Morphological measurements. Indicating measurements of the worker (a, b, c) and male (d). HWe Head width including the eyes HW head width just above the eyes HW1 head width at the top of the head HW2 posterior head width HW3 anterior head width HL1 head length 1 HL2 head length 2 SL scape length IFW1 interfrontal width 1 IFW2 interfrontal width 2 WL Weber’s length PL petiole length PPL postpetiole length GL gaster length PW pronotal width HFL hind femur length IOD interocular distance.