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Article title
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Abstract
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Keywords
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Introduction
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The history and nature of the sphragis
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Formation of the sphragis
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Behavioral specialization
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Function of the sphragis
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The taxonomic distribution of the sphragis
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The present state of knowledge
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Methods
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Terms related to sphragis-like structures used in this study
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Results and discussion
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Variation in the morphology and color of the sphragis
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Towards an understanding of sphragis evolution
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Conclusions
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Acknowledgments
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References
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Supplementary material
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