Corresponding author: Alexander Anichtchenko (
Academic editor: A. Casale
Species of the genus
This study aims to clarify the circumscription of the subtribe
The most recent catalogue covering the study area (
This study was based on a total of about 270 specimens, including a number of primary types. Measurements: body length, from anterior margin of clypeus to apex of elytra along suture; length of pronotum, along midline; width of pronotum, at widest point; length of elytra, from his base to apex along suture; and width of elytra, at widest point.
The material from the following institutional and private collections has been examined:
Alexander Anichtchenko private collection (Spain)
Andreas Pütz private collection (Germany)
Daugavpils University Beetle Collection (Latvia)
Dawid W. Wrase collection (Germany)
Institute of Zoology Collection (Armenia)
K. Orszulik private collection (Czech Republic)
Moravske Muzeum, Brno (Czech Republic)
National Museum of Natural History in Paris (France)
Collection of Zoology & Ecology Department, Moscow State Pedagogical University (Russia)
Naturhistorisches Museum Wien (Austria)
National Museum of Natural History, Prague (Czech Republic)
Ron Felix private collection (The Netherlands)
Natural History Museum ‘Naturalis', Leiden (The Netherlands)
Staatliches Museum für Naturkunde Stuttgart (Germany)
United Arab Emirates Invertebrate Collection (UAE)
Zoological Institute, St. Petersburg (Russia)
Zoological Museum of Moscow University (Russia)
Zoologische Staatssammlung München (Germany)
Highresolution habitus images of
The exact composition of this subtribe has not been settled. Debate on the taxonomic status of
The recent catalogues (
Now we are not in position to give diagnostic features of this genus, for we do not know the other related genera well enough. The limits of these generic groups are not yet defined. Details of relationships among genera and subtribes remain to be worked out. In lieu of a definitive treatment of classification of
The genus
AFGHANISTAN: ♂ – TYPE, red label: Agatus afganus sp. n. det. Ing. Jedlička; white label: Afganistan, Bhougavi; Cotype ♂, Afghanistan, Douchi; Cotype ♀, Afghanistan, Dehran (NMPC). KAZAKHSTAN: S Kazakhstan, Karatau Mt. Rng., 40 km N Igilik vill., Kurkal,
This species is most similar to the sympatric
Habitus of
Length 4.3–5.3 mm. Body elongate; head and pronotum redbrown, elytra black with redbrown basal band and a common preapical round red macula reaching interval 6. Venter light brownish yellow, sterna sometimes darker; legs yellow (
Head microsculptured, deeply and irregularly punctate, punctures sometimes almost confluent in frontal depressions, on the frons separated by 3–5 their diameters, towards the base of head, by 1–2 diameters. Clypeus with a few punctures at lateral margins. Eyes large and bulging, with numerous strong setae at posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples smooth, 2.4 times shorter than length of eye. Scape with a very long seta and a few long thin setae; pedicel with a band of apical setae; antennomere III with scattered setae on apical half. Antennae pubescent from the midlength of antennomere IV.
Pronotum shinier than head, microsculptured, cordate, 1.08–1.11 times as wide as head, 1.2–1.25 times as wide as long, widest just behind the marginal setae. Anterior margin straight, anterior angles effaced, sides slightly sinuate towards posterior angles, which are rectangular or slightly acute. Disc sparsely irregularly punctate, somewhat less densely than head; punctures at lateral margins confluent, surface rugose. Disc convex to lateral margins, which are very narrow. Posterior pore right in front of angle. Basal grooves shallow, rugose, confluently punctate. Pronotal base extended in a rounded median lobe. Furrow short and shallow.
Each interval with a row of setiferous pores along the middle from base to apex. Setae as long as the width of interval 2. Microsculpture almost isodiametric. Apices slightly sinuate. Striae deep, punctate. Intervals weakly convex at base, flat at apex.
Legs brownish yellow. Tarsomere V with 4 pairs of ventral setae. Propleuron smooth, even, mes and metepisterna strongly microsculptured. Claws with 4 teeth, 3 relatively long and one (basal) minute. All abdominal sterna with long pubescence.
Aedeagus –
Varies in size, ventral colour, and the extent of dark elytral pattern (may be reduced).
Types collected in “Samarkand" (now Uzbekistan) and near Shahrud (n. Iran) (Solsky, 1874: 35).
Afghanistan, Iran, Kyrgyzstan, Kazakhstan, Tajikistan, Turkmenistan, Uzbekistan.
