2urn:lsid:arphahub.com:pub:45048D35-BB1D-5CE8-9668-537E44BD4C7Eurn:lsid:zoobank.org:pub:91BD42D4-90F1-4B45-9350-EEF175B1727AZooKeysZK1313-29891313-2970Pensoft Publishers10.3897/zookeys.1208.127725127725Research ArticleOonopidaeTaxonomyAfricaFour new species of the genus Camptoscaphiella Caporiacco, 1934 (Araneae, Oonopidae) from Xizang, ChinaWangXiaohanhttps://orcid.org/0009-0000-4614-95621TongYanfengtyf68@hotmail.comhttps://orcid.org/0000-0002-4348-70291LiShuqianghttps://orcid.org/0000-0002-3290-54162College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, ChinaShenyang Normal UniversityShenyangChinaInstitute of Zoology, Chinese Academy of Sciences, Beijing 100101, ChinaInstitute of Zoology, Chinese Academy of SciencesBeijingChina
20242607202412088198CCF6B926-24B8-53CF-9B63-BA792C88978664AFF550-64DD-4EBD-AEA5-2C38D0F6083A1705202407072024Xiaohan Wang, Yanfeng Tong, Shuqiang LiThis is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.http://zoobank.org/64AFF550-64DD-4EBD-AEA5-2C38D0F6083A
Four new species of the genus Camptoscaphiella Caporiacco, 1934 are described from Xizang, China, i.e., C.metok Tong & Li, sp. nov. (♂), C.shannan Tong & Li, sp. nov. (♂♀), C.trifoliata Tong & Li, sp. nov. (♂♀) and C.zayu Tong & Li, sp. nov. (♂♀). Morphological descriptions, photographic illustrations and a distribution map of the four new species are given.
Wang X, Tong Y, Li S (2024) Four new species of the genus Camptoscaphiella Caporiacco, 1934 (Araneae, Oonopidae) from Xizang, China. ZooKeys 1208: 81–98. https://doi.org/10.3897/zookeys.1208.127725
Introduction
The family Oonopidae is one of the most diverse spider families worldwide, comprising 115 extant genera and 1952 species, and 3 fossil genera and 45 species (WSC 2024). Camptoscaphiella Caporiacco, 1934 is a small genus, distributed mainly in montane tropical and subtropical regions in Asia, mostly within the Himalayan Plateau (WSC 2024). Baehr and Ubick (2010) revised this genus, with two species re-described and nine new species described from China, India, Nepal, Sri Lanka and Thailand, and a key to the known 15 species provided. Two other species were later recorded in the Pacific Island of New Caledonia (Baehr and Harvey 2013) and five species were documented from Yunnan, China (Huang et al. 2021; Wang et al. 2023). Altogether, 23 species have been recorded in the world (WSC 2024).
Up to now, seven species belonging to three genera of oonopid spiders have been recorded in Xizang of China (Cheng et al. 2021; Tong et al. 2023). Although there are no records of the genus Camptoscaphiella from Xizang, many species have been described from the adjacent areas, including one from Bhutan, three from India, seven from Nepal, and eight from Yunnan Province of China (Tong and Li 2007; Baehr and Ubick 2010; Huang et al. 2021; Wang et al. 2023).
In this paper, Camptoscaphiella is recorded for the first time from Xizang and four new species of the genus are described and photographed.
Materials and methods
The specimens were examined using a Leica M205C stereomicroscope. Details were studied under an Olympus BX51 compound microscope. Photos were captured with a Canon EOS 550D zoom digital camera (18 megapixels) mounted on an Olympus BX51 compound microscope. Endogynes were cleared in lactic acid. Scanning electron microscope (SEM) images were taken under high vacuum with a Hitachi S-4800 after critical-point drying and gold-palladium coating. All measurements were taken using an Olympus BX51 compound microscope and are in millimeters. The type material is deposited at the
Shenyang Normal University (SYNU) in Shenyang, China.
The following abbreviations are used in the text and figures:
ALE = anterior lateral eyes;
ap = apodemes;
as = anterior sclerite;
cd = copulatory duct;
dp = dorsal process;
mp = median plate;
mpr = median process;
np = narrow process;
PLE = posterior lateral eyes;
PME = posterior median eyes;
pro = prolateral outgrowth;
rlf = retrolateral fold;
tfp = trifurcate process;
tls = thread-like structure;
vo = ventral outgrowth;
vp = ventral process;
wlo = wing-like outgrowth.
