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  • Article title
  • Abstract
  • Key words
  • Introduction
  • Materials and methods
    • Study area and sampling
    • DNA extraction and purification
    • mtDNA sequencing
    • Microsatellite analyses
    • Morphological analyses
    • Geometric morphometrics of landmark data
    • Analysis of distance measurements
    • Analysis of scale counts and colour markings
    • Linear discriminant analysis
  • Results
    • Distribution of mtDNA lineages
    • Microsatellite clusters
    • Morphology
  • Discussion
  • Acknowledgements
  • Additional information
    • Conflict of interest
    • Ethical statement
    • Funding
    • Author contributions
    • Author ORCIDs
    • Data availability
  • References
  • Supplementary material
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