East slope of Mountain Kapudschikh, Armenia.

Holotype, ♂, pinned, in ZIN. The holotype is labelled: ‘Vostochnyi skl. g. Kapudschikh, Armenia, V. Zaitzev, 17.VIII.1959’ [East slope of mtn. Kapudschikh, Armenia]; ‘Holotypus Cheilosia armeniaca Stackelberg’.

♂: Face in frontal view slightly divergent from level of antennal insertion to lower mouth edge, covered with very weak narrow strip of pollinosity (Fig. 2A). Facial knob rounded, not very protruding, shining. Parafacia pollinose in upper part and shining in lower part, in its broadest part nearly 1/2 the width (in its narrowest part about 1/3) of basoflagellomere. Frons slightly swollen, shining, with dense white wavy pile. Frontal angle ≈ 90° (Fig. 1, indicated with letter C). Lunula dark-brown. Antennal pits confluent. Vertical triangle with white pile. Eye-contiguity shorter than length of frons without lunula (A < B, see Fig. 1). Antenna dark-brown, basoflagellomere rounded (Fig. 2B). Arista with very short pile. Scutum, scutellum and pleura shining, with only very pale yellowish (almost whitish) pile, longer pile on pleura and posterior part of scutum distally wavy. Scutum and scutellum completely without black bristles or bristle-like pile. Postalar callus without black bristles. Anepisternum and katepisternum shining, with slight pollinosity, with white pile. Katepisternum with dorsal and ventral hair patches connected. Legs mainly black, knees narrowly yellow, tibiae also narrowly yellow at the extreme tip, 1–3 segments of meso- and 2–3 segments of metatarsi reddish (in the paratype metatarsi black with reddish tips of tarsomeres). Apical half of posterior surface of mesofemur with long light pile. Metafemur with short and long white pile ventrally. Halter yellow. Wing with all cross-veins darkened. Cell bm and cell cup of wing with areas bare of microtrichia. R_{4 + 5} of wing not distinctly curved. Abdomen oval, shining, densely punctated. Pile erect and white, longer on the lateral parts and shorter on the central parts of tergites. Sternite I with pollinosity, sternites II–IV shining. Pile erect, white on sternites. Hypopygium as on Fig. 2C–E.

Size. Body length 8.5 mm.

Georgia 1♂; ‘Georgia, Gornaia Tushetia, Omalo village 30.VII.1959 (Zaitsev)’ [ZIN].

Caucasus: Armenia, Georgia*.

Female unknown.

Österreich [Austria].

Lectotype, ♂, in MNHN. The original description is for both sexes. Becker (1902: 354) mentions only one male specimen which agrees with the original description and he designates it ‘Die Type’. The lectotype bears the following labels: ‘Meigen’ [handwritten in ink, round greyish collection label]; ‘caerulescens’ [Meigen’s handwriting].

Male: Fig. 4A. Face in anterior view slightly divergent from level of antennal insertion to lower mouth edge, with lower part protruding, with pollinosity laterally and below antenna (Fig. 3A). Facial knob moderate in size, shining. Parafacia rather narrow, in width slightly more than 1/2 the width of basoflagellomere, shining in lower part and with pollinosity on upper part narrowly along the eye. Frons slightly swollen, shining, with white and posteriorly some black pile, yellow pile anteriorly along eye inclined. Frontal angle about 90°. Lunula dark brown. Antennal pits confluent. Vertical triangle with black pile. Eye-contiguity about equal or slightly shorter than the length of frons without lunula. Scape and pedicel dark-brown, basoflagellomere small, dark brown to yellowish brown. Arista rather long, with short pile (Fig. 3C). Scutum shining (with bluish tinge), rather coarsely punctated, with whitish to yellowish erect, rather long pile of about even length, intermixed with black pile of same length or slightly longer. Scutellum with yellowish erect pile of same length as on scutum, margin with some longer black or yellow bristles. Postalar callus with 3–5 black bristles. Anepisternum and katepisternum shining, with very slight pollinosity, and with white long pile. Katepisternum with dorsal and ventral hair patches connected. Wing completely microtrichose, hyaline, crossveins darkened. R_{4 + 5} of wing not distinctly curved, angle of R_{4 + 5} and M₁ about 90°. Halter yellow. Legs with distal part of femora and tibiae, and basal 1/3–1/2 of tibiae yellowish, tarsi dorsally dark-brown. Mesofemur posteriorly with white pile, metafemur anteroventrally and dorsally with long white pile, and ventrally with shorter (stronger) black pile. Abdomen slightly oval, shining (bluish tinge), with coarse punctation, with long erect yellow pile on tergite I, tergites II–IV with erect yellow pile laterally (longest on tergite II), and somewhat shorter erect yellow and black pile intermixed medially. Yellow pilosity on postero-lateral corners tergites II–IV inclined. Sternite I with pollinosity, sternites II–IV shining, all with long yellowish pile. Hypopygium as in Fig. 3E–G.

