Diagrams of the mouthparts of adult horned beetles. A from an oblique anterior side of a Trypoxylus dichotomus male B from an oblique anterior side of a T. dichotomus female; names of mouthparts were omitted C View from an oblique ventral side of T. dichotomus male D diagram of two types of adult mandible tips; left, the tip of the left mandible of a T. dichotomus male from the left side; this tip does not branch off; right, the tip of a left mandible of a Dynastes hercules male from the left side; this tip branches off and is forked.