Concatenated mitochondrial cox1 and nuclear 28S genes phylogeny. Araneus bonali sp. n. underlined in green within the Holartic Araneus sequences available at GenBank or BOLD (accession codes shown in Table 3). The red-coloured frame shows the clade corresponding to the Araneus diadematus group. Tree topology was inferred using Bayesian inference analysis (GTR + I + Gamma substitution model).