HOLOTYPE, 1 golden foil circle, 1 handwritten label: “cingulatus Gebl. Sibir. Or." (indeed = Alakul') (ZIN); AFGHANISTAN:
This species is most similar to
Length 4.5–5.2 mm. Body elongate, head and pronotum redbrown, elytra black with redbrown basal band and two rounded preapical spots, often confluent. Femora and tibial apices piceous to black (
Head microsculptured, deeply irregularly punctate, punctures sometimes almost confluent in frontal depressions, on the frons separated by 3–5 times their diameters, towards the base of head, by 1–2 diameters. Clypeus with a few punctures at lateral margins. Eyes large and bulging, with numerous moderately long setae at posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples smooth, 4.2 times shorter than length of eye. Scape with a very long seta and a few long thin setae; pedicel with a band of apical setae; antennomere III with scattered setae on apical half. Antennae pubescent from the midlength of antennomere IV.
Pronotum shinier than head, microsculptured, cordate, 1.07 times as wide as head, 1.25–1.31 times as wide as long, widest just behind the marginal setae. Anterior margin straight, anterior angles effaced, sides slightly sinuate towards posterior angles, which are rectangular or slightly acute. Disc sparsely irregularly punctate, somewhat less densely than head; punctures at lateral margins confluent, surface rugose. Disc convex to lateral margins, which are very narrowly explanate. Posterior pore right in front of angle. Basal grooves shallow, rugose, confluently punctate. Pronotal base extended in a rounded median lobe. Furrow shallow and short.
Each interval with a row of setiferous pores along the middle from base to apex. Setae as long as the width of interval 2. Microsculpture almost isodiametric. Apices slightly sinuate. Striae deep, punctate. Intervals weakly convex at base, flat at apex.
Femora and apical parts of tibiae piceous to black. Tarsi sparsely pubescent, with long dorsal setae. Tarsomere V with 4 pairs of ventral setae. Propleuron rugose, mes and metepisterna strongly microreticulate. Claws with 5 teeth. Abdominal sterna black, venter otherwise redbrown. Abdominal sterna with long pubescence.
Aedeagus –
The study of the holotype of
Afghanistan, Iran, Iraq, Israel, Kyrgyzstan, Kazakhstan, s. Russia, Tajikistan, Turkmenistan, Uzbekistan.
Species described from a single specimen labeled “Sibiria" (from collection of Eversmann) as similar to
Syntypes: Turkestan, W. Balassoglo leg. (1♂ 1♀, ZIN); TURKMENISTAN: SW Turkmenistan, BekiBent, 15.V.1952, Romadina leg. (1♂, ZIN); KaraBogaz, 40 km N KizylArvat, 14.IV.1952, Il'ichev leg. (1 ex., ZIN); W Kopetdag, 15 km S Iskender, 15.V.52, Kir'janova leg. (1♀, ZIN); SW Turkm., vall. of Divan, 15 km W Chat, 4.V.1952 (1♀, ZIN); Turkm., 13 km S KizilArvat, 25.IV.52 (1♂, ZIN); Chardzhou, AmuDar'ja riv., 8.VIII.1910, N. Androsov leg. (2♂♂ 2♀♀, ZIN); Kushka, Zakaspiisk reg., 16.VI.1912, leg. V. Kozhanchikov (2♂♂, ZIN); Kushka, 29.VII.1910 (3♂♂ 5♀♀, ZIN); Turkm., EroilanDuz, 1.V.1968, G. Medvedev leg. (1♀, ZIN); Tamdytau, Aktau, 4.V.1965, Medvedev leg. (1♂, ZIN); Turkm., KaraKala, Sjumy, IX.1931 (1♂, ZIN); Turkmenia (1♀, NMPC). UZBEKISTAN: Termez, Buhara, 3.VII.1912, Kirichenko leg. (1♀, ZIN).
Instantly recognizable by the combination of cordate pronotum, elongate and flat elytra, intervals with single uninterrupted row of conspicuous setiferous pores bearing long setae. In sympatric
Length 5.9–6.2 mm. Redbrown, with wide piceous postmedian band on elytra (
Head smooth, very distinctly microsculptured, finely punctulate, punctures separated by 2–4 diameters. Clypeus with few punctures at base. Eyes normal, with 3–5 setae at posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples long, smooth. Scape with a very long subapical seta and several short setae; pedicel and antennomere III each with a single band of apical setae. Antennae pubescent from the midlength of antennomere IV.
Pronotum cordate, smooth, 1.1 times as wide as head and 1.27 times as wide as long, widest right behind marginal setae. Anterior margin straight, anterior angles effaced, sides strongly sinuate towards base. Posterior angles acute, prominent. Lateral margin narrow. Discal punctation sparse and shallow, somewhat sparser than on head. Posterior pore right in front of angle. Lateral and basal setae long. Basal grooves punctiform. Pronotal base rugosely punctate, extended in a rounded median lobe. Microsculpture as on head.