Camptoscaphiellafulva Caporiacco, 1934, by monotypy.
Diagnosis.
Males of this genus are similar to those of Opopaea Simon, 1892 in the extremely large, club-shaped palpal patella, but can be distinguished by the spination of the legs I–II (tibia I and II with four pairs of long spines, and metatarsus I and II with two pairs of long spines) vs. without spination; the cymbium not fused with the bulb vs. fused; and the reduced abdominal scuta vs. complete scuta. Females of this genus are similar to those of Ischnothyreus Simon, 1893, but can be separated by lacking the distinct, darkly sclerotized, strongly winding duct of endogyne and the uniquely shaped atrium.
Distribution.
China (Xizang, Yunnan), South Asia (Bhutan, India, Nepal, Pakistan, Sri Lanka), Southeast Asia (Thailand) and New Caledonia.
Holotype China • ♂ (SYNU-1134); Xizang, Nyingchi, nr. Metok Co.; 29°19.382'N, 95°19.016'E, 980 m; 2.VIII.2013; Y. Lin leg.
Paratype: China • ♂ (SYNU-1135); same data as for holotype, Motuo Petrol Station; 22.IX.2013; Z. Gao leg.
Diagnosis.
This new species is similar to C.linyejiei Tong & Li, 2021, but can be distinguished by their normal-sized eyes (Fig. 1B, G) vs. reduced (Huang et al. 2021: fig. 4B, G); and the thread-like structure (tls) on the tip of the psemblous (Fig. 2G, H), vs. lacking; and having a plate-like process (Huang et al. 2021: fig. 5G, H).
Camptoscaphiellametok sp. nov., male holotype A, C, E habitus, dorsal, ventral and lateral views B, D, F, G prosoma, dorsal, ventral, lateral and anterior views, arrow shows the pointed anterolateral bumps. Scale bars: 0.4 mm (A–G).
Camptoscaphiellametok sp. nov., male left palp A, I prolateral view B, E bulb, prolateral and retrolateral views C, K dorsal view D, J retrolateral view F, G, H distal part of bulb, dorsal, prolateral and retrolateral views. Abbreviations: np = narrow process; rlf = retrolateral fold; tls = thread-like structure; vo = ventral outgrowth; wlo = wing-like outgrowth. Scale bars: 0.2 mm (A–E, I–K); 0.05 mm (F–H).
Male (holotype). Body: uniformly colored, yellow; habitus as in Fig. 1A, C, E; length 1.29. Carapace (Fig. 1B, F): 0.66 long, 0.54 wide; pars thoracica strongly elevated in lateral view, entire surface finely reticulated. Eyes (Fig. 1B, G): ALE: 0.05; PME: 0.04; PLE: 0.04; ALE circular, PME oval, PLE oval; posterior eye row procurved from both above and front; ALE separated by less than one radius. Clypeus (Fig. 1F, G): margin unmodified, straight in front view, sloping forward in lateral view. Mouthparts (Fig. 1D, G): chelicerae unmodified; anterior-median part of endites strongly sclerotized. Sternum (Fig. 1D): as long as wide, with pointed anterolateral bumps. Abdomen (Fig. 1A, C, E): 0.63 long, 0.34 wide; oval, scuta pale orange; dorsal scutum covering about 5/6 of abdomen length, about 3/4 of abdomen width, fused to epigastric scutum; postgastric scutum covering about 3/4 of abdominal venter. Palp (Fig. 2A–K): reddish brown; patella extremely long club-shaped, length/width = 3.04, about 5.6 times longer than femur, and 2.5 times longer than bulb; cymbium narrow (length/width = 1.68) in dorsal view; distal part of bulb with narrow process (np), broad ventral outgrowth (vo), a narrow wing-like outgrowth (wlo), a small retrolateral fold (rlf) and cluster of thread-like structure (tls).
Female. Unknown.
Notes.
Seven species [C.glenniei (Fage, 1946) from India, C.martensi Baehr, 2010, C.panchthar Baehr, 2010, C.silens Brignoli, 1976, C.strepens Brignoli, 1976 and C.taplejung Baehr, 2010 from Nepal and C.gunsa Baehr, 2010 from India and Nepal] in adjacent areas are known only from females and this new species can be potentially conspecific with one of them.
Etymology.
The specific name refers to the type locality and is a noun in apposition.
