The possible distribution of Leistus punctatissimus during the last ice age maximum, (red triangles). Green triangles: northern “sentinels” of Duvalius breiti, a relict of the “Massifs de refuge”. The blue lines indicate the approximate height of the alpine “inlandsis” of the Dolomites around the Rolle Pass. Reconstructed after the map of Pellegrini et al., 2004. The Pale di San Martino massif represents one of the largest nunatak area near the Rolle Pass.