.2 mm (n=2).
Radicle color and shade: darker than scape. Pedicel color: at least partially darker than scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: with medially interrupted transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: granulate. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: broadly angular, with rounded median peak. Occiput sculpture: umbilicate foveate. Extra carina ventral to occipital carina: present, medially incomplete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate; rugose. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: granulate. Median mesoscutal carina: absent. Major sculpture of mesoscutellum centrally: absent. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: with notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: reaching middle of T5. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: longitudinally striate; umbilicate foveate. Microsculpture of T6: absent. T6 medially: with medially truncate emargination, sloping down to apical rim. T6 metasomal flanges: present as slightly expanded lateral rims, truncate posteriorly. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep, without carinae ventrally, dorsal separator complete, some complete carinae present above dorsal separator; submedian carina very weak. Hyperoccipital carina absent. Occipital carina complete, medially sinuate. Mesoscutellar rim not expanded, with median notch. Metascutellum broad, weakly emarginate apically, projecting dorsally. Coxa darker than rest of leg. Postmarginal vein absent. Female: A4, A5 broader than long. T4, T5 without metasomal flanges. T6 with narrowly expanded lateral margins. Main surface of T6 weakly emarginate medially, not sloping down to apical rim. S6 not exposed to dorsal view.
Latin noun, genitive case, meaning “stubble.”
[
Holotype, female: AUSTRALIA: SA, Belair National Park, 4.II–11.II.1996, malaise trap, J. T. Jennings, OSUC 439590 (deposited in SAMA). Paratype: AUSTRALIA: 1 female, OSUC 148617 (SAMA).
Female. Body length 3.25–3.9 mm (n=6).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: concave. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: present. Occipital carina: omicron-shaped, with sharp corners where median portion meets lateral portions. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, medially incomplete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate punctate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent; granulate.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally and ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T6. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: narrowly separated, angled anteriorly to become parallel. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: with median areole or pair of pits. Median propodeal carina: present.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: straight. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: densely foveate, longitudinal sculpture irregular. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: granulate. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep, transverse carinae absent; submedian carina strong. Hyperoccipital carina indicated by rugae. Occipital carina complete, laterally with strong corners and medially broadly sinuate. Metascutellum short, concave, rugose. Postmarginal vein present. Coxa not darker than rest of leg. Metasomal depression with a pair of submedian foveae. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female only: A4 about as long as broad, A5 broader than long. Mesoscutellum without granulate sculpture. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach T6. T6 broader than long.
Named in honor of Danielle Stringer, for the large amount of excellent work that she contributed in packing and shipping specimens for this project.
[
Holotype, female: AUSTRALIA: QLD, Mount Moffatt Section, Carnarvon National Park, creek bed, Peawaddy Gorge,
Female. Body length 3.3–4 mm (n=20).
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: with medially interrupted transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: umbilicate foveate. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: absent; umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T6. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Body entirely dark brown, including antennae and legs. Frontal depression shallow, transverse carinae absent or broadly interrupted ventrally, absent dorsally; submedian carina absent medially. Hyperoccipital carina absent. Occipital carina complete, weakly convex or forming a rounded peak medially. Metascutellum deeply concave, emarginate apically, projecting dorsally. Postmarginal vein absent. T1 lateral carina expanded laterally. Metasomal flanges absent. Female: A3 longer than pedicel. A4 as long or slightly longer than broad, A5 nearly as long as broad. Mesoscutellum without granulate sculpture. T1 midlobe with 4 longitudinal carinae. T6 without metasomal flanges, main body of tergum not abruptly separated from apical rim. Fore wing long enough to reach middle of T6. Male: All flagellomeres longer than broad. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach to or beyond T7. T7 emarginate apically, with rounded posterior lobes.
Latin noun, meaning “thinness.”
[
Holotype, female: AUSTRALIA: TAS, Collinsvale, 12.II.1983, malaise trap, M. A. Williams, OSUC 438303 (deposited in ANIC). Paratypes: AUSTRALIA: 32 females, 54 males, Australian Museum K245265 (AMSA); ANIC DB 32-020902, 32-020903, OSUC 438282, OSUC 438283, OSUC 438284, OSUC 438285, OSUC 438286, OSUC 438287, OSUC 438288, OSUC 438289, OSUC 438290, OSUC 438291, OSUC 438292, OSUC 438293, OSUC 438294, OSUC 438295, OSUC 438296, OSUC 438297, OSUC 438298, OSUC 438299, OSUC 438300, OSUC 438301, OSUC 438311, OSUC 438312, OSUC 438313, OSUC 438314, OSUC 438315, OSUC 438316, OSUC 438317, OSUC 438318, OSUC 438319, OSUC 438320, OSUC 438321, OSUC 438322, OSUC 438324, OSUC 438325, OSUC 438326, OSUC 438327, OSUC 438328, OSUC 438329, OSUC 438330, OSUC 438331, OSUC 438332, OSUC 438333, OSUC 438334, OSUC 438335, OSUC 438336, OSUC 438337, OSUC 438338, OSUC 438339, OSUC 438340, OSUC 438341, OSUC 438343, OSUC 448574, OSUC 448575, OSUC 448576, OSUC 448577, OSUC 448578, OSUC 448579, OSUC 448580, OSUC 448581, OSUC 448582, OSUC 448583, OSUC 448584, OSUC 448585, OSUC 448586, OSUC 453952, OSUC 453953, OSUC 453954, OSUC 453955, OSUC 453956 (ANIC); OSUC 438345-438346 (MVMA); OSUC 438304-438310, 438323, 448587 (TDAH); OSUC 438302, 438344, 453957 (WINC).
