Description of a new species of Coelosis Hope from Guajira Peninsula, northern Colombia (Coleoptera, Scarabaeidae, Dynastinae, Oryctini)

Abstract A new species of Coelosis is described from the Macuira Mountains, Guajira Peninsula, northern Colombia. A character comparison between this new and other previously known Colombian species in the genus is presented. A key for the identification and distributional map for Colombian species of Coelosis is provided, as well as a key for the genera included in the tribe Oryctini in Colombia.


Introduction
The genus Coelosis is a Neotropical dynastine group of mainly nocturnal forest beetles that can be collected at lights (Endrödi 1985, Iannuzzi and Marinoni 1996, Ratcliffe 2003. The larvae of Coelosis biloba (L.) is the only one described in the genus and is known to be associated with the nests of the leaf-cutter ant Atta cephalotes (L.) (Hymenoptera: Formicidae) (Pardo-Locarno et al. 2006).
The purpose of this work is to describe a new species of Coelosis from a tropical dry forest in the protected area of Macuira, Guajira Pensinsula, to provide new biological data and distribution records for other species Coelosis, and to provide diagnostic characters and illustrations for all three Colombian species in the genus Coelosis as well as a key to the genera of Oryctini from Colombia.

Materials and methods
Internal and external morphological characters were studied using a dissecting microscope (6.5-40.0×). For measurements, an ocular micrometer was used. Internal sclerotized structures were dissected after relaxing the specimen in hot (75 °C) water. Heavily sclerotized parts were soaked in a 15 % solution of potassium hydroxide and neutralized in a 15 % solution of acetic acid. Genitalia were card-mounted or placed in a glycerin-filled vial beneath the specimen.
Specimens were characterized using body length, puncture density, setation, and color as described in Orozco (2012).

Material examined
138 specimens were reviewed and label data were obtained from the following Colombian collections except where noted otherwise (curators in brackets):  Mandibular teeth widely separated at base (Fig. 2H). Males and females with cephalic horns (Fig. 1A)
Allotype (Fig. 3G). Female. Similar to holotype except for the following: body length 22.4 mm; width 12.0 mm, frons and disc of pronotum smooth, without fovea or horns, pygidium slightly concave in lateral view, and genital plate as Fig. 5L.
Etymology. The specific epithet wayuorum refers to the Wayuu indigenous group inhabiting the Guajira Peninsula.
Temporal distribution. Three specimens collected in September 2014, during the dry season.
Life history. The type material was collected at night with lights during the dry season.

The following are new biological data and distribution records for Coelosis species in Colombia
Coelosis bicornis.