Corresponding author: Liang Tang (
Academic editor: Volker Assing
Chinese species of
The members of
Up to the present, 59 species of
Specimens examined in this paper were all collected near streams through forests and killed with ethyl acetate. For examination of male genitalia, the last three abdominal segments were detached from the body after softening in hot water. The aedeagus together with other dissected pieces were mounted in Euparal (Chroma Geselschaft Schmidt, Koengen, Germany) on plastic slides. Photos of sexual characters were taken with Cannon G7 attached to Olympus SZX 16 stereoscope; habitus photos were taken with a Cannon macro photo lens MP-E 65mm attached to Cannon EOS40D camera.
The type specimens treated in this study are deposited in the following public and private collections:
Department of Biology, Shanghai Normal University, P. R. China
private collection of V. Puthz, Schlitz, Germany
private collection of G.-M. de Rougemont, London, England
The measurements of proportions are abbreviated as follows:
body length, measured from the anterior margin of the clypeus to the posterior margin of 10th abdominal tergite
forebody length, measured from the anterior margin of the clypeus to the apex of the elytra (apicolateral angle)
width of head including eyes
width of pronotum
width of elytra
length of pronotum
length of elytra, measured from humeral angle
1 | Pronotum bicolorous with golden bands along the anterior and posterior margins | 2 |
– | Pronotum unicolorous | 3 |
2 | Head distinctly broader than elytra; femora bicolorous. Habitus ( |
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– | Head narrower than elytra; femora unicolorous. Habitus ( |
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3 | Body, at least head and pronotum, metallic green or golden green | 4 |
– | Body, at least head and pronotum, metallic blue or black with plumbeous lustre | 6 |
4 | Elytra relatively narrow, head about as broad as elytra; punctures on head and pronotum moderate in size and distinctly separated, interstices can be as broad as half the diameter of a puncture. Habitus ( |
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– | Elytra relatively broad, head distinctly narrower than elytra; punctures on head and pronotum very coarse and very dense, interstices narrow and sharp | 5 |
5 | Frons between eyes sharply inclined inward forming a deep and broad concavity; punctures on elytra mostly distinctly delimited; paratergites of abdominal tergite 4 broad, slightly declivous. Habitus ( |
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– | Frons between eyes gently inclined inward forming a shallow and broad concavity, traces of two lateral longitudinal furrows can be recognized at posterior portion of the concavity; punctures on elytra mostly transversely or diagonally confluent; paratergites of abdominal tergite 4 narrow, slightly reflexed. Habitus ( |
|
6 | Forebody distinctly metallic blue; femora bicolorous | 7 |
– | Forebody black with plumbeous lustre, sometimes elytra with brassy reflection; femora unicolor | 8 |
7 | punctures on frons deep and dense; posterior half of elytra with vorticose sculpture. Habitus ( |
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– | punctures on frons shallow and sparse; posterior half of elytra with transverse sculpture. Habitus ( |
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8 | Punctation of pronotum and elytra coarser and less confluent; posteromedian part of 7th male sternite ( |
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– | Punctation of pronotum and elytra smaller and confluent; posteromedian part of 7th male sternite ( |
Adult habitus of
China (Yunnan, Hunan), Vietnam, Thailand, Borneo, Indonesia.
Body entirely black, head, pronotum, elytra and basal abdominal tergites with a blue metallic lustre. Antennae blackish brown, with club segments lighter. First two segments of maxillary palpi brownish yellow, last segment brown. Legs blackish with tibiae and tarsi slightly lighter, femora yellowish in basal third.
BL: 4.5–4.9mm; FL: 2.5–2.6 mm.
Proportions of holotype: HW: 63.5, PW: 47, PL: 53, EW: 61, EL: 67.
Head 1.04 times as wide as elytra; interocular area gently inclined inward forming a shallow and broad concavity; punctures round, distinctly delimited, slightly larger on median area than near dorsal margins of eyes, diameter of large punctures about as wide as widest cross section of 2nd antennal segment, interstices smooth, mostly smaller than half diameter of punctures. Antennae when reflexed exceeding posterior margin of pronotum; length of segments from base to apex: 10.0: 7.0: 17.5: 11.0: 9.0: 8.0: 8.0: 7.0: 7.0: 8.0: 10.0.
Pronotum 1.28 times as long as wide, widest slightly before middle and constricted at base; punctures round, partially slightly confluent, distinctly larger than those on frons, interstices smooth, mostly smaller than half diameter of punctures.
Elytra nearly rectangular; punctation on average slightly coarser than that of pronotum, punctures on humeral area mostly distinctly delimited, and those on inner 2/3 portion of elytra (especially those on posterior half) obliquely confluent, interstices similar to those on pronotum.
Length of metatarsi from base to apex: 11.5: 8.5: 5.5: 3.5: 10.5.
Abdomen subcylindrical; 3rd to 6th segments with broad and densely punctate paratergites, paratergites of tergite 4 narrower than greatest width of hind tibia; 7th tergite with an apical membranous fringe; punctures on 3rd tergite as large as one eye facet, interstices smooth.