Female: Fig. 4D. Face as in the male. Eye-margin width about 1/2 the width of basoflagellomere. Frons coarsely punctuated, with whitish erect pile anteriorly, and with black erect pile on posterior part (Fig. 3B). Whitish pile on anterior part of frons inclined along eye. Vertical triangle with black and white pile. Basoflagellomere rounded, reddish brown to dark-brown (Fig. 3D). Scutum shining, with white pile of even length, with few scattered black pile of about the same length or slightly longer intermixed. Postalar callus with 0–3 black or yellow bristles. Scutellum margin with 2–4 black or yellow bristly-like pile. Metafemur with only yellow pile. Tibiae in basal 1/3 yellow, apically narrowly and obscurely yellow, with broad dark ring, tarsi dorsally brown or yellow, with brown segments 4–5 of protarsi, segment 5 of mesotarsi and segments 3–5 of metatarsi. Abdomen oval. All sternites shining. Otherwise as the male.

Size. Body length 6.5–12 mm.

Switzerland 2 ♂, 2 ♀ Zermatt, 1 600–1 700 m, 30.VII.1964 (v.d. Goot; Lucas) [RMNH]; 1 ♂ Zermatt, 30.7.1930 [MNB]; Italy 1 ♂, 1 ♀ Italia, Südtirol, W Tubre di Avigna 1 300–1 800 m 6.VII.1988 (Claussen) [ISEA], 1 ♂ Stilfserjoch [MNB], 2 ♂ Italia, Alpen, Vinschgau Martelltal, Lyfi-Alm, 2 000–2 500 m, 1.7.1995 (Doczkal) [coll. Doczkal], 1 ♂ Italy, Stelvio Park, Trafoi, 1 900–2 000 m, 20.6.1993 (Ståhls) [MZH]; France 1 ♂ ‘Le Lautaret, 4.VIII.26’ [MNHN], 1 ♀ ‘Le Lautaret, 2.VIII.26’ [MNHN], 1 ♂ ‘Le Lautaret, 12.VI.1919’ [MNHN]; Austria ‘Wien, Egger’ [MNB]. See also Table 2 for data molecular specimen vouchers also used for morphological study.

Austria, Belgium, Czech Republic, France, Germany, Great Britain, Italy, Netherlands, Montenegro, Poland, Romania, Spain, Slovakia, Switzerland.

Color of pile on scutum varies from almost yellow with some black pile to yellow with broad stripe of black pile between wing bases (Fig. 4B–C). Color of tarsi varies from dark to mainly yellow (especially in female). Some studied females do not have yellow bristles or bristle-like pile on scutellum hind margin. The color of basoflagellomere varies from dark brown to yellowish brown. We treat the taxon as subspecies until evidence to the contrary is presented.

‘Lipetzk Region, Galichia Gora reserve’ (Russia).

Holotype, ♂, in ZIN. The holotype is labelled: ‘Lipetzkaya obl. Galichia gora 18.V.1963 Skufjin’.