Each interval of elytra with a single regular row of long sparse setae, interval 8 with two rows of setae. Setae as long as the width of interval 3 at its widest. Pores deep, punctiform, separated by three diameters. Microsculpture of elytra and scutellum irregularly polygonal, shallower than on head and pronotum. Elytral apices weakly obliquely sinuate, rounded at suture. Striae deep and punctate. Intervals almost flat throughout.
Legs pale brownish. Tarsomere V with 4 pairs of ventral setae, two basal pairs very short. Claws with 3 denticles on basal half. Venter entirely brownish yellow. Episterna smooth. All abdominal sterna with long pubescence.
Holotype: ♀, red label:
This species is most similar to the sympatric
Length 6.8–8.0 mm. Head, pronotum, basal third of elytra, thorax, legs, and antennae redbrown; suture, apical 2/3 of elytra, and abdominal sterna black (
Head microsculptured, deeply irregularly punctate, punctures near eyes sometimes almost confluent, on the frons separated by more than three diameters and on the vertex by 1–2 diameters. Clypeus sparsely punctate. Eyes large and bulging, with numerous strong setae at posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples smooth. Scape with a very long seta and a few short thin setae scattered throughout; pedicel with a band of apical setae; antennomere III with numerous setae on apical half. Antennae pubescent from the basal third of antennomere IV.
Pronotum shinier than head and elytra, 1.18 times as wide as head, 1.34 times as wide as long, widest in front of marginal setae. Anterior margin straight, anterior angles effaced, sides very broadly and evenly rounded, slightly sinuate towards posterior angles, which are rectangular. Disc shiny, very sparsely and irregularly punctate, punctures separated by 2–5 diameters, transversely confluent towards lateral margins. Lateral margins narrow, flat towards base. Posterior pore right in front of angle. Basal grooves shallow, rugose, confluently punctate. Pronotal base extended in a rounded median lobe. Furrow fine and reduced. Microreticulation faint.
Elytra pubescent. Each interval with 1–2 irregular rows of setiferous pores from base to apex. Setae as long as mesotarsomere IV. Interval 7 flat, as wide as adjacent intervals. Microsculpture polygonal, deep and irregular, same as on head. Apices slightly sinuate. Striae slightly punctate, shallower on disc and at apices. Intervals slightly convex at base, flat at apex.
Tarsomere V with 5 pairs of ventral setae. Propleura rugose. Mes and metepisterna smooth. Claws with 5 teeth, basal teeth very small. All abdominal sterna pubescent; pubescence long and dense, as on elytra.
Aedeagus –
Afghanistan, Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan.
This species was described as
Named after the Russian entomologist Semyon Solsky who first described this species.
Holotype: ♂, Tajikistan, Petr 1 mt. rng., western Sangvor, LuliHarvi, deciduous forest, 23.VIII.1975, leg. V. Yanushev (MPU). Paratype: ♀, same label (MPU).
This species is most similar to
Length 6.0 mm. Head, pronotum, and legs redbrown. Venter redbrown, except black abdominal sterna. Elytra black, with basal third redbrown (
Head densely punctate, very distinctly microsculptured and pubescent; pubescence as long as on pronotum and elytra, and equal to the width of the sutural elytral interval. Punctures near eyes sometimes almost confluent, on the frons separated by over twice their diameter. Clypeus smooth, very distinctly microsculptured. Eyes large and bulging, with 4–5 long setae at posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples very short, smooth. Scape with a very long subapical seta and several more, rather long, setae; pedicel with a band of setae; antennomere III with two bands of apical setae. Antennae pubescent from the basal third of antennomere IV.
Pronotum evenly setose, 1.22 times as wide as head, 1.36 times as wide as long, widest near marginal setae. Anterior margin weakly emarginate, anterior angles effaced, sides broadly evenly rounded, sinuate towards posterior angles, which are acute and protrude as minute denticle. Punctation uniform and deep, somewhat shallower than on the head, especially on the disc of pronotum. Lateral margin moderately explanate in basal half only. Posterior pore right in front of angle. Basal grooves shallow, small, punctiform. Pronotal base extended in a rounded median lobe. Furrow fine, not reaching apical and basal margins. Microsculpture as on head.
Elytra: All intervals with confused punctation, conspicuous polygonal microsculpture and long, pale pubescence. Apices weakly obliquely sinuate, rounded at suture. Striae deep and crenulate. Intervals almost flat on disc, slightly convex at base and flat towards apex. Scutellum smooth.