Holotype China • ♂ (SYNU-1143); Xizang, Shannan City, Cona Co., Lemenba Ethnic Township, 17–20 km section from Lewang Bridge to Liulian Hwy; 27°47.700'N, 91°46.417'E, 3706 m; 5.VI.2016; J. Wu leg.
Paratypes: China • ♀ (SYNU-1144); same data as for holotype • 6♀ (SYNU-1123–1128); same data as for holotype.
Diagnosis.
The new species is similar to C.tuberans Tong & Li, 2007 and can be distinguished by the single strong bristle behind cheliceral paturons (Fig. 3D–H), vs. bristle absent (Tong and Li 2007: fig. 23); the strongly sclerotized median process (mpr) and a broad ventral outgrowth (vo) of the psembolus (Fig. 4D, H, K) vs. lacking the median process and having instead a very narrow, bifurcated ventral outgrowth (Tong and Li 2007: figs 25–27); and narrow forcipate median plate (mp) of endogyne (Fig. 5F, H) vs. circular median plate (Tong and Li 2007: fig. 22).
Camptoscaphiellashannan sp. nov., male holotype A, C, E habitus, dorsal, ventral and lateral views B, D, F, H prosoma, dorsal, ventral, lateral and anterior views, arrow shows the pointed anterolateral bumps G labium and endites, ventral view. Scale bars: 0.4 mm (A–G).
Camptoscaphiellashannan sp. nov., male left palp A, K prolateral view B, E bulb, prolateral and retrolateral views C, L retrolateral view D, G, H distal part of bulb, prolateral, dorsal and retrolateral views F, J dorsal view I same asG showing the details (arrow). Abbreviations: mpr = median process; pro = prolateral outgrowth; rlf = retrolateral fold; vo = ventral outgrowth. Scale bars: 0.2 mm (A–C, E, F, J–L); 0.05 mm (D, G, H); 0.01 mm (I).
Camptoscaphiellashannan sp. nov., female paratype (SYNU-1144) A, C, E habitus, dorsal, ventral and lateral views B, D, G prosoma, dorsal, ventral and anterior views F epigastric region, ventral view H, I endogyne, ventral and dorsal views. Abbreviations: ap = apodemes; as = anterior sclerite; cd = copulatory duct; mp = median plate. Scale bars: 0.4 mm (A–E, G); 0.2 mm (F, H, I).
Male (holotype). Body: uniformly colored, yellow; habitus as in Fig. 3A, C, E; length 1.83. Carapace (Fig. 3B, F): 0.81 long, 0.72 wide; pars thoracica slightly elevated in lateral view, entire surface finely reticulated. Eyes (Fig. 3B, H): ALE: 0.08; PME: 0.06; PLE: 0.06; ALE circular, PME oval, PLE oval; posterior eye row straight from above, procurved from front; ALE separated by less than one radius. Clypeus (Fig. 3F, H): margin unmodified, straight in front view, sloping forward in lateral view. Mouthparts (Fig. 3G, H): with single strong bristle behind paturons; anterior-median part of the endites strongly sclerotized. Sternum (Fig. 3D): as long as wide, with pointed anterolateral bumps. Abdomen (Fig. 3A, C, E): 1.02 long, 0.73 wide; oval, scuta pale orange; dorsal scutum covering about 2/3 of abdomen, 1/2 of abdomen width, fused to epigastric scutum; postgastric scutum small. Palp (Fig. 4A–L): reddish brown; patella extremely long club-shaped, length/width = 3.30, about 5.8 times longer than femur, and 3.2 times longer than bulb, cymbium narrow (length/width = 1.90) in dorsal view; psembolus with prolateral and ventral outgrowths (pro and vo respectively), strongly sclerotized median process (mpr) and small retrolateral fold (rlf).
Female (SYNU-1144). As in male except for the following. Body: habitus as in Fig. 5A, C, E; length 1.98. Carapace: 0.86 long, 0.73 wide. Eyes: ALE 0.07; PME 0.05; PLE 0.07. Abdomen: 1.12 long, 0.80 wide. Postgaster (Fig. 5F, H): with stick-like anterior sclerite (as) and forcipate median plate (mp). Endogyne (Fig. 5I): copulatory duct (cd) long, narrow, straight with tip well beyond postgastric scutum; apodemes (ap) short.
Etymology.
The specific name refers to the type locality and is a noun in apposition.