♂: Face in frontal view slightly divergent from level of antennal insertion to lower mouth edge, slightly elongated, shining with stripe of pollinosity. Facial knob moderate, shining (Fig. 5A). Parafacia shining with slight pollinosity, moderate in width, in broadest part about 2/3 of the width of basoflagellomere. Frons very slightly swollen, shining, with white and posteriorly some black pile. Frontal angle slightly < 90°. Lunula dark-brown. Antennal pits separated. Vertical triangle with black pile. Eye-contiguity approximately equal to length of frons without lunula. Basoflagellomere oval, yellow, arista brownish, almost bare, with very short pile (Fig. 5B). Scutum black, shining with fine punctuation, covered with dense erect bright yellow pile of equal length, with black pile of same length intermixed. Postalar callus with black bristles. Scutellum hind margin with some long fine yellow and black pile. Pleurae with rather dense gray pollinosity and yellow waving on the end pile. Dorsal and ventral hair patch of katepisternum connected. Femora black with yellow extreme tips; mesofemur with long yellow pile on posterior surface of basal 4/5 and black pile of apical 1/5. Pile on anterior surface of metafemur long and yellow in basal 2/3 and short and depressed in apical 1/3. Tibiae yellow with black ring, tarsi yellow with black fifth segment. Halter yellow. Wing completely covered with microtrichia, with brown spot on cross-veins. R_{4 + 5} of wing not distinctly curved. Abdomen oval, shining black covered with erect yellow pile. Sternite I and IV with pollinosity, sternites II–III shining. Genitalia as in Fig. 5E–G.

Female: Face as in male, parafacia shining, narrow, about 1/3 of width of basoflagellomere. Frons comparatively broad with two furrows (Fig. 5D). Vertical triangle with black pile. Basoflagellomere slightly broader than in male (Fig. 5C). Pilosity of scutum and scutellum as in male, but all bristles on scutellum hind margin yellow. Mesofemur with long yellow pile on posterior surface. Tibiae yellow with rather narrow black ring. Tarsi yellow with darkened 5th segment. Wing and abdomen as in male.

Size. Body length 9–10.2 mm.

Russia 4 ♂, 2 ♀ ‘Lipetzk Region, Galichia Gora reserve, 24–28.IV.1966, Kuznetzova’ [VSU]; 1 female ‘Lipetzk Region, Galichia Gora reserve, 12.5.1974, V. Kuznetsova’ [ZIN]; 1♂, 1 ♀ same place 24–25.IV.1966, (V. Kuznetzova) [ISEA].

European part of Russia.

We treat the taxon as subspecies until evidence to the contrary is presented.

Holotype, ♂, pinned, with genitalia dissected in microvial, in ZIN. The holotype is labelled: ‘Teberdinskij zapov., okr. Usad’by, Kavkaz, Gorodkov, 9.VII.[1]968’, ‘g. Mal. Khatipara, verch. granitsa lesa, 2400 m’, ‘subalpinskij lug’ [Teberdinskii reserve, mountain Malaia Khatipara, close to border forest, 2400 m, subalpine meadow] [North Caucasus, Karastschaevo-Tscherkessia].

Paratype, 1 ♂, Teberdinskij reserve, mountain Malaia Khatipara, 1400–2000 m 14.VII.1982 (Lukashova) (ISEA).

♂: Face in frontal view slightly divergent from level of antennal insertion to lower mouth edge, with weak narrow strip of pollinosity (Fig. 6A). Facial knob rounded, not very protruding, shining. Parafacia pollinose in upper part and shining in lower part. Parafacia in its broadest part slightly more than 1/2 the width of basoflagellomere. Frons very slightly swollen, shining, with dense white wavy pile. Frontal angle ≈ 90°. Lunula brown. Antennal pits confluent. Vertical triangle with white pile. Eye-contiguity shorter than the length of frons without lunula. Basoflagellomere yellowish brown, rounded (Fig. 6B). Arista with short pile. Scutum, scutellum and pleura shining, with only very pale yellowish (almost whitish) pile, longer pile on pleura and posterior part of scutum distally wavy. Scutum and scutellum completely without black bristles or bristly like pile. Postalar callus without black bristles. Anepisternum and katepisternum shining, with white pile. Katepisternum with dorsal and ventral hair patches connected. Legs mainly black, knees narrowly yellow, tibiae also narrowly yellow at the extreme tip, 1–3 segments of meso and 2–3 segments of metatarsi yellowish. Apical half of posterior surface of mesofemur with long light pile. Metafemur with short and long white pile ventrally, apicoventrally with a few black pile. Halter yellow. Wing with all cross-veins darkened, completely microtrichose, R_{4 + 5} of wing not distinctly curved. Abdomen oval, shining, densely punctated. Pile erect and white, longer on the lateral parts and shorter on the central parts of tergites. Sternite I with pollinosity, sternites II–IV shining, pile erect, white. Hypopygium as on Fig. 6D–F.