Tarsomere V with 4–5 pairs of ventral setae. Claws with 5 teeth, of which apical three very long. Prosternum, propleura and mesepisterna smooth. Abdominal sterna pubescent.
Aedeagus –
Habitus of
Named after Dr. Kirill V. Makarov.
Tajikistan.
Holotype: ♂, Afghanistan, Herat, NW Adraskan, 20.XI.1971, leg. Kabakov (ZIN). Paratypes: Afghanistan, Ghazni, W Moqur, 2300 m, 11.IX.1972, leg. Kabakov (1♀, ZIN); Iran, Khorasan, TorbateHeydariyeh, 5 km S Zharf, 14.V.2007, leg. A. Anichtchenko (1♂, AAC); small green rectangle and a white label “Christoph collection" (in Russian) meaning Schahrud, Persia 1870–1873 (1♀, ZIN).
This plus
Length 5.2–5.8 mm. Redbrown, with postmedian sutural infuscation on elytra (
Head smooth, microsculptured, uniformly punctate, punctures separated by 1–2 their diameters. Clypeus impunctate. Eyes moderately convex, with a few relatively long setae at posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples short and smooth. Scape with a very long seta at 2/3 of its length and a few thin setae towards the apex; pedicel with a single, apical band of setae; antennomere III with a few additional setae towards apex. Antennae pubescent from the basal fourth of antennomere IV.
Pronotum smooth, shinier than head, 1.14 times as wide as head, 1.2 times as wide as long, widest at marginal setae. Anterior margin straight, anterior angles slightly prominent, sides very broadly and evenly rounded, conspicuously sinuate towards posterior angles, which are acute. Punctation irregular, sparser and shallower than on the head. Explanate lateral margin rapidly widened from apex, broad and flat to slightly reflexed at base. Lateral and apical setae very long. Posterior pore right in front of angle. Basal grooves punctate. Pronotal base extended in a rounded median lobe. Furrow variable. Microsculpture subtle, slightly transversely polygonal.
Elytra infuscate behind middle, infuscation may reach just one or two innermost intervals, or extend to lateral margins. Intervals 1, 3, 5, 7, 8 pubescent throughout, and interval 6 pubescent in apical half, each with an irregular row of large setiferous pores. Setae as long as the width of interval 4 at its widest. All intervals convex at basal third and slightly convex from there to apex. Interval 7 convex from base to midlength. Microsculpture deep, irregularly polygonal. Apex slightly sinuate. Striae deep, punctate, shallower at apex.
Legs brownish yellow. Tarsomere V with 3 pairs of ventral setae. Mes and metepisterna smooth. Claws with 4 moderately long teeth. Venter evenly brownish yellow. All abdominal sterna pubescent, pubescence as long as on elytra.
Aedeagus –
Habitus of
Named after Oleg Kabakov, a famous Russian coleopterist.
Afghanistan, Iran.
Holotype: ♀, Iran, Kerman, Sirjan, 8 km N Balvard, 11–12.V.2007, leg. A. Anichtchenko (ZIN).
Among the species consisting of smallbodied beetles, with convex, subovate elytra and long pubescence,
Length 4.6 mm. Uniformly redbrown, with venter and legs paler (
Head smooth, distinctly microsculptured, very sparsely minutely punctulate, punctures sometimes barely visible. Clypeus and labrum distinctly microsculptured. Eyes moderately large, each with 3 long setae behind. Second supraocular seta located immediately basad the posterior margin of eye. Temples long, smooth. Scape with a very long subapical seta and several small setae; pedicel and antennomere III each with a single band of apical setae. Antennae pubescent from the midlength of antennomere IV.
Pronotum smooth, cordate, 1.07 times as wide as head, 1.3 times as wide as long, widest just behind marginal setae. Anterior margin straight, anterior angles effaced, sides broadly and regularly rounded, slightly sinuate towards base, posterior angles obtuse. Disc very sparsely punctate. Anterior margin with a few big punctures, basal part with large punctation reaching along lateral margins 1/3 of pronotal length. Lateral margin narrowly explanate from apex, slightly wider at base. Posterior pore right in front of angle. Basal grooves shallow. Pronotal base extended in a rounded median lobe. Furrow long and fine. Microsculpture as on the head.
Elytra suboval, convex, with strong polygonal microsculpture. Each interval with a series of very large setiferous pores in a single row. In the discal part of intervals 2–5 pores may be smaller or rows interrupted. Setae as long as the combined width of intervals 1 and 2. Scutellum smooth and shiny, without microsculpture. Elytral apices truncate, weakly obliquely sinuate, rounded at suture. Striae crenulate at base, finely punctate elsewhere. Intervals slightly convex.
