Corresponding author: Simon van Noort (
Academic editor: B. Santos
We describe
Rousse P, Broad GR, van Noort S (2016) Review of the genus
“To find the details of the life history of [
Natural History Museum, London, UK (Gavin Broad & David Notton)
Muséum National d’Histoire Naturelle, Paris, France (Claire Villemant & Agnièle Touret-Alby)
Iziko South African Museum, Cape Town, South Africa (Simon van Noort)
Museum für Naturkunde, Humboldt Universität, Berlin, Germany (Frank Koch & Viola Richter)
Zoologische Staatsammlung, München, Germany (Stefan Schmidt)
At
(updated after
The gradual curve of the propodeum in profile is typical of the tribe
Interestingly, Tom Huddleston’s note in the
Online Lucid identification keys are available at:
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1 | Male: known only for |
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– | Female (a, b) |
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2 | Metasoma yellowish-orange with tergite 1 dark reddish-brown (A); hind femur and tibia yellowish-orange (A); antennal scrobe extends to eye margins (B) |
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– | Metasoma uniformly dark reddish-brown (a); hind femur and tibia dark reddish-brown (a); antennal scrobe less extensive, ending an ocellar diameter from eye margins (b) |
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3 | Frons with pronounced, horn-like projections pointed upward (A, B) |
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– | Frons with a bisinuate transverse lamella (a), that may have minor projections (b) |
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4 | Lower gena produced into a long spade-like protrusion (A, B); clypeus exceptionally expanded (A, B) |
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– | Lower gena not expanded (a), or distinctly less expanded into a backward curved collar (b) |
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5 | Clypeus apically slightly convex, gena moderately protruded (A); metasoma brownish-black (B) |
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– | Clypeus apically deeply emarginate, gena dramatically protruded (a); metasoma yellowish-orange (b) |
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6 | General coloration black (A); lower gena expanded into a backward curved collar (B) |
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– | General coloration dark red (a); lower gena not expanded (b) |
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7 | Frons with a forked, horn-like projecting carina above the antennal scrobes (A); second tergite medially punctate (B) |
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– | Frons with a broad to acute cordate lamella above the antennal scrobes (a); second tergite medially longitudinally strigose (metasoma lacking in ♀ allotype of |
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8 | Metasoma, hind femora and tibiae yellowish-orange (A); frons with a broad cordate lamella above the antennal scrobes (B) |
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– | Metasoma, hind femora and tibiae reddish (a); frons with an acute cordate lamella above the antennal scrobes (b) |
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9 | Lower gena expanded into a backward curved, bluntly triangular, protuberance (A); wings hyaline (B) |
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– | Lower gena not expanded into a backward curved protuberance (a); wings infuscate (b) |
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The uniquely shaped cordate, sublamelliform raised carina dorsally demarcating the scrobe readily separates females of this new species from all the other described species, each of which has a diagnostically shaped horizontal carina, with various uniquely shaped projections in this region. The male is distinguishable from the only other known male (
From the latin “cossus” for “worm or grub found in wood”, which is the likely host (
FEMALE.
MALE. Yellow markings more expanded, encompassing also mandible, clypeus, median face, genal orbits and pronotal collar; flagellum not widened with 33 flagellomeres, flagellomeres 7–14 with tyloids on outer surface and 6–32 with differentiated bristle mid-ventrally; lower frons transversely striate, transverse carina strongly attenuated; parameres simple; otherwise similar to female.
Reared from logs of
South Africa.
The unique clypeus, which is deeply emarginate with long, sharp corners, and the exceedingly long, strongly sculptured genal protuberance, makes this a distinctive species, with the most extreme head ornamentation that we have seen in
The name is derived from the genus name for warthogs, after the resemblance of the head to that of a warthog. Noun in apposition.
FEMALE.
MALE. Unknown
Unknown, but see discussion on biology under the genus treatment.
Tanzania.
Amongst
The name is derived from the Latin for "smoky", in reference to the infuscate wings. Noun in apposition.
FEMALE.
MALE. Unknown
Unknown, but see discussion on biology under the genus treatment.
South Africa.
Associated with
Kenya.
Unknown, but see discussion on biology under the genus treatment.
Kenya.
Unknown, but see discussion on biology under the genus treatment.
Uganda.
Unknown, but see discussion on biology under the genus treatment.
South Africa.
A female from South Africa (Natal: Kloof, 1500ft, VIII.1926, R.E. Turner) in
Unknown, but see discussion on biology under the genus treatment.
South Africa, Tanzania.
Thanks to Brett Hurley and Bernard Slippers (Tree Protection Co-operative Programme (TPCP) and NRF/DST Centre for Tree Health Biotechnology (CTHB), University of Pretoria) for making the reared specimens available to us. We are grateful to the curators and collections managers of the above mentioned museums for loaning specimens. Simon van Noort was funded by South African NRF (National Research Foundation) grants: GUN 81139; GUN 98115. Pascal Rousse was funded by SABI (South African Biodiversity Initiative) NRF post-doctoral fellowship GUN 81609, Claude Leon Foundation post-doctoral fellowship, and Société Entomologique de France (Bourse Germaine Cousin). We are grateful to the reviewers for their useful comments, especially Alexander Tereshkin, who drew our attention to