While
Female. Body length 2.95–4 mm (n=20).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long; as long as broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: absent. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: granulate. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: broadly angular, with rounded median peak. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, medially incomplete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: reaching middle of T5. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as strong posterior corners. T6: broader than long. Major sculpture of T6: umbilicate punctate; longitudinally striate. Microsculpture of T6: absent. T6 medially: with medially truncate emargination, sloping down to apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep, carinae interrupted medially, no carinae above dorsal separator; submedian carina absent or weak and irregular. Hyperoccipital carina indicated by rugae. Occipital carina complete, sinuate medially. Mesoscutellar rim not expanded, without median notch. Metascutellum scoop-shaped, deeply concave, slightly emarginate apically, projecting dorsally. Coxa darker than rest of leg. T1 lateral carina expanded laterally. Female: A3 longer than pedicel. A4 as long or longer than broad, A5, A6 broader than long. T1 midlobe with 4 longitudinal midlobe carinae anteriorly or with dense sculpture obscuring them. Fore wing long enough to reach middle of T5. T4, T5 without metasomal flanges. Main body of T6 raised above apical rim and abruptly separated from it; lateral margins of T6 slightly expanded, forming a truncate or slightly emarginate shelf-like apex.
Latin noun, genitive case, meaning “a truncation.”
[
Holotype, female: AUSTRALIA: QLD, 15km NE Mareeba, 20.XII–7.I.1985, Storey & Titmarsh, OSUC 429933 (deposited in QMBA). Paratypes: AUSTRALIA: 46 females, ANIC DB 32-020082, 32-020153, OSUC 359643, OSUC 429944, OSUC 429945, OSUC 429946, OSUC 429950, OSUC 429952, OSUC 429956, OSUC 429957, OSUC 429958, OSUC 429959, OSUC 429960, OSUC 429961, OSUC 453995 (ANIC); OSUC 148377, 429948-429949, 429955, QM Reg. No. T35151 (QMBA); OSUC 449090 (UQIC); OSUC 429922-429932, 429934-429943, 429947, 429951, 429953, 453994 (WINC).
Female. Body length 4.35–4.8 mm (n=2).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: longer than pedicel. A4: longer than broad. A5: longer than broad.
Ventral clypeal margin: concave. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: with transverse ledge, face sharply receding below it. Transverse curved rugae extending from frontal depression to eye: present. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: present as multiple regular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: smooth; transversely rugose; umbilicate punctate. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate punctate. Major sculpture of gena posteroventrally: umbilicate punctate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate punctate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; irregularly rugose. Microsculpture of mesoscutal midlobe anteriorly: absent; granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent. Major sculpture of mesoscutellum centrally: umbilicate punctate. Major sculpture of mesoscutellum peripherally: umbilicate punctate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with a median carina, otherwise weakly sculptured. Postmarginal vein: present. Fore wing apex at rest: reaching middle of T5. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: present. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: straight. T2 sculpture: densely foveolate, longitudinal sculpture irregular. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: longer than broad. Major sculpture of T6: umbilicate punctate; longitudinally striate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression nearly flat, surrounded by many transverse curved rugae; submedian carina indicated by a set of rugae. Hyperoccipital carina indicated by many transverse carinae. Occipital carina incomplete, lateral portions approaching hyperoccipital carina. Occiput mostly smooth, with many rugae dorsally, otherwise smooth and with setiferous puncta. Metascutellum broad and very short, concave. Postmarginal vein present. Coxa and rest of leg bicolored. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female: A3 longer than pedicel. A4, A5 longer than broad. Mesoscutellum with setiferous puncta. T1 midlobe with strong anterior horn, no complete longitudinal carinae. Fore wing long enough to reach to middle of T5. T6 longer than broad. Male: A4, A11 longer than broad. Mesoscutellum without extensive granulate sculpture. Fore wing long enough to reach apex of T5. T7 tiny, truncate or very weakly emarginate.
Latin noun, genitive case, meaning “tube.”
[
Holotype, female: NEW CALEDONIA: Nord Prov., 5km WSW Pouebo, Mount Mandjelia,
Female. Body length 2.4–3.4 mm (n=20).