Legs pale brownish yellow. Tarsomere V with 2 pairs of short ventral setae. Claws with 4 minute denticles near base. Abdominal sterna smooth, shiny, with sparse short setae.
Iran.
Holotype: ♂, SE Iran, Sistan, 100 km SE Zahedan, Tamin, 2100m, 3.V.2006, leg. A. Klimenko (ZIN); Paratype: ♂, same label (AAC).
Among the quartet of species consisting of smallbodied beetles, with subovate elytra and long pubescence (
Length 4.7–5.1 mm. Uniformly yellowbrown, legs paler (
Head smooth, very distinctly microsculptured, finely punctulate, punctures separated by 2 to 4 diameters. Clypeus impunctate. Eyes large and bulging, with 2–3 short setae at posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples short, smooth. Scape with a very long subapical seta and several short setae; pedicel and antennomere III each with a single band of apical setae. Antennae pubescent from midlength of antennomere IV.
Pronotum smooth, 1.08 times as wide as head, 1.38 times as wide as long, widest right behind marginal setae. Anterior margin straight, anterior angles effaced, sides almost straight basad the widest point. Posterior angles obtuse. Lateral margin narrow, 2–3 transverse wrinkles at base. Discal punctation sparse and shallow, somewhat sparser than on head. Posterior pore right in front of angle. Lateral and basal setae long. Basal grooves punctiform. Pronotal base extended in a rounded median lobe. Furrow deep and short. Microsculpture as on head.
Elytra shinier than head and pronotum. Each interval with a single irregular row of sparse setae; setae longer than the width of interval 3 at its widest. Setiferous pores large and flat, sparser on intervals 3 and 5 than elsewhere. Microsculpture polygonal, more delicate than on head and pronotum. Apices weakly obliquely sinuate, rounded at suture. Striae very finely punctate. Intervals slightly convex at base and flat at apex. Scutellum microsculpture as on elytra.
Tarsomere V with 4 pairs of ventral setae, two basal pairs very short. Claws with 3 denticles in basal half. Venter brownish yellow throughout. Episterna smooth. Abdominal sterna with long pubescence.
Aedeagus –
Named after Alexei Klimenko who collected the specimens.
Iran.
Holotype: ♂, AmanKutan, merid. versus, ab Samarkand, 14.V.65 (ZMM).
Among the quartet of smallbodied species, with convex, subovate elytra and long pubescence (
Length 5.1 mm. Redbrown with apical half of elytra black; anterior margin of the dark area blurry and perpendicular to suture (
Head sparsely feebly punctate on the frons and between eyes, with punctures separated by 4 to 7 diameters, deeply and densely punctate elsewhere, with punctures separated by their diameter towards head base. Head and clypeus very distinctly microsculptured throughout. Clypeus smooth. Head pubescent, pubescence as long as on pronotum and elytra and as long as width of second interval. Eyes moderately large and bulging, with 4–5 long setae at posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples short, smooth. Scape with a very long subapical seta and several more, rather long, setae; pedicel with a band of setae; antennomere III with two apical bands of setae. Antennae pubescent from the basal third of antennomere IV.
Pronotum 1.22 times as wide as head, 1.33 times as wide as long, widest near marginal setae. Anterior margin straight, anterior angles effaced, sides broadly and regularly rounded, sinuate at base, posterior angles rectangular. Punctation confused and coarse, punctures large, shallower and larger towards the furrow and anterior angles. Disc convex, uniformly setose. Lateral margin slightly explanate in basal half only. Posterior pore right in front of angle. Basal grooves flat, densely punctate. Pronotal base extended in a rounded median lobe. Furrow deep, almost reaching anterior margin and ended short of the base by length of antennomere III. Microsculpture as on head.
Elytra subovate, with polygonal microsculpture. Each interval with a series of large setiferous pores in a single irregular row. Scutellum smooth. Elytral apices truncate, weakly obliquely sinuate, rounded at suture. Striae deep and slightly crenulate. Intervals slightly convex.
Legs redbrown. Protarsomere IV small and narrow. Tarsomere V with 4 pairs of ventral setae. Claws with 3 long apical teeth and one tiny denticle. Abdominal sterna black, pubescent throughout. Prosternum, propleura and mesepisterna smooth.
Aedeagus –
Named after Timur (Tamerlane), the fourteenthcentury conqueror of Asia.
Uzbekistan.
