Corresponding author: Kyohei Watanabe (
Academic editor: J. Fernandez-Triana
The Japanese species of the genus
Watanabe K (2017) Revision of the genus
The genus
In Japan, two species have been recorded, i.e.
Materials used were from the collections of
The holotype and paratype of
A stereomicroscope (Nikon S800) was used for observations. Photographs were taken by OLYMPUS TG-4 digital camera joined with the stereo microscope. Digital images were edited using Adobe Photoshop® CS6.
Morphological terminology mainly follows that established by
The following abbreviations are used in the material data and distribution:
The Japanese
Body covered with short setae. Head and mesosoma mat, covered with dense punctures. Ventral margin of clypeus blunt, without a median notch. Occipital carina complete. Lower end of occipital carina connected to hypostomal carina distant from base of mandible. Antenna with tyloid-like carina on ventral surface of basal 9–14 segments in female. Mesoscutum without a raised anterolateral area on each side. Epomia absent. Epicnemial carina present laterally and ventrally. Hind rim of metanotum without a sublateral triangular projection. Fore wing with: junction of vein
Eastern Palaearctic and Oriental regions.
The above description is partly referred from diagnosis provided by
1 | Base of T1 with a conspicuous yellow area or spots (Figs |
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– | Base of T1 without yellow area or spots (Figs |
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2 | Base of T1 with a pair of lateral yellow spots. Posterior segments of metasoma reddish brown. Antenna with 52 flagellomeres. Scutellum black with two yellow spots. Ovipositor sheath 0.4 times as long as hind tibia. Male unknown. Myanmar and India | |
– | Base of T1 with a wide yellow area or if with a pair of lateral yellow spots, propodeum with four yellow spots anteriorly and posteriorly (Fig. |
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3 | Propodeum with four yellow spots anteriorly and posteriorly in both sexes (Figs |
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– | Propodeum with two posterior yellow spots. Mesopleuron without two large yellow spots in female. |
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4 | T2-T7 entirely red. T1 ca. 2.0 times as long as maximum width |
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– | T2-T7 black with or without white band(s), without red area. T1 various in length |
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5 | Body large and elongate. T1 2.2–2.8 times as long as maximum width. T2 1.1–1.3 times as long as maximum width (Figs |
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– | Body small and not elongate. T1 1.8–2.1 times as long as maximum width. T2 0.9–1.0 times as long as maximum width (Figs |
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6 | Ovipositor sheath 0.3 times as long as hind tibia. Clypeus 0.5 times as long as wide. Face 0.6 times as long as wide, without a narrow longitudinal depression between eye and antennal socket (Figs |
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– | Ovipositor sheath 0.4–0.5 times as long as hind tibia. Clypeus 0.4 times as long as wide. Face 0.5 times as long as wide, with a narrow longitudinal depression between eye and antennal socket (Fig. |
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Clypeus 0.5 times as long as wide. Face 0.6 times as long as wide, without a narrow longitudinal depression between eye and antennal socket. Antenna with 45–47 flagellomeres. Hind femur 5.7–6.1 times as long as maximum depth in lateral view, black. T1 1.8–1.9 times as long as maximum width. Base of T1 without yellow area or spots. T2 0.9–1.0 times as long as maximum width. T2-T7 black with or without white band(s), without red area. Ovipositor sheath 0.3 times as long as hind tibia. Face sometimes without a pair of yellow stripes along inner orbit in female, or entirely yellow in male. Metasomal tergite nearly entirely black in female.
JAPAN: [Holotype]
Japan (Hokkaido, Honshu, Shikoku and Kyushu).
The specific name is from my friend, Mr. Shunpei Fujie, who is a taxonomist of Japanese
Host is unknown. The specimens collected from Hyogo (Mt. Hyonosen) and Shikoku were collected by light trap.
This species resembles
Clypeus 0.4 times as long as wide. Face 0.5 times as long as wide, with a narrow longitudinal depression between eye and antennal socket. Antenna with 46–49 flagellomeres. Hind femur 6.4–6.9 times as long as maximum depth in lateral view, black or reddish brown. T1 1.9–2.0 times as long as maximum width. Base of T1 without yellow area or spots. T2 0.9–1.0 times as long as maximum width. Each metasomal tergite with a posterior white band. Ovipositor sheath 0.4–0.5 times as long as hind tibia.
JAPAN: [Holotype]
Japan (Honshu, Kyushu, Tsushima Is. and Yakushima Is.),
The specific name is from a Japanese term “Miyabi”, which means elegant.
Host is unknown.
This species resembles
JAPAN: 1
Japan (Hokkaido, Honshu, Shikoku and Kyushu); Far East Russia.
Host is unknown. Some specimens collected from Shikoku and Kyushu were collected by light trap.
This is the first record of the male. Males were collected only from Hokkaido and Honshu.
Metasoma. T1 2.0–2.2 times as long as maximum width. T2 0.9–1.0 times as long as maximum width. Ovipositor sheath 0.4 times as long as hind tibia.
JAPAN: 1 F (holotype of
Japan (Tsushima Is. and Amamioshima Is.); Far East Russia (Primorye Kray) and Kazakhstan.
Host is unknown.
The distribution data for this species is relatively sparse as compared to other species. The locality of Kazakhstan is distant from Japan and Far East Russia (Primorye Kray) while no differences of character states were found between both specimens.
The author would like to express his cordial thanks to David Wahl (