Radicle color and shade: same as scape, both yellowish or reddish. Pedicel color: same as scape; at least partially darker than scape. A3: shorter than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; transversely rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: uniformly rounded dorsally. Occiput sculpture: umbilicate foveate. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate punctate; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: with shallow irregular carinae, posterodorsal corner with dense microsculpture. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: deeply emarginate, with the resulting pair of posterior processes subtriangular and directed dorsally. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: exceeding metasomal apex; reaching middle of T6. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 6. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as strong posterior corners. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: with broad emargination between protruding posterolateral corners, separated from apical rim. T6 metasomal flanges: present as spine-like structures posterolaterally. T6 raised peripheral rim: absent. S4 sculpture: densely setose, setal pits between very weak longitudinal rugae. S5 sculpture: densely setose, setal pits between very weak longitudinal rugae. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: exposed to dorsal view by T6 emargination. S6 apex: rounded or acuminate.
Mesoscutum and mesoscutellum black. Frontal depression shallow, transverse carinae absent; submedian carina absent or incomplete. Hyperoccipital carina indicated by strong rugae. Occipital carina complete, convex medially. Metascutellum deeply concave, projecting dorsally. T1 lateral carina expanded laterally. Female: A4, A5 broader than long. T1 midlobe with 6 or more longitudinal carinae, or these obscured by a smooth elevation. T5 with strong but rounded posterior corners. T6 abruptly narrower than T5, with sharp protruding posterior corners and a steep median slope leading to apical rim. Fore wing long enough to reach middle of T6 or beyond metasomal apex. Main body of T6 not abruptly separated from apical rim. Male: A3 not longer than pedicel. All flagellomeres between A4 and A12 broader than long. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to exceed metasomal apex. T6 with strong posterior corners. T7 abruptly narrower than T6, with sharp, protruding posterior corners.
Latin noun, genitive case, meaning “a bump.”
[
Holotype, female: AUSTRALIA: QLD, Heathlands,
Female. Body length 3.15–3.75 mm (n=17).
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: shorter than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present only as a weak shift in elevation. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: omicron-shaped, with sharp corners where median portion meets lateral portions. Occiput sculpture: transversely rugose. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: with large smooth areas. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: absent. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: with slight, indistinct sculpture dorsally, smooth ventrally. Metascutellum shape: deeply emarginate, with the resulting pair of posterior processes subtriangular and directed dorsally. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, but parallel for a short distance anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: with median areole or pair of pits. Median propodeal carina: present.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: with medially truncate emargination, sloping down to apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Body entirely dark brown, including antennae and legs. Mesoscutum and mesoscutellum black. Frontal depression shallow, transverse carinae absent; submedian carina indicated by weak rugae. Hyperoccipital carina indicated by rugae. Occipital carina complete, omicron-shaped medially. Metascutellum deeply concave, emarginate apically, projecting dorsally. Tibiae with flattened spines. T1 lateral carina expanded laterally. Metasomal sterna with longitudinal rugae. Female: A3 shorter than pedicel. A4, A5 broader than long. T1 midlobe with 4 longitudinal carinae. T6 without metasomal flanges. Fore wing long enough to reach beyond metasomal apex. Main body of T6 not abruptly separated from apical rim. Male: A3 longer than pedicel. A4 broader than long, A11 longer than broad. T1 midlobe with 4 longitudinal carinae that may be obscured by raised sculpture. Fore wing long enough to exceed metasomal apex. T7 with elongate spine-like posterior projections.
Latin noun, genitive case, meaning “barbs.”
[
Collected on
Holotype, female: AUSTRALIA: ACT, Honeysuckle Creek,
Female. Body length 2.75–3.35 mm (n=4).
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: longer than pedicel. A4: longer than broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: absent. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: granulate. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: uniformly rounded dorsally. Occiput sculpture: umbilicate foveate; transversely rugose. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; umbilicate punctate. Microsculpture of gena anteroventrally: granulate. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: with large smooth areas. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: punctate. Microsculpture of mesoscutellum peripherally: punctate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: with slight, indistinct sculpture dorsally, smooth ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: reaching near apex of T5; reaching middle of T6. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: with median areole or pair of pits. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: obscured by other raised sculpture. T1 lateral carina: straight. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression shallow, transverse carinae absent; submedian carina absent medially. Genal carina expanded, with large foveae between it and gena laterally. Hyperoccipital carina indicated by rugae. Occipital carina complete, convex. Epomial corner slightly protruding. Mesoscutum with raised longitudinal smooth area postero-medially. Metascutellum broad, deeply concave, slightly emarginate apically. Postmarginal vein present. Entire leg dark brown. Metasomal depression with irregular sculpture. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female: A3 longer than pedicel. A4 longer than broad, A5 broader than long. Mesoscutellum without granulate sculpture, densely foveate medially. T1 midlobe carinae obscured by raised area. Fore wing long enough to reach apex of T5 or middle of T6. Male: Fore wing long enough to reach beyond metasomal apex. T7 truncate apically, without apical protrusions.