Size. Body length 9 mm.

The specific name means “from Tscherkessia” in Latin and English.

No additional material available.

Russia (Caucasus).

Female unknown.

“Roumaine, Carpates Meridionales, Monts Mehedinti, Baile Herculane, Vallee du ruisseau Feregari” [Herculane, Romania].

Holotype, male, in MGAB [not available for study]. Paratypes, a male and a female. Paratypes are labelled: ’Romania, Baile Herculane, 5.IX.1980, leg. VL Brădescu’, in coll. Claussen.

♂: Face in frontal view slightly divergent from level of antennal insertion to lower mouth edge, shining, with very weak stripe of pollinosity, lower part somewhat protruded (Fig. 7A). Parafacia rather narrow, approximately 1/2 the width of basoflagellomere, shining, with slight pollinosity on upper part. Frons slightly swollen, shining, with white and posteriorly some black, pile. Frontal angle ≈ 90°. Lunula black. Antennal pits confluent. Vertical triangle with black and a few yellow pile. Eye-contiguity slightly shorter than length of frons without lunula. Basoflagellomere small, pale-brown to dark-brown, arista rather long, with short pile (Fig. 7B). Scutum shining, rather coarsely punctated, all covered with whitish pile of even length, also on scutellum. Scutellum margin without stronger bristles. Anepisternum and katepisternum partly shining, with white long pile. Katepisternum with dorsal and ventral pile patches broadly divided posteriorly and narrowly connected anteriorly. Halter brownish. Wing with all cross-veins infuscated. Cell bm and cell cup of wing with obvious bare areas (Fig. 7H). R_{4 + 5} of wing not distinctly curved. Postalar callus without black bristles. Legs with distal part of femora and apical 1/3–1/2 and distal 1/6 of tibiae yellowish, tarsi yellow to yellowish brown, pile mainly white. Mesofemur posteriorly with long yellow pile. Metafemur ventrally with short black (rather strong) pile, distally dorsally with yellow pile, its anterior surface with long yellow pile. Abdomen shining, with pollinosity on medial anterior 2/3 of tergite II, and on medial anterior 1/3 of tergite III. Abdomen with white erect pile, longer on the sides and shorter medially. Hypopygium Fig. 7E–F.

♀: Face and parafacia as in the male. Frons shining, coarsely punctated, with white erect pile, and mixed with black erect pile on posterior part. Frontal furrows moderate (Fig. 7D). Lunula dark-brown. Basoflagellomere yellowish brown. Scutum shining, with white pile of even length. Otherwise as the male.

Size. Body length 10–11 mm.

No additional material available.

Romania, Montenegro, FYR Macedonia.

“Der Hartz, Oesterreich” [sic][Austria].

Lectotype, ♂, pinned, in MNB, here designated to fix the concept of Cheilosia hercyniae Loew and to ensure the universal and consistent interpretion of the same. The original description concerns only the male sex. The lectotype is labelled: ‘Austria, Schiner’, ‘Coll. H. Loew’, ‘Zool. Mus., Berlin’, ‘Lectotype Cheilosia hercyniae Loew, Ståhls & Barkalov des.’.

♂: Face in frontal view moderately divergent from level of antennal insertion to lower mouth edge, with slight to moderate pollinosity, but facial knob shining. Facial knob moderately protruding (Fig. 8A). Parafacia rather broad, approximately 2/3 of width of basoflagellomere, with slight pollinosity in upper part and shining in lower part and with short white pile. Frons moderately swollen, shining, with black pile. Frontal angle slightly > 90°. Lunula reddish brown. Antennal pits separated. Vertical triangle with black pile. Eye-contiguity nearly equal to the length of frons without lunula. First and second antennal segment dark-brown, third segment somewhat quadratic, bright orange with dorsal margin black (Figs 8B, 12C). Arista pubescent, with very short pile. Scutum shining (bluish ting), finely punctated, with whitish to yellowish erect, rather long, pile of about even length, intermixed with black pile of same length or slightly longer, also on scutellum, margin with 8–10 longer black bristles. Postalar callus with 3–5 bristles. Anepisternum and katepisternum shining, with slight pollinosity, and with whitish and black, rather long, pile intermixed. Katepisternum with dorsal and ventral hair patches broadly divided. Wing brownish. Wing completely microtrichose. R_{4 + 5} of wing not curved. Halter yellow, knob dark-brown. Legs with distal part of femora, tibia with basal 1/3–1/2 and distally narrowly yellow, tarsi dorsally dark-brown. Mesofemur posteriorly with long yellow pile and with some black ones on the tip; metafemur ventrally with short black (rather strong) pile, distally dorsally with longer black and yellow pile, its anterior surface with long yellow pile in basal 2/3 and with black pile in apical 1/3. Abdomen slightly oval, shining tergites I–III with brownish pollinosity medially, tergite IV with stripe of pollinosity antero-medially. Pile yellow, erect on all tergites, tergite IV on postero-lateral corners also with black pile. Pregenital segments with pollinosity, with yellow pile. Sternites pollinose with yellow erect pile, sternite IV also with short appressed black pile medially. Hypopygium as on Fig. 8E–F.

