Two new species of Pselaphodes Westwood and new record of Taiwanophodes minor Hlaváč from South China (Coleoptera, Staphylinidae, Pselaphinae)

Abstract Two new species, Pselaphodes linae Yin & Li, sp. n. (Hainan, Fujian) and Pselaphodes shii Yin & Li, sp. n. (Hainan) are described from South China. Taiwanophodes minor Hlaváč is reported from outside Taiwan for the first time. Illustrations of major diagnostic features are provided for all treated taxa. The latest key to Chinese Pselaphodes is modified to include the new species.


Introduction
A number of tyrine specimens have been submitted for determination since the publication of our previous papers of the genus Pselaphodes Westwood (Yin et al. 2010(Yin et al. , 2011(Yin et al. , 2012. Among this material were two new Pselaphodes species, and two males and three females of Taiwanophodes minor Hlaváč, a species previously known only from Taiwan. This information is reported herein.

Methods
The terminology of foveal system follows Chandler (2001), except for using 'ventrite' instead of 'sternite' when concerning meso-and metathoracic structures.
A slash (/) is used to separate lines on the same label, and a double slash (//) is used to separate different labels on the same pin.
Measurements are in millimeter, the following acronyms are used in the text: ALlength of the abdomen along the midline; AW-maximum width of the abdomen; BLlength of the body (= HL + PL + EL + AL); EL-length of the elytra along the sutural line; EW-maximum width of the elytra; HL-length of the head from the anterior clypeal margin to the occipital constriction; HW-width of the head across eyes; PLlength of the pronotum along the midline; PW-maximum width of the pronotum.
The type series are deposited in the Insect Collection of Shanghai Normal University, Shanghai, China (SNUC). Diagnosis. Reddish brown; medium-sized; genae rounded; antennomeres IX-XI strongly modified; pronotum with rounded lateral margins; with thin and long metaventral processes (in lateral view); pro-and metatibiae sinuate; aedeagus with asymmetric median lobe and small parameres.

Pselaphodes linae
Description. Male (Fig. 1). Length 2.65-2.87. Head as long as wide, HL 0.56-0.58, HW 0.56-0.58; eyes large, each composed of about 40 facets. Antennal clubs as in Fig  Distribution. South China: Hainan, Fujian. Comparative notes. Very distinctive species, easily separable from all other Pselaphodes species by the sinuate pro-and metatibiae, the metatibiae with a blade-shaped ventral tooth near apical 1/3, and the serrate ventral margin for the distance posterior to the tooth. The new species shares with P. bomiensis Yin et al., 2011 andP. condylus Yin et al., 2011 a similar placement of the antennal modification, but can be readily separated from both by the pronotal lateral margins being rounded. On the contrary the lateral margins of the pronotum are more or less angularly expanded laterally in P. bomiensis and P. condylus.
Etymology. The species name is dedicated to Mei-Ying Lin for the collection of part of the type series. Diagnosis. Reddish brown; medium-sized; postocular margins rounded; antennomeres IX strongly modified; pronotum angularly expanded laterally; with robust and long metaventral processes (in lateral view); aedeagus with asymmetric median lobe and long parameres.
Female. Unknown. Distribution. South China: Hainan. Comparative notes. This species is placed close to P. bomiensis, P. condylus and P. jizushanus Yin et al., 2011 by sharing a similar placement of the antennal modification and projected anterolateral margins of the pronotum. It can be readily separated from P. bomiensis and P. condylus by the much smaller size and different form of the antennomeres IX, from P. jizushanus by the metaventral processes being slightly curved posteriorly, the metatrochanters being protuberant ventrally and the metatibiae being simple. Pselaphodes jizushanus has the metaventral processes being curved anteriorly, the metatrochanters are simple, and the metatibiae are bluntly projected at apical 1/5.
Etymology. The species is named after Hong-Liang Shi for the collection of the holotype.

Identification key
The latest key to the Chinese Pselaphodes (Yin et al. 2012: 36) is modified as the following to include the two new species described in the present paper.
Female. Similar to male in general; each eye composed of about 30 facets; lacking antennal modification; lacking metaventral process; apices of protibiae simple.
Distribution. South China: Taiwan, Hainan (New Province Record). Comparative notes. Readily distinguished from the only other congener T. magnus Bekchiev, 2010 by the smaller size, antennomeres X being less modified, pronotum being angularly expanded anterolaterally, profemora and protibiae being simple, different conformation of the metaventral processes, and aedeagus with more elongate median lobe and reduced parameres.
Comments. Taiwanophodes minor was originally described from central Taiwan (Hlaváč, 2002). It is somewhat surprising that this species also occurs in the Hainan Island, suggesting geographic affinities between the two islands.