Holotype China • ♂ (SYNU-1145); Xizang, Rikaze City, Jilong Co., Zalong Vill.; 28°22.865'N, 85°21.158'E, 2715 m; 31.VII.2014; Y. Li leg.
Paratypes: China • 1 ♀ (SYNU-1100); Rikaze City, Dingjie Co., Chentang Town; 27°54.875'N, 87°28.869'E, 3267 m; 3.VIII.2014; Y. Li leg. • 1 ♂ (SYNU-1099); Rikaze City, Dingri Co., Rongxia Town; 28°03.450'N, 86°21.148'E, 3383 m; 27.VII.2014; Y. Li leg. • 2 ♂ (SYNU-1054–1055); Nyingchi, Bayi Distr., Bayi Town, Biri Mt; 28°51.334'N, 94°47.941'E, 2900 m; 11.VII.2013; Y. Lin leg. • 11♀ (SYNU-1057–1067); same data as above • 4♀ (SYNU-1146–1149); same data as above • 2 ♂ 4 ♀ (SYNU-1113–1118); Nyingchi, Bayi Distr., Lulang Town; 29°41.449'N, 94°43.605'E, 3530 m; 14.VII.2013; Y. Lin leg. • 9 ♂ 2 ♀ (SYNU-1101–1112); Nyingchi, Bayi Distr., Lulang Town; 29°21.449'N, 94°43.605'E, 3530 m; 14.VII.2013; Q. Cao leg. • 5 ♂ 1 ♀ (SYNU-1068–1073); Nyingchi, Mainling Co.; 29°13.310'N, 94°13.309'E, 3050 m; 13.VIII.2013; Y. Lin leg. • 2 ♂ 1 ♀ (SYNU-1152–1154); same data as above • 6 ♂ 4 ♀ (SYNU- 1074–1083); Nyingchi, Mainling Co.; 29°12.316'N, 94°12.649'E, 3060 m; 13.VIII.2013; Y. Lin leg. • 4♂10♀ (SYNU-1084–1097); Nyingchi, Mainling Co., Pai Town; 29°30.264'N, 94°53.868'E, 3321 m; 6.VIII.2015; J. Wu leg.
Diagnosis.
Males of this new species are similar to those of C.yujufeng Tong & Yang, 2023, but can be distinguished by lacking a cluster of black, strong setae on the labium (Fig. 6D) vs. setae cluster present (Wang et al. 2023: fig. 5D) and psemblous with trifurcate process (tfp, Fig. 7D, K) vs. with semicircular, prolateral rim and trifurcate ventral process (Wang et al. 2023: fig. 6E, J). Females of the new species are similar to those of C.zayu sp. nov., but can be distinguished by the fusiform median plate (Fig. 8H) vs. pear-shaped (Fig. 11H) and the long copulatory duct, which extends beyond the postgastric scutum (Fig. 8I) vs. just reaching groove between posterior spiracles (Fig. 11I).
Camptoscaphiellatrifoliata sp. nov., male holotype A, C, E habitus, dorsal, ventral and lateral views, arrow shows the pointed anterolateral bumps B, D, F, G prosoma, dorsal, ventral, lateral and anterior views. Scale bars: 0.4 mm (A–G).
Camptoscaphiellatrifoliata sp. nov., male left palp A, K prolateral view B, E bulb, prolateral and retrolateral views C, L retrolateral view D, G, H distal part of bulb, prolateral, dorsal and retrolateral views F, J dorsal view I same asG showing the details (arrow). Abbreviations: rlf = retrolateral fold; tfp = trifurcate process; vo = ventral outgrowth. Scale bars: 0.2 mm (A–C, E, F, J–L); 0.05 mm (D, H, G); 0.01 mm (I).
Camptoscaphiellatrifoliata sp. nov., female paratype (SYNU-1146) A, C, E habitus, dorsal, ventral and lateral views B, D, G prosoma, dorsal, ventral and anterior views F epigastric region, ventral view H, I endogyne, ventral and dorsal views. Abbreviations: ap = apodemes; as = anterior sclerite; cd = copulatory duct; mp = median plate. Scale bars: 0.4 mm (A–E, G); 0.2 mm (F, H, I).
