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Article title
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Abstract
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Keywords
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Introduction
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Materials and methods
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Insects
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Observations of the mandible muscles
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Measurements of mandible strengths and resistance to breakage
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Paraffin surface scratching tests with horned beetle heads
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Video recording and mandible gear-like structures filing procedures
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Results
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Bark-carving behaviors and mandibles
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Gear-like structures on mandibles
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Measurements of mandible break-resistance strengths
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Discussion
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Acknowledgments
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References
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Supplementary materials
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