Omoglymmius (s. str.) wukong sp. n., a new species from Xizang, China (Coleoptera, Rhysodidae, Omoglymmiini)

Abstract Omoglymmius (s. str.) wukong sp. n. (Coleoptera: Rhysodidae: Omoglymmiini) is described from Xizang, China. Relevant morphological characters of the new species are illustrated with colour plates, and known distribution of the subgenus Omoglymmius in the Himalayan region is mapped.


Introduction
Omoglymmius Ganglbauer, 1891 is the most speciose genus of Rhysodidae (Coleoptera), almost cosmopolitan, but absent from Madagascar, New Zealand, and South America. Bell and Bell (1982) excellently revised Omoglymmius and established eleven subgenera to classify the congeneric species. The nominotypical subgenus is the largest with 97 species (Lorenz 2005, Bell and Bell 2009, Hovorka 2015. However, in the fauna of East Asia, only two species in the subgenus Omoglymmius had been recorded before this study, namely O. (s. str.) sakuraii (Nakane, 1973) (China (Taiwan), Japan, Vietnam) and O. (s. str.) laticeps Bell, 1977 (Bhutan, India). In this paper, a new species, O. (s. str.) wukong sp. n., is described and illustrated from Xizang Autonomous Region, China. The new species is compared to the two related species, with some selected and important morphological characters presented in a table.

Materials and methods
Specimens were relaxed and softened in a hot saturated solution of potassium hydroxide for 4 minutes (for mounted dry specimens) or 8 minutes (for alcohol-preserved specimens), and then transferred to distilled water to rinse the residual potassium hydroxide off and stop any further bleaching. The softened specimens were placed in glycerine and dissected to observe morphological details. After examination, the body parts were mounted on a glass slide with Euparal Mounting Medium for future studies. Habitus photographs were taken using a Canon macro photo lens MP-E 65mm on a Canon 550D. Observations, photographs, and measurements of morphological details were performed using a Zeiss Axio Zoom.V16 motorized stereo zoom microscope with a Zeiss AxioCam MRc 5. Photographs in Figure 6 were taken with an Olympus BX53 microscope with an Olympus DP73 camera. The final deep focus images were created with Zerene Stacker 1.04 stacking software. Adobe Photoshop CS6 was used for final processing. Precise label data are cited, while authors' remarks and addenda are placed in square brackets; separate label lines are indicated by a slash (/), and separate labels are indicated by a double slash (//). Measurements are averages taken from five specimens. The morphological terminology follows Bell (1978, 1982). Rhysodid beetles are treated as an independent family, following the publications of Bell (2003), Bousquet (2012), andMakarov (2008).
The material examined for this study is deposited in the following collections and museums (with names of curators in parentheses):

BITS
Bin Insect Taxonomy  Antennal length length between the antennal base and the apex.

Body length
length between the mandibular apex (mandibles closed) and the elytral apex.

Elytral length
length between the basal border of elytra and the apex along suture.

Elytral width
widest part of both elytra combined.

Eye length
length of a single compound eye in lateral view.

Eye width
width of a single compound eye in lateral view.

Head length
length between the anterior apex of clypeus and the posterior margin of temporal lobe along the midline.