♀: Face and parafacia as in male. Basoflagellomere big, bright orange-yellow, quadratic, with a long (≈ 1/3 of the length of antenna) groove on the inner ventral side (Fig. 8C). Anterior 1/3 of frons very shiny, with fine punctation, and very short pile, posterior 2/3 shining, coarsely punctated, with short yellow and longer black erect pile (Fig. 8D). Scutum shining, rather coarsely punctated, with short erect yellow and slightly longer black pile. Scutellum margin with 4–6 rather strong black bristles. Legs as in the male. Wing yellowish. Abdomen oval, shining, densely punctated, tergite I and antero-medial part of tergite II with pollinosity. Pile erect, yellow on lateral parts, long on lateral part of tergite II, and short, black, appressed pile medially. Sternites with slight pollinosity. Otherwise as the male.

Size. Body length 8–11 mm.

Italy 1 ♂ Italy, Val Gardena, Col Raiser-n. Regensb. Hütte, 2150–2200 m, 46°35'N 11°44'E, 22.06.2013, leg. T. & W. Romig [coll. Romig]; France 1 ♂ ‘Le Lautaret, 10.VII.24’ [MNHN], 1 ♂ ‘Le Lautaret, 31.VII.26’ [MNHN], 1 ♂ ‘Le Lautaret, 5.VIII.23’ [MNHN], 1 ♂ ‘Le Lautaret, 4.VIII.25’ [MNHN]; Montenegro 1 ♂ Montenegro, Durmitor, Krecmani, 30.VII.1999, leg. Vujić [FSUNS]; Switzerland 1 ♂ ‘Helvetia, GR, Ftan /Clünas, 2 100 m, 5.VIII.1996, Merz & Bächli’ [ETH]. See also Table 2 for data molecular specimen vouchers also used for morphological study.

Austria, Czech Republic, France, Germany, Italy, Montenegro, Poland, Romania, Spain, Switzerland,

“Ostungarn, Szászka und Rév” [Romania].

Type material could not be located, apparently lost.

♂: Face in frontal view slightly divergent from level of antennal insertion to lower mouth edge, shining with very slight pollinosity, lower part somewhat protruded (Fig. 9A). Facial knob rounded, shining, moderately protruded. Parafacia rather narrow, in the upper part approximately 1/2 the width of basoflagellomere, in lower part slightly narrowed. Parafacia shining, or with slight pollinosity on upper part. Frons slightly swollen, shining, with predominantly yellow and a few black pile. Frontal angle ≈ 90°. Lunula black. Antennal pits separated. Vertical triangle with black and a few yellow pile. Eye-contiguity slightly shorter than length of frons without lunula. Basoflagellomere small, brownish, arista rather long, almost bare (or with very short pile) (Fig. 9B). Scutum shining, rather densely punctated, all covered with erect yellow and black pile of even length, pile becoming slightly longer towards scutellum. Postalar callus with few black bristles. Scutellum with only yellow pile, margin without longer black bristles. Anepisternum and katepisternum with slight pollinosity, with yellow long pile. Ventral and dorsal pile patches on katepisternum narrowly connected posteriorly. Halter yellow. Wing with all cross-veins infuscated, completely covered with microtrichia. R_{4 + 5} of wing not curved. Legs dark, with apical (1/8) part of femora, and basal 1/2 and apical 1/6 of tibiae yellowish, tarsi yellow, fifth tarsal segment dorsally blackish or brownish. Mesofemur posteriorly with white pile, metafemur anteroventrally and -dorsally with long white pile, and ventrally with shorter (stronger) black pile. Abdomen shining laterally, mid parts of tergites I–III with pollinosity. Tergites with yellow erect pile, longer on the sides and shorter medially. Sternite I pollinose, other sternites shining, pile yellow, erect. Hypopygium as on Fig. 9E–G.