Male (holotype). Body: uniformly colored, yellowish brown; habitus as in Fig. 6A, C, E; length 1.75. Carapace (Fig. 6B, F): 0.84 long, 0.68 wide; pars thoracica strongly elevated in lateral view, entire surface finely reticulated. Eyes (Fig. 6B, G): ALE: 0.07; PME: 0.05; PLE: 0.04; ALE circular, PME oval, PLE oval; posterior eye row straight from above, procurved from front; ALE separated by less than one radius. Clypeus (Fig. 6F, G): margin unmodified, straight in front view, sloping forward in lateral view. Mouthparts (Fig. 6D, G): chelicerae unmodified; anterior-median part of the endites slightly sclerotized. Sternum (Fig. 6D): as long as wide, with pointed anterolateral bumps. Abdomen (Fig. 6A, C, E): 0.91 long, 0.53 wide; oval, scuta pale orange; dorsal scutum covering about 3/4 of abdomen length, about 2/3 of abdomen width, fused to epigastric scutum; postgastric scutum small, covering about 1/2 of abdominal venter. Palp (Fig. 7A–L): reddish brown; patella extremely long club-shaped, length/width = 3.41, about 5.5 times longer than femur, and 2.5 times longer than bulb; cymbium (length/width = 1.92) narrow in dorsal view; psembolus with trifurcate process (tfp), ventral outgrowth (vo) and small retrolateral fold (rlf).
Female (SYNU-1100). As in male except for the following. Body: habitus as in Fig. 8A, C, E; length 1.94. Carapace: 0.83 long, 0.74 wide. Eyes: ALE 0.07; PME 0.05; PLE 0.06. Abdomen: 1.11 long, 0.76 wide. Postgaster (Fig. 8F, H): with rounded anterior sclerite (as) and fusiform median plate (mp). Endogyne (Fig. 8I): copulatory duct (cd) long, narrow, sinuous with tip well beyond postepigastric scutum; apodemes (ap) slender.
Etymology.
The specific name, derived from Latin word trifoliatus, refers to the three-forked processes on the distal part of the bulb; adjective.
Holotype China • ♂ (SYNU-1141); Xizang, Zayu Co., Chawalong Town, Long Vill.; 28°28.941'N, 98°28.193'E, 2883 m; 8.IX.2014; J. Liu leg.
Paratypes. China • 1 ♀ (SYNU-1142); same data as for holotype.
Diagnosis.
The new species is similar to the type species, C.fulva Caporiacco, 1934, but can be distinguished by the strongly sclerotized dorsal (dp) and ventral (vp) processes of the psembolus (Fig. 10G, I) vs. with two spinelike processes (Baehr and Ubick 2010: figs 161–163), and long copulatory duct (cd, Fig. 11H, I) vs. short inverted droplet-shaped copulatory duct (Baehr and Ubick 2010: figs 182, 183).
Description.
Male (holotype). Body: uniformly colored, pale yellow; habitus as in Fig. 9A, C, E; length 1.91. Carapace (Fig. 9B, F): 0.85 long, 0.78 wide; pars thoracica strongly elevated in lateral view, whole surface finely reticulated. Eyes (Fig. 9B, G): ALE: 0.08; PME: 0.06; PLE: 0.06; ALE circular, PME oval, PLE oval; posterior eye row procurved from both above and front; ALE separated by less than one radius. Clypeus (Fig. 9F, G): margin unmodified, straight in front view, sloping forward in lateral view. Mouthparts (Fig. 9D, G): chelicerae unmodified; anterior-median part of the endites slightly sclerotized. Sternum (Fig. 9D): as long as wide, with pointed anterolateral bumps. Abdomen (Fig. 9A, C, E): 1.06 long, 0.68 wide; oval, scuta pale orange; dorsal scutum covering about 5/6 of abdomen length, about 3/4 of abdomen width, fused to epigastric scutum; postgastric scutum small, covering about 1/2 of abdominal venter. Palp (Fig. 10A–K): reddish brown; patella extremely long club-shaped, length/width = 2.96, about 4.2 times longer than femur, and 2.5 times longer than bulb; cymbium narrow (length/width = 1.64) in dorsal view; psembolus with strongly sclerotized dorsal process (dp), strongly sclerotized ventral process (vp), broad ventral outgrowth (vo) and a small retrolateral fold (rlf).
Camptoscaphiellazayu sp. nov., male holotype A, C, E habitus, dorsal, ventral and lateral views B, D, F, G prosoma, dorsal, ventral, lateral and anterior views, arrow shows the pointed anterolateral bumps. Scale bars: 0.4 mm (A–G).