EGYPT: Aegyptus, Faggala (1♀, NMPC); Le Caire (1♀, MNHN). ISRAEL: AravaTal, S Zofar, 18.II.1987, 0 m, leg. Schawaller & Schmalfuss (1♂, SMNS). YEMEN: Lahj, X.2000, malaise trap, leg. A. v. Harten & A. Sallam (1♂ 1♀, SMNS).
Most similar in overall appearance and small size to
Length 4.5–4.8 mm. Winged. Ferrugineous or testaceous red with a dark preapical sutural spot reaching interval 4 or 5 (
Head obtuse, rather wide, slightly convex to flat between the eyes, with numerous large and deep punctures (mixed with wrinkles near eyes) that become sparser towards the neck. Frons without punctures. Eyes large and convex.
Pronotum punctate, slightly wrinkled, transverse, subconvex, considerably broader than head, strongly rounded in front, with anterior angles effaced. Sides rounded, with a long sinuation in front of posterior angles, which are acute. Pronotal margin broadly explanate, basal impressions fairly deep. Pronotal base projected at the middle, median furrow moderately wide and deep.
Elytra approximately 1/3 longer than broad, sides subparallel, almost obliquely truncate at apex which is weakly sinuate. Striae, including the scutellar stria, fairly deep and finely punctate; intervals weakly convex, with two pores on interval 3.
Microsculpture strong, especially on forebody where the cells are small and rather isodiametric, but cells become more transverse towards the sides of pronotum. Elytra with finer microsculpture of larger, subquadrate cells, and thus appearing shinier than the forebody.
In male, protarsomeres I–III slightly dilated.
Aedeagus –
Egypt (country record), Israel (country record), Saudi Arabia, Yemen.
Holotype, ♀, red label:
This species is most similar to
Length 4.3–5 mm. Yellowish redbrown, elytra with black pattern behind middle that varies from continuous band reaching lateral margins to small grey sutural spot (
Head microsculptured, deeply and uniformly punctate, punctures separated by about their diameter but denser in frontal depressions. Clypeus with few scattered punctures. Eyes large and bulging, with 4–5 very short setae at posterior margin. Temples short and smooth. Scape with a very long seta at 2/3 of its length and a few short thin setae towards the apex; pedicel with a band of apical setae; antennomere III with two bands of setae (at midlength and at apex). Antennae pubescent from midlength of antennomere IV.
Pronotum 1.21–1.27 times as wide as head, 1.42–1.45 times as wide as long, widest just behind the marginal setae. Anterior margin straight, anterior angles effaced, sides very broadly and regularly rounded, slightly sinuate towards posterior angles, which are rectangular or slightly acute, protruded as minute denticle. Discal punctation similar to that of head, but shinier due to more delicate microsculpture. Pronotum at apex and base rugulose and densely punctate. lateral margins narrowly explanate from anterior angle to lateral setae, then widened basally, broad and flat at base. Posterior pore anterad angle. Pronotal base extended in a rounded median lobe. Furrow short and fine.
Elytra in the middle 1.36–1.4 times as long as wide. Intervals 1, 3, 5 and 7 each with minute setae visible only in lateral view at high magnification, and two irregular rows of minute sparse setiferous pores from base to apex. Interval 7 convex in basal half and narrower than interval 6. Microsculpture polygonal. Apices slightly sinuate. Striae slightly punctate, shallower on disc and towards apex. Intervals slightly convex at base, flat at apex.
Legs brownish yellow. Tarsomere V with 5 pairs of ventral setae. Propleuron weakly rugose towards coxae. Mes and metepisterna smooth. Claws with 4 teeth. Venter brownish yellow. All abdominal sterna with short pubescence, no longer than mesotarsomere IV.
Aedeagus –
Habitus of
Varies in body size and extent of dark elytral pattern (complete band to faint spot).
Armenia (country record), Iran (country record), Israel (country record), Turkey.