Latin noun, genitive case, meaning “strong chain.”
[
Holotype, female: AUSTRALIA: WA, 5km SE Nornalup, coastal heathlands, Walpole-Nornalup National Park, 17.XII-18.XII.1990, pan trap, A. D. Austin, OSUC 448998 (deposited in WAMP). Paratypes: AUSTRALIA: 3 females, 1 male, OSUC 462579-462580 (CNCI); OSUC 448999-449000 (WINC).
Female. Body length 3.4–3.5 mm (n=4).
Radicle color and shade: darker than scape. Pedicel color: same as scape. A3: shorter than pedicel; as long as pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: with medially interrupted transverse carinae. Dorsal portion of frontal depression: with medially interrupted transverse carinae. Submedian carina: present. Frontal depression dorsally: hood-like, dorsally protruding. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: absent. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: with shallow irregular carinae, posterodorsal corner with dense microsculpture. Posterior border of central pronotal area: directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: with large smooth areas. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: absent; umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: punctate. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: not emarginate, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent; but marginal vein curving slightly at apex. Fore wing apex at rest: reaching base of T5. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: present as slightly protruding sharp corners. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: present as strongly protruding acuminate flanges. T6: broader than long. Major sculpture of T6: longitudinally striate; umbilicate foveate. Microsculpture of T6: absent. T6 medially: slightly emarginate, not separated from apical rim. T6 metasomal flanges: present as slightly expanded lateral rims, rounded posteriorly. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: present, posteriorly complete. S6 apex in relation to T6: exposed to dorsal view by T6 emargination. S6 apex: rounded or acuminate.
Both sexes: Frontal depression deep, without carinae ventrally, dorsal separator interrupted medially, one or more interrupted carinae present above dorsal separator; submedian carina weakly defined or absent medially. Hyperoccipital carina absent. Occipital carina complete, medially weakly convex. Mesoscutellar rim not expanded, without median notch. Metascutellum tiny, projecting dorsally. Coxa darker than rest of leg. Postmarginal vein absent. Female: A3 longer than pedicel. A4, A5 broader than long. T4, T5 without distinct metasomal flanges. Main surface of T6 with slightly expanded lateral margins, truncate or very slightly emarginate apically, but sharply separated from apical rim. S6 exposed to dorsal view, rounded apically.
Latin noun, genitive case, meaning “a covering.”
[
Holotype, female: AUSTRALIA: SA, coastal dunes, 15km SSW Streaky Bay, 10.XI–15.XI.1987, pan trap, A. D. Austin, OSUC 436034 (deposited in SAMA). Paratypes: AUSTRALIA: 4 females, OSUC 436030-436033 (WINC).
Female. Body length 3.5–4 mm (n=11).
Radicle color and shade: same as scape, both dark brown. Pedicel color: at least partially darker than scape. A3: longer than pedicel. A4: longer than broad; as long as broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: with some transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: present as a single carina. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate; umbilicate punctate. Major sculpture of gena posteroventrally: umbilicate foveate; umbilicate punctate. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: smooth anteriorly, densely setose posteriorly. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: transversely rugose; umbilicate punctate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent. Major sculpture of mesoscutellum centrally: umbilicate foveate; obliquely rugose. Major sculpture of mesoscutellum peripherally: umbilicate punctate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: deeply emarginate, with the resulting pair of posterior processes subtriangular and directed dorsally. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: absent. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, but parallel for a short distance anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: anteriorly longitudinally rugose with setal pits spanning interspaces, posteromedially sparsely foveate. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: densely foveate, longitudinal sculpture irregular. T4 metasomal flanges: absent. T5 sculpture: densely foveate, longitudinal sculpture irregular. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: smooth, with tiny umbilicate foveae. S5 sculpture: smooth, with tiny umbilicate foveae. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression small and shallow, transverse carinae absent; submedian carina sharp. Hyperoccipital carina indicated by a set of sharp carinae. Occipital carina complete or incomplete, sometimes gradually becoming indistinguishable medially; occiput with many transverse rugae. Mesoscutum and sometimes mesoscutellum with deeply sunken midline, causing surrounding areas to apparently protrude. Metascutellum broad and deeply concave, laterally with only a narrow longitudinally striate area, weakly emarginate or truncate apically, projecting dorsally. Postmarginal vein absent or extremely short. T1 lateral carina not expanded laterally. Female: A3 longer than pedicel. A4 longer than broad, A5 nearly as long as broad. T1 midlobe with 4 longitudinal carinae that may be obscured by a smooth raised area. Fore wing long enough to reach middle of T6. T6 broader than long. Male: All flagellomeres past A3 about as long as broad. T1 midlobe with 5 longitudinal carinae (submedian pair weak). Fore wing long enough to reach beyond T7. T7 tiny, truncate or weakly convex.
