Two new species of terrestrial frogs of the Pristimantis gladiator complex (Anura, Strabomantidae) from the Ecuadorian Andes, with insights on their biogeography and skull morphology

Colour variation of Pristimantis kayi sp. nov. A MZUTI 2209, adult female, LRC: 20.15 mm, from Cordillera de Guacamayos B MZUTI 2210, adult female, LRC:19.90 mm, from Cordillera de Guacamayos C MZUTI 2211, adult male, LRC: 14.30 mm, from Cordillera de Guacamayos D MZUTI 2213, adult male, LRC: 15.10 mm, from Cordillera de Guacamayos E MZUTI 2214, adult female, LRC: 18.73 mm, from Cordillera de Guacamayos F ZSFQ 774, adult female, LRC:18.96 mm from Sumaco Volcano G ZSFQ 782, adult female, LRC: 30.03 mm, from Sumaco Volcano. Photographs by Lucas Bustamante (A, B, C, D, E) and Jose Vieira (F, G).