Corresponding author: Jiri Hulcr (
Academic editor: A.I. Cognato
Following the recent reclassification of the Palaeotropic xyleborine genera (
The following species are synonymized:
The following species are given new status:
List of abbreviations
BMNH Natural History Museum, London
FRI Forestry Research Institute, Dehra Dun, India
MCg Museo Civico Genova, Genova
MNB Museum fur Naturkunde der Humboldt University, Berlin, Germany
MSUC Michigan State University Arthropod Collection, East Lansing, MI, USA
NHMW Natürhistorisches Museum, Wien, Austria
RAB Roger A. Beaver’s private collection, Chiang Mai, Thailand
SMTD Staatliches Museum fur Tierkunde, Dresden, Germany
UCD Bohart Museum, University of California-Davis, CA, USA
USNM United States National Museum, Smithsonian Institution, Washington, D.C., USA.
Indonesia, Java (
Lectotype of
Indonesia, Java, Buitenzorg (
1.5 mm.
Type specimen of
Malaysia, Sabah, Danum Valley (2, R.A. Beaver det., MSUC); Sumatra (USNM).
Prolonged large representative of
Malaysia, Burma (2 indiv., BMNH).
Very similar to
Malaysia, Sarawak (holotype, BMNH); Malaysia, Sabah, Danum Valley (9 indiv., Hulcr det., MSUC.).
Elytral declivity slightly with much higher number of declivital tubercles than other
Elytral declivity superficially different from other
Indonesia, Kalimantan (2, holotype & allotype, NSMT); Indonesia, Kalimantan (2 paratypes, NHMW).
Elytral apex not emarginate, but all other diagnostic characters of
Holotype of
Reported from India and Thailand, and by
Malaysia, Sarawak, Mt. Penrissen, 4500 ft. (lectotype, NHMW).
All diagnostic features of genus
Indonesia, Borneo (lectotype, NHMW).
All diagnostic features of
Lectotype of
Malaysia (BMNH); Malaysia, Sabah, Danum Valley (3, Hulcr det.).
An “elegant” form of
Holotypes of
India, Sunderbans Div. (holotype, BMNH).
All diagnostic features of
Australia, Queensland, Geagana (lectotype, NHMW).
All diagnostic features of
Malaysia, Selangor (lectotype, BMNH); Malaysia, Selangor (Schedl det., BMNH); Malaysia, Sabah, Danum Valley, (13, Hulcr det., MSUC).
Very similar to
Creates irregularly branching galleries with transverse brood chambers (Beaver and Browne 1978).
Holotype of
Malaysia, Peninsula (holotype, NHMW).
Diagnostic characters of
Type species of
Indonesia, Java, Buitenzorg (holotype, NHMW).
Type of
Uganda (
The only known African
Described as
Malaysia, Perak (holotype, BMNH).
Transferred to
Malaysia, N.S. Triang (homotype, NHMW); Malaysia, Perak (BMNH); Malaysia, Borneo (BMNH).
Type in Hamburg museum lost (
Lectotype of
My utmost thanks belong to Dr. S. L. Wood for creating a classification of scolytine beetles upon which all current and future bark beetle taxonomists can build. I thank Dr. A. I. Cognato for helping organize this series of publications, and for his support of my work. The work was sponsored by USDA-APHIS grant, National Science Foundation (PEET DEB-0328920), National Geographic Society (7922–05), and the Ministry of Education fo the Czech Republic(MSM 60076605801). I am indebted to curators of the museums who readily loaned specimens for examination: M. Barclay (BMNH), R. Poggi (MCG), J. Frisch and J. Willers (MNB), H. Schoenmann (NHMW), and O. Jaeger (SMTD). S. Heydon and L. Kimsey kindly supported my extended work in the UCD collection. I thank R. A. Beaver for specimens and unceasing advice.