Corresponding authors: Petr Bulirsch (
Academic editor: A. Casale
The genera
Bulirsch P, Stachowiak M (2017) Overview and new records of the species of the tribes Dyschiriini and Clivinini from Iraq (Coleoptera, Carabidae, Scaritinae). ZooKeys 672: 135–144.
The scaritine tribes
This article is based on the collections listed below. The specimens collected by Z. Stebnicka and J. Pawłowski during the 1977–1978 expedition of
Abbreviations used in the key are as follows:
This genus comprises approximately 650 species distributed almost worldwide.
1 specimen: Hatra, 3 v 1978, at light in a hotel, leg. Z. Stebnicka, (
A widespread, common and variable species reported from exact localities in Iraq for the first time.
A poorly known species described from Iraq.
A common species widespread in the Mediterranean area to the Middle Asia.
This genus in the sense of
1 specimen: Iraq, Kirkuk, lgt. W. Schors, (
A common subspecies widespread in NW Africa to the Middle Asia.
Unlike
1 specimen: Iraq, Tekrit, ii-v.1979, (
A very common species widespread from NW Africa to W Kazakhstan.
5 specimens: Iraq, Tekrit, ii-v.1979, (
A rather common species widespread from NW Africa to the Middle Asia.
1 specimen (ab.
A rather common species populating Sardinia, Sicilia, North Africa and extending eastward as far as Iraq and Iran.
Not a frequent species in Turkey, Iraq and Iran.
1 specimen: Iraq, Euphrat, (
A rare species in Turkey and Iraq (one old record).
Not a common species, distributed from Turkey to Middle Asia.
1 specimen: Iraq, Tekrit, ii-v.1979, (
Some specimens of
1 specimen: Iraq: Razeza Lake, 15 km W Kerbala, 14 v 1978, leg. Z. Stebnicka, (
A very common subspecies, very largely distributed from CE Europe, N Africa to Mongolia.
1 specimen: Iraq: Hatra, 3.v.1978, leg. Z. Stebnicka, (
A common subspecies distributed from Italy to Mesopotamia.
Paratypus, Egitto, Helwan, 18.ii.1935, W. Wittmer; 5 specimens: Iran Prov. Markazi, Kavir Desert, Houz-e Soltan, 830m, 3 km S Kushk-e Nosrat;
1 specimen: Iraq: Tharthar Lake, vii-viii.1977, leg. J. Pawłowski, on banks of lake, in detritus, together with
The genus includes a single species described from Heluan in Egypt and then reported from Karaman Island in Yemen (
Habitus of
The genus is recognized as including one rare species from North Africa (with two subspecies) and the other one from the Middle Asia.
1 specimen: Iraq: Hatra, 3.v.1978, leg. Z. Stebnicka, (
The locality in Iraq is subequally distant from those of
(including possible species in square brackets)
1 | Head tumid; body cylindrical, redish to testaceous; pronotum distinctly longer than broad |
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– | Head not tumid. Other combination of characters. |
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2(1) | Head and pronotum anteriorly coarsely rugose. |
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– | Head and pronotum smooth. |
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3(1') | Clypeus tridentate; elytra with basal ridge, fore tibia strongly dentate. |
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– | Clypeus not tridentate, at most broadly vaulted in middle; elytra without, rarely with gently basal ridge, fore tibia less strongly dentate | |
4(3') | Fore tibia with indistinct lateral teeth and with distinctly curved apical spine. |
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– | Fore tibia with distinct marginal teeth (at least lower one) |
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5(4) | Pronotum and elytra broader, elytral striae finer. |
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– | Pronotum and elytra narrower, elytral striae coarser. |
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6(4') | Basal |
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– | Basal |
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7(6) | Elytral striae not weakened apically; lateral channel of pronotum not shortened. |
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– | Elytral striae weakened to obliterated apically |
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8(7) | Head with frons coarsely rugose; elytral base with small tubercle. |
[ |
– | Head with straight or obsolete clypeofrontal suture; elytral base with two distinct tubercles. |
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9(7') | Lateral channel of pronotum not shortened. |
[ |
– | Lateral channel of pronotum strongly shortened. |
[ |
10(6') | Lateral channel of pronotum shortened, not reaching posterior |
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– | Lateral channel of pronotum not shortened, at least reaching posterior |
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11(10) | Clypeofrontal suture V-shaped, elytral striae obliterated apically |
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– | Clypeofrontal suture straight; lateral channel of pronotum strongly shortened, disappeared just below anterior |
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12(11) | Lateral channel of pronotum strongly shortened, disappeared just below anterior |
[ |
– | Lateral channel of pronotum slightly shortened, disappeared just above posterior |
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13(10') | Elytra cylindric; striae deep apically |
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– | Elytra shorter, ovate to elliptic |
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14(13) | Each elytron with two distinct, often fused tubercles at base. |
[ |
– | Elytra without or with a small basal tubercle |
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15(14) | Larger species. |
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– | Smaller species. |
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16(15) | Clypeofrontal suture prolonged posteriorly by distinct keel; pronotum and elytra in average narrower. |
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– | Clypeofrontal suture not prolonged posteriorly by distinct keel; pronotum and elytra in average broader. |
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17(13') | Elytral base with two tubercles; striae deep throughout. |
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– | Elytral base with 0-1 tubercles. |
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18(17) | Elytral base with one tubercle; elytra longer, with striae not to slightly weakened apically. |
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– | Elytral base without distinct tubercle; elytra shorter, with striae disappeared to strongly weakened apically. |
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We would like to thank Prof. Jerzy Pawlowski for supplying material for this publication and for additional information about the circumstances of its collection.