Genus
Habitus ( Fig. 2A-B) elongate, rather narrow, lustrous. Body colour mostly blackish brown to black; antennae and legs somewhat reddish brown; mouthparts reddish brown to yellowish brown.
Head ( Fig. 3A-C) broad, as wide as long. Median lobe short, broad, subtruncate at tip. Frontal space short, nearly V-shaped, margins only shallowly sinuate. Temporal lobes longer than wide; medial angles rounded, contiguous; posteriomedial margin evenly rounded into posteriolateral margin; occipital angle scarcely evident; orbital groove impressed, extended before or near the middle of eye, following one or two separate coarse dorsal punctures far away from posterior margin of temporal lobe (red arrow in Fig. 3A); remainder of temporal lobe smooth except micropunctures; temporal setae absent; postorbital tubercle minute, not pilose, appearing as a slight convexity in lateral view. Eye entire, curvilinearly triangular, length/width = 1.1. Mentum surface coarsely and continuously punctate, with many setae. Antenna (Fig. 3D) without stylet; antennomeres V-X with minor setae in form of subapical rings; basal setae absent; all antennomeres impunctate. Pronotum (Fig. 3E) subelliptical, distinctly narrowed anteriorly and posteriorly, widest at about basal 4/9, length/width = 1.1-1.2. Sides gently curved, hardly sinuate before hind angle; hind angles broadly rounded. Carinae subequal at middle; outer carina with base distinctly narrowed, with medial margin sinuate before base and with a distinct oblique microgroove at about basal 1/4 of its length; inner carina distinctly narrowed in basal part, gradually narrowed in apical part, and weakly undulated at Elytra (Figs 2A; 3F) elongate, narrow, length/width = 2.2-2.3. Striae impressed, coarsely punctate, punctures relatively small and deep; intervals only slightly convex; stria IV with one seta at about basal 2/9, one seta at about apical 2/7 of its length and one seta subapically; subapical striole with one seta; stria VII with four setae near apex (some specimens with one seta behind the insertion level of hind leg). Metathoracic wings fully developed.
Protibia (Fig. 4A) nearly cylindrical, not swollen at middle; profemur with a large and somewhat rounded tooth at medial position of ventral side. Mesotibia (Fig. 4A) with one curved spur and one minute calcar. Metatibia (Fig. 4C) with one straight spur and one calcar small, subtriangular, obtusely rounded at apex.
Ventral surfaces of pterothorax and abdomen (   num with only a few coarse punctures sparsely located along the midline; more coarse punctures closely arranged almost into a row near lateral margins; remainder of disc smooth; a shallow median pit present posteriorly. Each abdominal sternum with coarse punctures arranged into two or three irregular transverse rows; sternum IV with deep, round lateral pits; sternum V without visible pits; sternum VI with two setae near apical margin. Genital ring (Fig. 5E) subquadrate, with long handle, nearly parallel-sided, and rounded at tip. Aedeagus ( Fig. 5A-C) with median lobe thick, tubular; opening of apical orifice (Fig. 5D) large, subelliptical; left paramere ( Fig. 6D) broad, subelliptical; right paramere ( Fig. 6E) small, simply curved at outer margin, expanded in apical part. Endophallus as shown in Fig. 6F.
Female. Similar to male in general appearance, but distinguished by the following characteristics ( Fig. 2C-D): mentum surface with fewer setae, less coarsely punctate; profemur without tooth on ventral side; meso-and metatibiae without calcars; abdominal sternum IV with lateral pits distinctly larger.
Etymology. The specific epithet is from the name of "Sun Wukong", also known as the Monkey King, a mythological figure who features in a body of legends, which can be traced back to the period of the Song dynasty.
Remarks. This new species is probably closely allied to the two known species of the subgenus Omoglymmius from East Asia, O. (s. str.) sakuraii (Nakane) and O. (s. str.) laticeps Bell. They resemble each other in general appearance, but detailed comparison of selected morphological characters of importance show their differences (Table 1).  relatively large (Fig. 1E, G)

Stria IV
with one seta at about basal 2/9, one seta at about apical 2/7 of its length and one seta subapically ( Fig. 2A, C) with one seta at about basal 1/5, one seta at middle of its length and one seta subapically (Fig. 1A, C) with one seta at about apical 2/7 of its length and one seta subapically (Fig. 1E, G) Metasternum with only a few coarse punctures sparsely located along the midline; more coarse punctures closely arranged almost into a row near lateral margins; remainder of disc smooth; a shallow median pit present posteriorly (Figs 2B,D;3G) with more coarse punctures located along the midline; more coarse punctures closely arranged almost into a row near lateral margins; remainder of disc also with a certain number of coarse punctures; a deep median pit present posteriorly (Fig. 1B, D) with more coarse punctures located along the midline; more coarse punctures closely arranged almost into a row near lateral margins; remainder of disc also with a certain number of coarse punctures; a deep median pit present posteriorly (Fig. 1F, H) Aedeagus right paramere simply curved at outer margin and more expanded in apical part (Fig. 6E) unknown undulate at outer margin and less expanded in apical part (Fig. 6B) Female profemur without tooth on ventral side (Fig. 2D) without tooth on ventral side ( Fig. 1C-D) with a small tooth on ventral side (Fig. 1H Distribution. Bhutan, India (Fig. 7). Diagnosis. See Table 1 under Omoglymmius (s. str.) wukong sp. n. above.