Latin noun, genitive case, meaning “wart.”
[
Holotype, female: AUSTRALIA: NSW, Point Lookout, New England National Park,
Female. Body length 3.4–3.7 mm (n=2).
Radicle color and shade: same as scape, both yellowish or reddish. Pedicel color: at least partially darker than scape. A3: longer than pedicel. A4: longer than broad. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: absent. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate. Upper frons microsculpture: absent. Hyperoccipital carina: absent. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: present laterally, absent medially. Occiput sculpture: smooth. Extra carina ventral to occipital carina: absent. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate; absent. Major sculpture of gena posteroventrally: umbilicate punctate; absent. Microsculpture of gena anteroventrally: absent. Microsculpture of gena posteroventrally: absent.
Lateral pronotal area sculpture: anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: not steep, forming less than a right angle. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate. Microsculpture of mesoscutal midlobe anteriorly: absent. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: absent. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate; umbilicate punctate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally, smooth ventrally. Metascutellum shape: deeply emarginate, with the resulting pair of posterior processes subtriangular and directed dorsally. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: same color as femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: with median areole or pair of pits. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 5. T1 lateral carina: straight. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: mostly smooth, with tiny umbilicate foveae. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: absent. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression nearly flat; submedian carina absent. Hyperoccipital carina absent. Occipital carina incomplete, lateral portions not approaching hyperoccipital carina. Occiput umbilicate punctate. Metascutellum broad and deeply emarginate. Postmarginal vein present. Coxa same color as rest of leg. T1 lateral carina not expanded laterally. Metasomal flanges absent. Female: A3 longer than pedicel. Mesoscutellum with small umbilicate foveae or pits. T1 midlobe without anterior horn, with 5 longitudinal carinae. Fore wing long enough to reach beyond metasomal apex. T6 broader than long.
Latin noun in apposition, meaning “traveller.”
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Holotype, female: VANUATU: Sanma Prov., Espiritu Santo Isl., 900A ground / moist lowland forest, MG09A2, Penaoru,
Female. Body length 3.2–3.55 mm (n=10).
Radicle color and shade: same as scape, both dark brown. Pedicel color: same as scape. A3: longer than pedicel. A4: broader than long. A5: broader than long.
Ventral clypeal margin: with slightly convex median lobe. Interantennal process: not elongate. Lower frons at dorsal margin of interantennal process: without transverse carina. Transverse curved rugae extending from frontal depression to eye: absent. Median longitudinal carina in frontal depression: absent. Ventral portion of frontal depression: smooth. Dorsal portion of frontal depression: without transverse carinae. Submedian carina: present. Frontal depression dorsally: not hood-like, open dorsally. Upper frons major sculpture: umbilicate foveate; irregularly rugose. Upper frons microsculpture: absent. Hyperoccipital carina: indicated by a set of irregular elevations. Carina connecting occipital carina to hyperoccipital carina: absent. Occipital carina: weakly arched dorsally, with rounded lateral corners. Occiput sculpture: irregularly sculptured. Extra carina ventral to occipital carina: present, complete. Gena length: shorter than eye. Major sculpture of gena anteroventrally: umbilicate foveate. Major sculpture of gena posteroventrally: umbilicate foveate; rugose. Microsculpture of gena anteroventrally: granulate. Microsculpture of gena posteroventrally: granulate.
Lateral pronotal area sculpture: densely covered with setiferous puncta. Posterior border of central pronotal area: directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum. Mesoscutum anteriorly: very steep and tall, descending at a right angle or protruding anteriorly. Major sculpture of mesoscutal midlobe anteriorly: umbilicate foveate; longitudinally rugose. Mesoscutal midlobe sculpture at midlength: not different from nearby sculpture. Major sculpture of mesoscutal midlobe posteriorly: umbilicate foveate; longitudinally rugose. Microsculpture of mesoscutal midlobe anteriorly: granulate. Microsculpture of mesoscutal midlobe posteriorly: absent. Median mesoscutal carina: present as a vague, occasionally interrupted elevation. Major sculpture of mesoscutellum centrally: umbilicate foveate. Major sculpture of mesoscutellum peripherally: umbilicate foveate. Microsculpture of mesoscutellum centrally: absent. Microsculpture of mesoscutellum peripherally: absent. Mesoscutellar rim: not expanded. Mesoscutellar rim medially: without notch. Mesofemoral depression: longitudinally striate dorsally and ventrally. Metascutellum shape: slightly emarginate posteriorly, concave but elevated posteriorly. Metascutellar setae: absent. Metascutellum sculpture: with large smooth posterior fovea. Postmarginal vein: present. Fore wing apex at rest: exceeding metasomal apex. Coxae color brightness: darker than femora. Spines along tibiae: absent. Lateral propodeal carinae: broadly separated, not parallel anteriorly. Setae in metasomal depression: absent. Anterior sculpture of metasomal depression: absent. Median propodeal carina: absent.