Male. Seventh sternite (
Female. Sternite 8 (
China (Hainan).
The new species is similar to
China (Yunnan), Myanmar, Thailand.
Body entirely black with a faint plumbeous lustre, elytra sometimes with brassy reflection. Antennae blackish brown. Maxillary palpi with first segment yellowish, second segment light brown and last segment brown. Legs black with a brownish tint, tibiae and tarsi slightly lighter.
BL: 3.7–4.4 mm; FL: 2.1–2.3mm.
Proportions of holotype: HW: 58.0, PW: 44.5, PL: 49.0, EW: 59.0, EL: 63.5.
Head about as wide as elytra; lateral portions of front slightly rising, medial portion concave; punctures round, distinctly delimited, slightly larger on median area than near dorsal margins of eyes, diameter of large punctures about as wide as apical cross section of 3rd antennal segment, interstices smooth, smaller than or as broad as half diameter of punctures. Antennae when reflexed extending to the posterior margin of pronotum; Length of segments from base to apex: 9.0: 6.5: 9.0: 8.0: 8.0: 5.5: 8.0: 6.0: 7.0: 6.5: 9.0.
Pronotum 1.10 times as long as wide, widest slightly before middle and constricted at base; punctures partially confluent, diameter of large punctures about as wide as apical cross section of 2nd antennal segment, interstices smooth, mostly smaller than or about as broad as half diameter of punctures.
Elytra nearly rectangular; punctation similar to that of the pronotum, punctures on humeral area mostly distinctly delimited, those on medial two thirds obliquely confluent, interstices similar to those on pronotum.
Length of metatarsi from base to apex as 18.5: 8.0: 5.5: 4.0: 11.5.
Abdomen subcylindrical; 3rd to 6th segments with broad and densely punctate paratergites, paratergites of 4th tergite as broad as greatest width of hind tibia; 7th tergite with an apical membranous fringe; punctures on 3rd tergite slightly smaller than one eye facet, interstices with very indistinct microsculpture.
Male. Seventh sternite (
Female. Eighth sternite (
Adult habitus of
China (Jiangxi, Guizhou)
In a few specimens the punctation of pronotum and elytra is strongly confluent as in
The new species slightly resembles
Body entirely black with a faint plumbeous lustre. Antennae blackish brown, antennal club slightly lighter than preceding segments. Maxillary palpi brownish. Legs black with a brownish tint, tibiae and tarsi slightly lighter.
BL: 4.6–5.0mm; FL: 2.6–2.7mm.
Proportions of holotype: HW: 59.5, PW: 44.0, PL: 50.5, EW: 66.0, EL: 69.5.
Head 0.9 times as wide as elytra; lateral portions of frons slightly raised, median portion concave; punctures round to elliptic, distinctly delimited, slightly larger on median area than near dorsal margins of eyes, diameter of largest punctures about as wide as basal cross section of 2nd antennal segment, interstices smooth, smaller than or as broad as half diameter of punctures. Antennae when reflexed extending to the posterior margin of pronotum; Length of segments from base to apex as 9.5: 6.5: 14.5: 8.5: 7.5: 6.5: 7.0: 5.5: 6.0: 5.5: 8.0.
Pronotum 1.15 times as long as wide, widest slightly before middle and constricted at base; punctures partially confluent, similar in size to those on head, interstices similar to those on frons.
Elytra nearly rectangular; punctation similar to that of the pronotum, punctures on humeral area mostly distinctly delimited, those on posterior half of elytra strongly confluent, forming a narrowly vorticose sculpture.
Relative length of segments of hind legs from base to apex as 15.0: 8.5: 5.5: 3.5: 14.5.
Abdomen subcylindrical; 3rd to 6th segments with broad and densely punctate paratergites, paratergites on 4th segment as broad as largest width of hind tibia; 7th tergite with an apical membranous fringe; punctures on 3rd tergite distinctly smaller than eye facet, interstices smooth.
Male. Seventh sternite (
Female. Eighth sternite (
China (Yunnan).
The new species is similar to
China (Xizang), Nepal.
The species was originally described from Nepal and not unsurprisingly was found in China near the border.
Adult habitus of
China (Guizhou), Myanmar, Thailand.
No Chinese material was examined by us; a photograph of a paratype (cPut) from Myanmar is provided here.
China (Yunnan).
This species was only known from the female holotype, which is actually deposited in “the collection of the Laboratory of Entomology, Tokyo University of Agriculture”, not in “Shanghai Institute of Entomology, Academia Sinica” (Present name: Shanghai Entomology Museum, the Chinese Academy of Science) as original published paper described.
China (Xizang), India, Nepal, Bhutan.
No Chinese material was examined by us, and a photograph of specimen (cPut) from Nepal is provided here.
We would like to express our sincere gratitude to dear Dr. V. Puthz (Germany) for his kind guidance on our study, to Mr. G.-M. de Rougemont (England) for the kind loan of specimens and other help, to Mr. Xian-Wei Liu (Shanghai), Dr. Y. Watanabe (Japan) and Dr. S. Nomura (Japan) for helping us to search for the holotype of