Pariaconus oahuensis and Pariaconus ohiacola immatures. A–DP. oahuensis: A 5th instar with anal ring detail (inset) B tarsi C antenna D anal ring EP. ohiacola anal ring F–H 1st instar: FP. oahuensis with dorsal tubercles detail (inset left) and marginal sectasetae detail (inset right) GP. ohiacola form ohiacola with marginal fan-shaped setae detail (inset) HP. ohiacola form angustipterusI–LP. oahuensis galls: I bud gall J, K, L cone galls on leaves opening with valves on lower leaf surface M–RP. ohiacola: M flat leaf galls open by valves on lower leaf surface N variation in discolouration of gall tissue O circular sutures in gall centres from 1st instar pits on lower leaf surface with detail (inset) P distribution of flat leaf galls Q 1st instar seated in pit before leaf tissue enclosure makes enclosed gall R 5th instar dissected from closed flat leaf gall.