Male adult Chordodes formosanus from novel hosts. A Posterior end B Anterior end C–D Variable crowned areole morphologies from different individuals E Close view of (C) with typical C. formosanus crowned areoles; F Close view of (D) with smaller crowned areoles. Ca, crowned areole. Scale bars 100 µm (A), 10 µm (B), 100 µm (C–D), and 10 µm (E–F).