♀: Face and parafacia as in the male. Frons coarsely punctated, with white erect pile, and also with black erect pile on posterior part (Fig. 9D). Parafacia < 1/2 the width of basoflagellomere. Basoflagellomere reddish (Fig. 9C). Postalar callus with no black bristles. Scutum shining, with yellow and more scattered black pile of even length. Abdomen shining, tergite I and anterio-medial part of tergite II with slight pollinosity. Otherwise as the ♂.

Size. Body length 10–11 mm.

Serbia 1 ♂, 1 ♀ ’Serbija, Klisura peka, 3.V.1993, leg. Milankov’ [FSUNS].

Montenegro, Romania, Serbia*.

“Oesterreich” [sic] [Austria].

C. laeviventris: Lectotype, ♂, pinned, with genitalia dissected in microvial, in ZMHU, here designated to fix the concept of Cheilosia laeviventris Loew and to ensure the universal and consistent interpretation of the same. The original description is only for the male sex. The lectotype is labelled: ‘Austria, Schiner’ [handwritten, faded inc], ‘laeviventris’ [handwritten, faded inc], ‘coll. H. Loew’, ‘11469’, ‘Cheilosia laeviventris m.’ [handwritten, faded inc], ‘Cheilosia laeviventris Loew, Ståhls & Barkalov des.’.

C. primulae: Lectotype, ♂, ZMHU, here designated to fix the concept of Cheilosia primulae Hering and to ensure the universal and consistent interpretation of the same. The lectotype is labelled: ‘Ebenstein, Hochschwabgruppe, leg. H. Franz’, ‘Cheilosia primulae m. det. Hering 1944’ [handwritten, faded inc], ‘Zool. Mus. Berlin’, ‘Lectotype Cheilosia primulae Hering, Ståhls & Barkalov des.’. Genitalia dissected. A female paralectotype, idem. Both C. primulae are reared from larvae found in Primula auricola L. and the pupae are mounted on the same pin. The lectotype is forced out from the pupa: left wing missing, right wing incomplete, only with right hind leg, face slightly demolished.

Description. ♂: Face in frontal view moderately divergent from level of antennal insertion to lower mouth edge, with pollinosity, lower part of face clearly protruded (Fig. 10A). Parafacia pollinose, moderate in width, approximately 2/3 of width of basoflagellomere. Frons shining, with black pile. Frontal angle slightly > 90°. Lunula dark-brown to black. Antennal pits separated or very narrowly connected. Vertical triangle with black pile. Eye-contiguity nearly equal to or slightly shorter than length of frons without lunula. Basoflagellomere small, rounded, reddish brown to dark-brown (Fig. 10B). Arista moderately long, with short pilosity. Scutum and scutellum shining, densely punctated, with erect, mixed yellow and black pile, of about equal lengths. Central part of scutum with a rather broad transversal band of black pile. Hind margin of scutellum with black bristles or bristle-like pile. Postalar callus with some black bristles. Pleura slightly pollinose, pile predominantly black or mixed. Katepisternum with dorsal and ventral hair patches only narrowly connected anteriorly. Legs black, knees sometimes very narrowly yellowish. Apical half of posterior surface of mesofemur with long black pile, and basal part of anterior surface of metafemur with short black pile. Wing completely microtrichose, with all crossveins infuscated, R_{4 + 5} not distinctly curved (Fig. 10H). Abdomen shining with erect, yellow longer pile on lateral parts of tergites, and erect, black pile on central parts of tergites II–III, IV tergite with depressed pile. Tergite I–II with pollinosity medially on all length of tergite, tergite III with pollinosity only narrowly anteriorly or on all length of tergite. Tergite II often with a patch of black pile laterally. Sternites shining. Hypopygium as on Fig. 10E–G.