AFGHANISTAN: S Jalalabad, S Agam 2000 m, 23.XII.1970, Kabakov leg. (1♀, ZIN); 20 km NO Kabul, 2000m, 10.IV.1970 Kabakov (1♂ 3♀♀, ZIN); same locality, 15.IV.1971 (1♂, ZIN); Ghazni W Moqur 2300m, 5.X.1972, Kabakov (1♂ 2♀♀, ZIN); same locality, 11.IX.1972 (1♀, ZIN); Kondahar, Ghbargay 2000 m, 20.III.1973, Kabakov (2♀♀, ZIN); Ghor Saghar 2500 m, 14.VIII.1970 (1♂, ZIN); Heart, NW Adraskan 26.X.1971 (1♂, ZIN); same locality, 20.XI.1071 (1♀, ZIN). IRAN: ChaharmahalvaBakhuyari prov., 10 km E Chaman Goli, 7–8.VI.2008, 3500m, Anichtchenko A. leg. (2♂♂, AAC); Kerman, Qohrud mts., 10 km E Korin, 3500m, 13.V.2007, Anichtchenko A. leg. (2♀♀, AAC); S Iran, Kerman prov., Balt area, Korin v., 17–19.V.2008, A. Klimenko leg. (1♂, AAC); Kerman prov., 15 km E Korin, 2800–3300m, KuheLalehzar mt., 17–19.V.2008, Anichtchenko A. leg. (1♂, AAC); Kerman, Sargad, 24–26.VI.1898, N. Zarudnyi leg. (1♀, ZIN); same locality, 25–27.VIII.1898 (1♂, ZIN); same locality, 30.IV.1901 (1♀, ZIN); Perse, BenderBouchir, Dr. Bussieres, 1905 (1♀, MNHN); S Iran, 65 km nördl. BandarAbbas, 30.III.1972, Exped. Mus. Vind. (1 ex, NHMW). IRAQ: Bagdad, coll. Kálalová (6♂♂ 4♀♀, NMPC); Bagdad, Irak, coll. Kálalová (4 ex., NMPC); Bagdad, Ex Musaeo H.W. Bates 1892, Museum Paris, 1952, Coll. R. Oberthür (2♂♂ 2♀♀, MNHN). PAKISTAN: NW Pakistan, prov. Swat, 71°90' L/ 35°70' B, Madyan, 1400 m, am Licht, 19.VI–4.VII.1971, leg. C. Holzschuh (2♂♂, NMPC); Northwest frontier Prov. Barseen,
Very similar in both external and aedeagal anatomy to
Length 5.0–6.3 mm. Yellowish brown, elytra behind the middle with piceous spot that may reach apical and lateral margins but sometimes not extended beyond interval 6 and often leaving elytral apices brownish red (
Head microsculptured and deeply irregularly punctate, punctures sometimes almost confluent in frontal depressions, on the frons separated by more than 3 diameters. Clypeus with a few punctures near base. Eyes large and bulging, with no setae at posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples short and smooth. Scape with a very long seta and a few short thin setae; pedicel with a band of apical setae; antennomere III with two bands of setae (at midlength and at apex). Antennae pubescent from the basal third of antennomere IV.
Pronotum pale yellow, sometimes shinier than head, 1.16–1.18 times as wide as head, 1.4–1.47 times as wide as long, widest just behind the marginal setae. Anterior margin straight, anterior angles effaced to slightly marked, sides very broadly and evenly rounded, slightly sinuate towards posterior angles, which are rectangular (normally) to subacute. Punctation, especially on disc, normally sparser and more delicate than on head; confluent and rugose towards lateral and posterior margins. Lateral explanate margin rapidly widens basad lateral setae, broad and flat at base. Posterior pore right in front of angle. Basal grooves shallow, punctate. Pronotal base extended in a rounded median lobe. Furrow variable. Microsculpture isodiametric.
Elytra 1.45–1.52 times as long as width in the middle. Intervals 1, 3, 5 and 7 each with minute setae and two irregular rows of pores from base to apex, often poorly visible. Even intervals may also bear a few setae in apical areas. Interval 7 convex and narrower than the adjacent intervals. Microsculpture nearly isodiametric. Apices slightly sinuate. Striae narrow and slightly punctate. Intervals slightly convex at base and flat behind the middle.
Legs brownish yellow. Metatarsomere V with 3–4 pairs of ventral setae. Propleura slightly wavy rugose towards sternum and coxae. Claws with 5 teeth. Venter brownish yellow throughout. All abdominal sterna pubescent; pubescence as short as on metatarsomere IV.
Aedeagus –
Varies in size, elytral pattern (dark spot may be reduced), shape of posterior pronotal angles (usually acute and prominent but may be rectangular and less prominent). In almost all specimens from Afghanistan the apical spot reaches elytral apices, and the series of punctures in intervals 5 and 7 are shorter. Pakistani specimens have more convex elytral intervals.
Afghanistan, Iran, Iraq (country record), Pakistan (country record), Turkey (country record), Turkmenistan.
HOLOTYPE: ♀, 5063 UAE, Hatta,
This species can be recognised easily by the wide and small size body (4.8–5.4 mm), strongly transverse, very densely punctate pronotum and uniformly yellow pale body colour.
