Corresponding author: Jiří Skuhrovec (
Academic editor: M. Thomas
The new species,
Skuhrovec J, Alonso-Zarazaga MA (2017) Revision of the genus
The phylogenetic and taxonomic position of hyperines is still unresolved. In the previous study, hyperines together with
Their only unique feature appears to be the specific meshed cocoon spun by the larvae from strands of protein secreted by the Malpighian tubules (
Only the Palaearctic fauna of
The genus
In this paper,
Label data are cited in the description, separate lines on labels are indicated by “/” and separate labels by “//”.
Body 2.5–4.6 mm; entire body densely covered with appressed scales of different shapes, from scales divided into two lobes to base up to entire scales. Eyes elliptical to oval. Rostrum long to very long, narrow; in dorsal view distinctly longer than its base width (ratio more than 3.00); enlarged anteriorly, tapered to basal third part and afterward almost parallel-sided; in side view slightly curved; as long as pronotum (ratio = 0.95–1.10). Antenna with 6 or 7 desmomeres. Pronotum distinctly wider than long, widest at middle. Elytra with very distinct prominent humeri. Apex of penis enlarged, sometimes partially to the tip, and always without projecting setae. Apodeme of sternite VIII in females relatively long, with distinct long lateral arms; plate wide, not very well sclerotized, upper part not connected and bearing apically many distinct setae.
These weevils occur in warm and dry habitats (calcareous hillsides, vineland, steppe, sandy habitats, meadows, clearings), and in mesophilic or moderately damp habitats of floodplains and hillsides (natural meadows) (
The genus
Altos de San Juan near El Escorial (Spain, limit between the provinces of Madrid and Ávila,
Holotype ♂: ‘Escorial / Puerto [printed label] // Altos de / San Juan [handwritten label] // Altos de S. Juan / debajo de pequeñas / piedras con terreno / de esta composición [translation: Altos de S[an] Juan (S. = San), under small stones with a ground of this composition. i.e., a stony or gravelly ground] [handwritten label]’ (
(Figs
This species is absolute unique not only in this genus, but also in whole tribe
Morphological detail of
The new species is named after a close friend of the authors, Herbert Winkelmann (Berlin, Germany), who mentored the first author in
Unfortunately, the host plant or any other biological data is not known. Weevils were collected probably at the beginning of the 20th century. We know only an exact locality of this weevil: Alto San Juan near Escorial – ca 35 km NW from Madrid, which is located in the mountains, 1734 m a.s.l. All known
Central Spain (provinces Madrid and Zaragoza).
1 | Desmomeres 7 (Fig. |
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– | Desmomeres 6 (Fig. |
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2 | Elytral scales entire, not divided in two lobes (Fig. |
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– | Elytral scales divided in two lobes apically, the emargination reaching at least midlength of each scale (Fig. |
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3 | Elytra only with a few projecting setae. (Fig. |
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– | Elytra with numerous projecting setae (Figs |
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4 | Pronotum widest behind midlength, near to base; lateral stripe of scales on each margin yellow (Fig. |
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– | Pronotum widest at midlength, lateral stripe of scales on each margin white (Fig. |
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Dorsal and lateral view of
Dorsal and lateral view of
Whereas identification of the species is, in contrast to the majority of other
The character of the number of desmomeres has high variability also within different genera in many weevil groups (e.g.
Taxonomic positions and relatives of genera, subgenera and species-groups within the tribe
The study of Jiří Skuhrovec was supported by a grant from the Ministry of Agriculture (Mze ČR) RO0416 and a SYNTHESYS (ES-TAF-3511). Miguel A. Alonso-Zarazaga was partly supported by project CGL2015‐66571‐P (MINECO/FEDER) (Ministerio de Economía y Competitividad, Spain). Special thanks to Jon Cooter (Oxford, UK) for linguistic help on the manuscript.