♀: Face as in the ♂. Frons with pollinosity laterally, otherwise shining, pile mixed black and white (Fig. 10D). Lunula brown. Basoflagellomere small, rounded, brown to dark-brown (Fig. 10C). Arista with short pile. Scutum shining, pile as in male. Otherwise as the ♂.

Size. Body length 8–11 mm.

Austria 1 ♂, 2 ♀ ‘Österreich, Tirol, Seefelder Joch, 2050 m, 15.VII.1969, v. d. Goot & Lucas’ [ZMA]; Germany 5 ♂ ‘Alpen Oberstdorf, Nebelhorn, Koblat, 1 920–2 220 m, 4.7.1994, D. Doczkal’ [coll. D. Doczkal].

Austria, France, Germany, Italy, Romania, Switzerland.

‘Oesterreich’ [sic] [Austria].

Lectotype ♂, pinned, with genitalia dissected in microvial, in MNB, here designated to fix the concept of Cheilosia venosa Loew and to ensure the universal and consistent interpretion of the same. The original descriptions is only for the male sex. The lectotype is labelled: ‘Austria, Schiner’ [handwritten, faded inc], ‘Coll. H. Loew’, ‘Zool. Mus., Berlin’, ‘Lectotype Cheilosia venosa Loew, Ståhls & Barkalov des.’.

♂: Face in anterior view moderately divergent from level of antennal insertion to lower mouth edge, with lower part not very prominent, with band of pollinosity (Fig. 11A). Facial knob prominent (nose-like). Parafacia with pollinosity, on lower part shining patch, slightly more than 1/2 the width of basoflagellomere. Frons shining, rather swollen, with black and white pile. Frontal angle ≈ 90°. Lunula dark-brown to blackish. Antennal pits confluent (occasionally only narrowly confluent). Vertical triangle with black pile. Eye-contiguity nearly equal to length of frons without lunula. Basoflagellomere small, rounded, brown to reddish brown. Arista short, thickened, clearly pubescent (Fig. 11B).

Scutum densely punctated, shining, with long, erect yellow pile, medially a transverse band of predominantly black pile. Scutellum on hind margin without (occasionally with some) black bristles. Postalar callus with two bristles. Pleura pollinose, with long black and yellow pile. Dorsal and ventral pile patches of katepisternum only narrowly connected anteriorly. Legs black, knees sometimes yellow, tibia sometimes black ringed. Apical half of posterior surface of mesofemur and basal part of anterior surface of metafemur with long black pile (Fig. 12A). Halter yellow. Wing with all crossveins infuscated, R_{4 + 5} distinctly curved (Figs 11I, 12A). Wing completely microtrichose. Abdomen shining, with pollinosity on middle part of tergites I–III. Tergites with long, erect, yellow pile laterally, and black short erect pile medially. Sternites I pollinose, II–IV shining. Hypopygium as on Fig. 11F–H.

♀: Face shining with fine pollinosity on the sides. Parafacia comparatively narrow, pollinose. Frons moderately broad, shining (Fig. 11D), with yellow and black pile (Fig. 11E). Basoflagellomere, small, rounded, black to brownish, with sensory pit (Fig. 11 C). Arista short, with dense, short pile, almost on whole length of arista. Scutum shining, with long, erect yellow and black pile of equal length. Pleura slightly pollinose. Scutellum margin without black bristles. Otherwise as the ♂.

Size. Body length 9–11 mm.

Additional material studied. Austria 1 ♂ ‘Österreich, Tirol, Seefelder Joch, 2 050m, 15.VII.1969, v.d. Goot’ [in ZMA], 1 ♂ ‘Austria, Brauer’ [MNB], 1 ♂ ‘Austria, Schiner’ [MNB]; Switzerland 1 ♀ ‘Helvetia, GR, Lenzerheide, Parpaner Rothorn, 2 850 m, 14.VII.96, B. Merz’ [ETH]; Italy 1 ♀ ‘Südtirol, Val di Planol, 2 000–2 400 m, 7.VII.1988, C. Claussen’ [in MZH], 1 ♀ ‘Stelvio Pass, 1 600–2 000 m, 22.VII.1988, Daccordi’ [in MZH], 1 ♀ ‘Valle Aurina, Bolzano, Val d. Vento, 2 400 m, 11.8.1987, Daccordi’ [in MZH].

Austria, Italy*, Germany, Romania, Switzerland*.