T1 horn: absent. Number of longitudinal carinae of T1 midlobe: 4. T1 lateral carina: protruding laterally, visible from ventral view. T2 sculpture: with longitudinal striae or rugae, setiferous puncta present between them. T2 sublateral longitudinal foveae: absent. T3 metasomal flanges: absent. T4 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T4 metasomal flanges: absent. T5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. T5 metasomal flanges: absent. T6: broader than long. Major sculpture of T6: umbilicate punctate; umbilicate foveate. Microsculpture of T6: absent. T6 medially: flat and tapering to a rounded apex, not separated from apical rim. T6 metasomal flanges: absent. T6 raised peripheral rim: absent. S4 sculpture: longitudinally striate or rugose, setal pits spanning interspaces. S5 sculpture: longitudinally striate to rugose, setal pits spanning interspaces. S5 median carina: present. S6 peripheral carina: absent. S6 apex in relation to T6: not exposed to dorsal view. S6 apex: rounded or acuminate.
Both sexes: Frontal depression with broadly interrupted transverse carinae, including some above dorsal separator; submedian carina weakly indicated but complete. Hyperoccipital carina indicated by weak rugae. Occipital carina complete, weakly convex medially. Metascutellum deeply concave, emarginate apically, projecting dorsally. Postmarginal vein present, short. Coxa darker than rest of leg. T1 lateral carina expanded laterally. Metasomal flanges absent. Female: A4, A5 much broader than long. Mesoscutellum without granulate sculpture. Fore wing long enough to reach middle of T6 or beyond metasomal apex. T1 midlobe with 4 longitudinal carinae. T6 without metasomal flanges, main body of tergum not separated from apical rim, strongly sloping apically but not concave. Male: A4 broader than long, A11 slightly longer than broad. T1 midlobe with 4 longitudinal carinae. Fore wing long enough to reach beyond metasomal apex. T7 tiny, truncate apically.
Arbitrary combination of letters, does not change spelling to agree with gender.
[
Holotype, female: AUSTRALIA: WA, Mount Cooke, 17.II-18.IV.1991, malaise trap, M. S. Harvey & J. M. Waldock, OSUC 449005 (deposited in WAMP). Paratypes: AUSTRALIA: 9 females, 7 males, OSUC 449006-449021 (WINC).
We thank L. Musetti, D. Stringer, J. Jennings, J. Cora, A. Tindall, A. Valerio, S. Hemly, S. Mantel, and other members of the PBI team, for assistance with specimens, photographs, and databasing. This work was funded by the National Science Foundation under grant No. DEB–0614764 to N.F. Johnson and A.D. Austin, and by the Australian Biological Resources Study grant 208-058 to A.D. Austin.
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Taxonomic records for all records used in the present paper. (doi:
Locality records for all records used in the present paper. (doi:
Characters. = used in phylogenetic analysis.
Radicle color and shade
same as scape, both yellowish or reddish
darker than scape
same as scape, both dark brown
Pedicel color
same as scape
at least partially darker than scape
A3 [female]
longer than pedicel
shorter than pedicel
as long as pedicel
A4 [female]
longer than broad
broader than long
as long as broad
A5 [female]
longer than broad
broader than long
as long as broad
Ventral clypeal margin
concave
uniformly convex
with slightly convex median lobe
with elongate median lobe
Interantennal process
not elongate
elongate
Lower frons at dorsal margin of interantennal process
without transverse carina
with transverse ledge, face sharply receding below it
with oblique carina extending towards mouth corner
Transverse curved rugae extending from frontal depression to eye
absent
present
Median longitudinal carina in frontal depression
absent
present
Ventral portion of frontal depression
with transverse carinae
with medially interrupted transverse carinae
with medially interrupted transverse foveae
smooth
Dorsal portion of frontal depression
without transverse carinae
with some transverse carinae
with medially interrupted transverse carinae
Submedian carina
present
absent
present only as a weak shift in elevation
Frontal depression dorsally
not hood-like, open dorsally
hood-like, dorsally protruding
Upper frons major sculpture
umbilicate foveate
irregularly rugose
transversely rugose
Upper frons microsculpture
absent
granulate
punctate
Hyperoccipital carina
absent
present as a single carina
indicated by a set of irregular elevations
present as multiple regular elevations
Carina connecting occipital carina to hyperoccipital carina
absent
present
Occipital carina
uniformly rounded dorsally
weakly arched dorsally, with rounded lateral corners
weakly arched dorsally, with sharp lateral corners
broadly angular, with rounded median peak
omicron-shaped, with sharp corners where median portion meets lateral portions
present laterally, absent medially
Occiput sculpture
smooth
umbilicate foveate medially, becoming smooth laterally
umbilicate foveate
transversely rugose
irregularly sculptured
umbilicate punctate
Extra carina ventral to occipital carina
absent
present, complete