Length 5.1–5.6 mm. Pale brownish yellow, with no dark elytral pattern (
Head pale yellowish redbrown, slightly darker than pronotum, very distinctly microsculptured and very coarsely and deeply irregularly punctate. Punctures sometimes almost confluent, sometimes separated by over two diameters. Clypeus twice as long as labrum. Eyes very large and bulging, with a few small setae at posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples very short, smooth. Scape with a very long subapical seta and several rather long setae; pedicel and antennomere III each with a single band of apical setae. Antennae pubescent from the midlength of antennomere IV. Antennomeres IV and V each slightly shorter than antennomere III. Antennomere V 2.5 times as long as wide at apex.
Pronotum 1.25 times as wide as head, 1.57 times as wide as long, widest just behind marginal setae. Anterior margin straight, anterior angles effaced, sides broadly and evenly rounded, slightly sinuate towards posterior angles, which are acute and form a minute denticle. Punctation coarse, irregular and deep, somewhat sparser than on head, especially on the disc. Punctures sometimes confluent, especially towards the sides of disc. Lateral explanate margin narrow at apex, rapidly widened basally. Posterior pore right in front of angle. Basal grooves shallow. Pronotal base extended in a rounded median lobe. Furrow very short and fine and does not reach anterior margin (by about the length of antennomere I) or the base. Microsculpture as on head.
Elytra generally concolorous with pronotum, with the apical third perhaps slightly darker. Intervals 1, 3, 5 and 7 with several minute, barely visible setae. Microsculpture lighter than on head and pronotum, so elytra seem shinier than head and pronotum. Scutellum with similar microsculpture. Elytral apices weakly obliquely sinuate, rounded at suture. Striae shallow, crenulate. Interval 8 convex over most of its length, slightly flattened towards apex; interval 7 convex only at shoulder; other intervals slightly convex near base and flat at apex.
Tarsomere V with 3 pairs of ventral setae. Claws with 4 teeth. Venter testaceous, abdomen slightly infuscated. Metathoracic process margined, with 4 long setae. Abdominal sterna pubescent; the last two also with scattered longer setae.
Habitus of
In the holotype, last abdominal sternum with 6 setae (3 on each side), irregularly placed. One of the paratypes has 4 setae (two on each side) and one has 5 setae: 3 on one side and 2 on the other.
Aedeagus –
UAE, Oman (country record).
GREECE: Attica, Dr. Krüper (1♂, NHMW); Graecia, Parnass, Collect. Hauser (2♂♂, NHMW); Dedeagač, Graecia sept. (1♂, NMPC). SYRIA: Al Lathqiyan env., Slenfe env., 1500m, 30–31.V.1998 Josef Mertlik leg. (1♂, DWWC); Syria (1♀, NMPC). TURKEY: Anatolia m., Toros Daglari 900m, 25 km NW Erdemli, 6.VI.1991, S. Kadlec leg. (1♂, NHMW); Bogaz Roy, 12.VII.1989, H. Schmid leg. (1♂, NHMW); Asia Minor, Akschehir (1♂, NHMW); vill. Icel 24–26.V.1995, Erdemli env., 8 km NW of Arslanli, Josef Mertlik leg. . (1♂, DWWC); Asia min., 1.V.1969, Burdur, Antalya, Wewalka leg. (1♂, NMPC); prov. Adana, 20 km NW of Erdemli, 13–15.VI.1992, V. Bíša & Z. Koščál leg. (1♂, DWWC).
This species is most similar to
Length 5.1–5.9 mm. Head and pronotum brown; elytra piceous to black, with paler lateral margins and a poorly defined triangular area in basal ¼, sometimes extended along suture (
Head very deeply and densely punctate, strongly microsculptured; punctures sometimes confluent near eyes, separated by a diameter on the vertex. Clypeus impunctate. Eyes large and bulging, with a few very short setae at posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples short and smooth. Scape with a very long seta at 2/3 of its length and a few short thin setae; pedicel with a band of apical setae; antennomere III with two bands of setae (at midlength and at apex). Antennae pubescent from midlength of antennomere IV.
Pronotum 1.23–1.27 times as wide as head, 1.45–1.5 times as wide as long, widest just behind marginal setae. Anterior margin straight, anterior angles slightly prominent, sides very broadly and evenly rounded, slightly to conspicuously sinuate towards posterior angles, which are acute. Punctation coarse, irregular and deep, somewhat sparser towards furrow, rugose and confluent towards lateral margins. Lateral explanate margin rapidly widened from the apex, broad and flat at base. Posterior pore right in front of angle. Basal grooves shallow, rugose, confluently punctate. