Corresponding author: Fenglong Jia (
Academic editor: Martin Fikácek
A revised checklist of
The studies of the material of the
The present paper is based predominantly on 600 specimens of the family
Details on distribution are partly adopted from
Holotype ♂, China, Guizhou, Guiyang, 6.x.1940, lgt. Zhe-long Pu (translation, labeled in Chinese) (SYSU). Paratypes (2 exs.): 1 ♂, same data as holotype (SYSU); 1 ♂, same data as holotype (NMW).
Length 2.9–3.0 mm, width 1.6–1.7 mm. Body oval, tapering backwards, widest before the middle (
Head. Dark brown, somewhat lighter between eyes, anterior margin of clypeus densely punctured, but with much stronger and sparser punctures between eyes. Labrum yellowish brown with dark spot in the middle. Distance between eyes 1.6× width of one eye. Antennae light yellowish brown, not darkened apically. Palpi yellowish brown.
Pronotum. Yellow to yellowish brown. Without basal plicae, strongly and densely punctured. Lateral sides margined, straight to slightly convex. Base a little narrower than elytra at base.
Elytra. Yellowish brown, with dark interrupted lines on primary punctures rows, darkened along suture, with vague dark marks connecting primary puncture rows, without dark band basally. Completely margined. Primary puncture rows moderately strong and dense, 38–39 punctures in row 1. Secondary punctures moderately strong and dense along suture, moderately strong and much sparse on intervals. All punctures darkened.
Ventral side. Brown red, with legs and anterior 1/fifth of prosternal process yellow brown; elytral epipleura yellowish brown with strong darkened punctures, reaching to abdominal sternite 6. Prosternal process narrowed between coxae, grooved along each side, anterior edge not margined, with moderately strong punctation. Metaventral process slightly bulbous with a row of strong punctures on each side that is slightly impressed, else moderately punctured (
Males. Pro- and mesotarsomeres moderately widened and provided with suction-pads. Mesotarsomere 1 not very strongly excised. Penis and parameres as
Female. Unknown.
The species is named in honour of Pierre André Latreille (1762–1833), a French entomologist who firstly used
This species is close to
Only known from the type locality.
Only valid names are given in this list. For complete synonymy and distribution outside China see
Widely distributed western and north-eastern Palaearctic species reaching the utmost north-west of China: Xinjiang. New to China.
Endemic to China: Sichuan.
Endemic to China: Tianjin.
Western and north-eastern Palaearctic species reaching China in the north-east: Heilongjiang, Inner Mongolia. New record for Inner Mongolia.
Endemic to China: Hubei, Hunan, Shaanxi. New records for Hunan and Shaanxi.
Additional material examined by Vondel:
Eastern Palaearctic species, known from several provinces in the east and south of China: Beijing, Chongqing, Guizhou, Hunan, Jiangsu, Shanghai, Sichuan, Yunnan, Zhejiang. New records for Sichuan and Zhejiang.
See type material in Systematics chapter.
Endemic to China: Guizhou.
Additional material examined by Vondel:
Endemic and widespread in the south-eastern part of China: Anhui, Fujian, Guangdong, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Shandong, Shaanxi, Taiwan, Yunnan, Zhejiang. New records for Guangxi, Hubei, Shaanxi and Yunnan.
A widespread western Palaearctic species reaching China in the utmost north-western part: Xinjiang. New to China.
Additional material examined by Vondel:
A widely spread western and north-eastern Palaearctic species, reaching the west of China: Qinghai, Xinjiang. New to China.
Additional material examined by Vondel:
Eastern Palaearctic species, known from several provinces in the eastern part of China: Anhui, Beijing, Guangdong, Heilongjiang, Inner Mongolia, Jiangsu, Jilin, Liaoning, Shaanxi, Shandong, Zhejiang. New records for Anhui, Inner Mongolia and Jilin.
North-eastern Palaearctic species, known from the north of China: Gansu, Heilongjiang, Inner Mongolia, Qinghai. New records for Gansu, Inner Mongolia and Qinghai.
The specimen from Kuerle was identified as
Southern Palaearctic reaching the western part of China: Xinjiang.
Additional material examined by Vondel: 1 ex. Djalantun St., 6.v.1939. lgt. P. Brinck [Butha Qi, Inner Mongolia] (CN)
North-eastern Palaearctic species, reaching China in the north-east: Heilongjiang, Inner Mongolia, Jilin, Liaoning.
Additional material examined by Vondel:
The older records of
Endemic to China, known from several provinces from west to east: Beijing, Fujian, Guizhou, Inner Mongolia, Jiangsu, Shandong, Shanghai, Sichuan, Shanxi, Xinjiang, Yunnan, Zhejiang.
Eastern Palaearctic and Oriental species, known from the north and south of China: Beijing, Heilongjiang, Tianjin, Yunnan. New record for Tianjin.
Additional material examined by Vondel:
Eastern Palaearctic and Oriental species, widely spread in the eastern part of China: Anhui, Beijing, Fujian, Guizhou, Hainan, Heilongjiang, Hong Kong, Hubei, Hunan, Jiangsu, Liaoning, Shaanxi, Shandong, Shanghai, Taiwan, Tianjin, Yunnan. New records for Anhui and Shaanxi.
Additional material examined by Vondel:
Endemic species in the south of China: Guizhou, Yunnan. New record for Guizhou.
Additional material examined by Vondel:
Eastern Palaearctic and Oriental species, known from the west, east and south-east of China: Beijing, Fujian, Guangdong, Guizhou, Hunan, Jiangsu, Liaoning, , Shanghai, Sichuan, Xinjiang, Yunnan, Zhejiang. New record for Yunnan.
Endemic to China: Yunnan.
Eastern Palaearctic/Oriental species, known from the Ryukyu Islands (Japan) and Taiwan.
Oriental species, reaching China in the south: Fujian, Guangxi. New record for Guangxi.
(examined by Vondel):
Eastern Palaearctic species, known from several provinces in the east and south of China: Anhui, Chongqing, Fujian, Guizhou, Liaoning, Shanghai, Taiwan, Yunnan.
Widely spread Palaearctic species, reaching to China in the utmost north-west: Xinjiang.
Oriental species, reaching to China in the south-east: Guangdong, Guangxi, Hainan. New record for Guangxi.
Additional material examined by Vondel:
North-eastern Palaearctic species, reaching to China in the north-east: Heilongjiang, Inner Mongolia, Liaoning.
Eastern Palaearctic species, known from the south-east of China: Beijing, Fujian, Guangdong, Shanghai, Sichuan, Taiwan, Zhejiang. New records for Sichuan and Guangdong.
Eastern Palaearctic species, known from the most-eastern part of Russia and several provinces in the east of China: Beijing, Fujian, Guangdong, Hebei, Liaoning, Shaanxi, Shandong, Tianjin. New records for Guangdong, Shaanxi and Tianjin.
Additional material examined by Vondel:
Eastern Palaearctic and Oriental species, widely spread in the eastern and south-eastern part of China: Anhui, Beijing, Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hebei, Henan, Hubei, Hunan, Jilin, Jiangsu, Jiangxi, Liaoning, Shaanxi, Shandong, Shanghai, Sichuan, Taiwan, Yunnan, Zhejiang. New records for Chongqing, Hubei and Shaanxi.
We are grateful to Miss Shuang Zhao for her help with preparing photographs. This work was supported by “Biodiversity of Mount Jinggangshan in China” (2010330007102993).