present, medially incomplete
Gena length
shorter than eye
as long as eye
Major sculpture of gena anteroventrally
umbilicate foveate
rugose
umbilicate punctate
absent
Major sculpture of gena posteroventrally
umbilicate foveate
rugose
umbilicate punctate
absent
Microsculpture of gena anteroventrally
absent
granulate
punctate
Microsculpture of gena posteroventrally
absent
granulate
punctate
Lateral pronotal area sculpture
with shallow irregular carinae, without microsculpture
irregularly foveate, with smooth area dorsally
with shallow irregular carinae, posterodorsal corner with dense microsculpture
with a series of arched carinae, posterodorsal corner with weak longitudinal rugae
anteriorly smooth, posterodorsal corner with dense microsculpture, ventral corner with irregular carinae
densely covered with setiferous puncta
irregularly sculptured
smooth anteriorly, densely setose posteriorly
mostly granulate, ventral corner with irregular carinae
Posterior border of central pronotal area
directed anteriorly, protruding at corner of epomial carina and transverse pronotal carina
directed posteriorly, epomial carina absent or meeting transverse pronotal carina at arch on lateral surface of pronotum
Mesoscutum anteriorly
very steep and tall, descending at a right angle or protruding anteriorly
not steep, forming less than a right angle
Major sculpture of mesoscutal midlobe anteriorly [female]
umbilicate foveate
longitudinally rugose
transversely rugose
umbilicate punctate
irregularly rugose
Mesoscutal midlobe sculpture at midlength [female]
not different from nearby sculpture
with large smooth areas
with narrow curved smooth elevations
Major sculpture of mesoscutal midlobe posteriorly [female]
umbilicate foveate
longitudinally rugose
obliquely rugose
umbilicate punctate
irregularly rugose
transversely rugose
Microsculpture of mesoscutal midlobe anteriorly [female]
absent
granulate
Microsculpture of mesoscutal midlobe posteriorly [female]
absent
granulate
punctate
Median mesoscutal carina
present as a narrow carina
absent
present as a flattened or rounded elevation
present as a vague, occasionally interrupted elevation
present as a ruga
Major sculpture of mesoscutellum centrally [female]
absent
umbilicate foveate
longitudinally rugose
obliquely rugose
umbilicate punctate
irregularly rugose
Major sculpture of mesoscutellum peripherally [female]
umbilicate foveate
longitudinally rugose
obliquely rugose
umbilicate punctate
irregularly rugose
Microsculpture of mesoscutellum centrally [female]
absent
granulate
punctate
Microsculpture of mesoscutellum peripherally [female]
absent
granulate
punctate
Mesoscutellar rim
not expanded
expanded
Mesoscutellar rim medially
without notch
with notch
Mesofemoral depression
longitudinally striate dorsally and ventrally
longitudinally striate dorsally, smooth ventrally
with slight, indistinct sculpture dorsally, smooth ventrally
smooth
Metascutellum shape
deeply emarginate, with the resulting pair of posterior processes subtriangular and directed dorsally
not emarginate, forming a flat, concave shelf
not emarginate, concave but elevated posteriorly
slightly emarginate posteriorly, concave but elevated posteriorly
not emarginate, convex dorsally
weakly emarginate apically, not concave dorsally
Metascutellar setae
absent
present
Metascutellum sculpture [female]
with large smooth posterior fovea
with a median carina ending in a broad granulate patch posteriorly
with a longitudinal median ruga and some weak transverse carinae
with three parallel and evenly spaced longitudinal carinae
with a median carina, otherwise weakly sculptured
with many longitudinal rugae
dense and irregular
densely umbilicate foveate
Postmarginal vein
absent
present
but marginal vein curving slightly at apex
Forewing apex position at rest [female]
reaching middle of T5
reaching near apex of T5
reaching base of T5
not reaching base of T5
exceeding metasomal apex
reaching middle of T6
Coxae
same color as femora
darker than femora
Spines along tibiae
absent
present
Lateral propodeal carinae
broadly separated, not parallel anteriorly
narrowly separated, subparallel anteriorly
meeting near propodeal midlength
broadly separated, subparallel anteriorly
Setae in metasomal depression
present
absent
Anterior sculpture of metasomal depression
absent
with median areole or pair of pits
with series of longitudinal carinae extending to lateral propodeal carinae
Median propodeal carina
absent
present
T1 horn [female]
absent
present
Number of longitudinal carinae of T1 midlobe [female]
3
4
5
6
obscured by other raised sculpture
more than 6
T1 lateral carina
straight
protruding laterally, visible from ventral view
T2 sculpture
densely foveate, longitudinal sculpture irregular
with longitudinal striae or rugae, setiferous puncta present between them
anteriorly longitudinally rugose with setal pits spanning interspaces, posteromedially sparsely foveate
densely foveate with granulate microsculpture, longitudinal sculpture irregular
mostly smooth with setiferous puncta medially, laterally with some longitudinal carinae
T2 sublateral longitudinal foveae [female]
