Two new species of terrestrial frogs of the Pristimantis gladiator complex (Anura, Strabomantidae) from the Ecuadorian Andes, with insights on their biogeography and skull morphology

Dorsolateral and ventral views in life of Pristimantis gladiator species complex and related species A Pristimantis donnelsoni sp. nov. female, DHMECN 14701, holotype, LRC: 15.42 mm B Pristimantis donnelsoni sp. nov. male, DHMECN 18854, LRC: 13.66 mm from Cerro Candelaria C Pristimantis kayi sp. nov. female, ZSFQ 0775, holotype, LRC: 19.7 mm D Pristimantis kayi sp. nov. male, DHMECN 14450 LRC: 13.58 mm, from Reserva Machay E Pristimantis gladiator sensu stricto female, DHMECN 12463, LRC: 20.38 mm F Pristimantis gladiator sensu stricto, male DHMECN 12474, LRC: 16.87 mm; from Río Quijos drainage G Pristimantis festae female, DHMECN 16280, LRC: 20.19 mm H Pristimantis festae, male (left), DHMECN 16837, LRC: 16.50 mm; female (right), DHMECN 16836, LRC: 19.80 mm; Photographers: JPRP (A, B), MYM (D, E, F, H); Patricia Bejarano (G); José Vieira (C).