Camptoscaphiellazayu sp. nov., male left palp A, I prolateral view B, E bulb, prolateral and retrolateral views C, K dorsal view D, J retrolateral view F, G, H distal part of bulb, dorsal, prolateral and retrolateral views. Abbreviations: dp = dorsal process; rlf = retrolateral fold; vo = ventral outgrowth; vp = ventral process. Scale bars: 0.2 mm (A–E, I–K); 0.05 mm (F–H).
https://binary.pensoft.net/fig/1102646
Female (SYNU-1142). As in male except for the following. Body: habitus as in Fig. 11A, C, E; length 2.37. Carapace: 0.92 long, 0.81 wide. Eyes: ALE 0.07; PME 0.05; PLE 0.05. Abdomen: 1.45 long, 0.90 wide. Postgaster (Fig. 11F, H): with rounded anterior sclerite (as) and pear-shaped median plate (mp). Endogyne (Fig. 11I): copulatory duct (cd) long, narrow, straight with tip reaching groove between posterior spiracles; apodemes (ap) short.
Camptoscaphiellazayu sp. nov., female paratype (SYNU-1142) A, C, E habitus, dorsal, ventral and lateral views B, D, G prosoma, dorsal, ventral and anterior views F epigastric region, ventral view H, I endogyne, ventral and dorsal views. Abbreviations: ap = apodemes; as = anterior sclerite; cd = copulatory duct; mp = median plate. Scale bars: 0.4 mm (A–E, G); 0.2 mm (F, H, I).
Distribution records of four new species of the genus Camptoscaphiella from Xizang, China. 1. C.metok sp. nov. 2. C.shannan sp. nov. 3. C.trifoliata sp. nov. 4. C.zayu sp. nov.
https://binary.pensoft.net/fig/1102648Etymology.
The specific name refers to the type locality and is a noun in apposition.
Distribution.
Known only from the type locality.
Acknowledgements
Thanks to Qi Cao, Zhizhong Gao, Yejie Lin, Jincheng Liu, and Jianglang Wu for collecting specimens. The manuscript benefitted greatly from comments by Yuri M. Marusik (Magadan, Russia) and Zhisheng Zhang (Chongqing, China). Joseph K. H. Koh (Singapore) kindly checked the English of the final draft.
Additional informationConflict of interest
The authors have declared that no competing interests exist.
Ethical statement
No ethical statement was reported.
Funding
This study was supported by the National Natural Science Foundation of China (NSFC32370479, 31972867).
Author contributions
YT designed the study. YT and XW finished the descriptions and took the photos. YT and SL drafted and revised the manuscript.
Author ORCIDs
Xiaohan Wang https://orcid.org/0009-0000-4614-9562
All of the data that support the findings of this study are available in the main text.
ReferencesBaehrBCHarveyMS (2013) The first goblin spiders of the genus Camptoscaphiella (Araneae: Oonopidae) from New Caledonia.Australian Journal of Entomology52(2): 144–150. https://doi.org/10.1111/aen.12010BaehrBCUbickD (2010) A review of the Asian goblin spider genus Camptoscaphiella (Araneae: Oonopidae).American Museum Novitates3697: 1–65. https://doi.org/10.1206/3697.2ChengWBianDTongYLiS (2021) A new genus and two new species of oonopid spiders from Tibet, China (Araneae, Oonopidae).ZooKeys1052: 55–69. https://doi.org/10.3897/zookeys.1052.66402HuangYBianDTongYLiS (2021) Two new species of the genus Camptoscaphiella from Yunnan, China (Araneae: Oonopidae).ZooKeys1052: 43–53. https://doi.org/10.3897/zookeys.1052.66743TongYLiS (2007) One new genus and four new species of oonopid spiders from southwest China (Araneae: Oonopidae).Annales Zoologici (Warszawa)57: 331–340.TongYBianDLiS (2023) Three new species of the genus Ischnothyreus Simon, 1893 and the discovery of the male of I.linzhiensis Hu, 2001 from Tibet, China (Araneae, Oonopidae).ZooKeys1152: 119–131. https://doi.org/10.3897/zookeys.1152.100341WangXWangZTongYBianDYangZ (2023) Three new species of Camptoscaphiella Caporiacco, 1934 (Araneae, Oonopidae) from Yunnan Province, China. Biodiversity Data Journal 11: e109679. https://doi.org/10.3897/BDJ.11.e109679WSC (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern. http://wsc.nmbe.ch [Accessed on: 30 April 2024]