absent
present
T3 metasomal flanges [female]
absent
present
T4 sculpture
longitudinally striate to rugose, setal pits spanning interspaces
densely foveate, longitudinal sculpture irregular
mostly smooth, with tiny umbilicate foveae
T4 metasomal flanges [female]
absent
present as slightly protruding sharp corners
present as strongly protruding flanges
T5 sculpture
longitudinally striate to rugose, setal pits spanning interspaces
densely foveate, interspaces not raised
densely foveate, longitudinal sculpture irregular
mostly smooth, with tiny umbilicate foveae
T5 metasomal flanges [female]
absent
present as slightly protruding acuminate flanges
present, blade-like with rounded margins
present, rounded and lobe-like
present as strongly protruding acuminate flanges
present as strong posterior corners
T6 shape [female]
longer than broad
broader than long
Major sculpture of T6 [female]
umbilicate punctate
longitudinally striate
umbilicate foveate
Microsculpture of T6 [female]
absent
granulate
T6 medially [female]
with medially truncate emargination, separated from apical rim
with deep emargination that is V-shaped medially, separated from apical rim
slightly emarginate, not separated from apical rim
with medially truncate emargination, sloping down to apical rim
with broad emargination between protruding posterolateral corners, separated from apical rim
with a median projection set off by an abrupt narrowing posterior to tiny metasomal flanges, not separated from apical rim
flat and tapering to a rounded apex, not separated from apical rim
tapering to a sharp point, not separated from apical rim
strongly convex, tapering and sloping down to a rounded apex, not separated from apical rim
truncate, separated from apical rim
T6 metasomal flanges [female]
absent
present as tiny apical sharp projections
present, very broadly rounded but with sharp apices
present, very broadly rounded, with rounded apices
present as slightly expanded lateral rims, rounded posteriorly
present as slightly expanded lateral rims, truncate posteriorly
present, broad and elongate, with slight lateral incisions near midlength
present as spine-like structures posterolaterally
present subapically
T6 raised peripheral rim [female]
present
absent
S4 sculpture
longitudinally striate or rugose, setal pits spanning interspaces
densely setose, setal pits between very weak longitudinal rugae
sparsely foveolate, with tiny pits in interspaces
smooth, with tiny umbilicate foveae
S5 sculpture
longitudinally striate to rugose, setal pits spanning interspaces
densely setose, setal pits between very weak longitudinal rugae
sparsely foveate, with tiny pits in interspaces
smooth, with tiny umbilicate foveae
S5 median carina
present
absent
S6 peripheral carina [female]
present, posteriorly complete
absent
S6 apex in relation to T6 [female]
exposed to dorsal view by T6 emargination
not exposed to dorsal view
S6 apex [female]
truncate
rounded or acuminate
A3 [male]
longer than pedicel
shorter than pedicel
as long as pedicel
A5 tyloid shape
narrow, linear
expanded, ovate or sinuate
A6 [male]
longer than broad
broader than long
A11 [male]
longer than broad
broader than long
as long as broad
Major sculpture of mesoscutal midlobe anteriorly [male]
umbilicate foveate
irregularly rugose
transversely rugose
umbilicate punctate
Major sculpture of mesoscutal midlobe posteriorly [male]
umbilicate foveate
irregularly rugose
transversely rugose
umbilicate punctate
obliquely rugose
longitudinally rugose
Microsculpture of mesoscutal midlobe anteriorly [male]
absent
granulate
Microsculpture of mesoscutal midlobe posteriorly [male]
absent
granulate
Major sculpture of mesoscutellum centrally [male]
umbilicate foveate
longitudinally rugose
umbilicate punctate
absent
irregularly rugose
Major sculpture of mesoscutellum peripherally [male]
umbilicate foveate
longitudinally rugose
umbilicate punctate
irregularly rugose
Microsculpture of mesoscutellum centrally [male]
absent
granulate
punctate
Microsculpture of mesoscutellum peripherally [male]
absent
granulate
punctate
Forewing apex position at rest [male]
reaching middle of T5
reaching apex of T5
exceeding metasomal apex
reaching middle of T6
T1 midlobe longitudinal carinae [male]
3
4
5
6 or more
obscured by other raised sculpture
T3 metasomal flanges [male]
absent
present
T4 metasomal flanges [male]
absent
present
T5 metasomal flanges [male]
absent
present as slightly protruding acuminate flanges
present, rounded and lobe-like
present as sharp corners that do not protrude
present as strongly protruding acuminate flanges
T6 metasomal flanges [male]
absent
present as slightly protruding acuminate flanges
present, rounded and lobe-like
present as sharp corners that do not protrude
present as strongly protruding acuminate flanges
present as subapical tubercles
T7 [male]
with a pair of sharply defined spine-like posterolateral projections
broadly and deeply emarginate, with rounded posterolateral margins
M-shaped, with a triangular median emargination
weakly emarginate
truncate
broadly emarginate, with sharply pointed posterolateral lobes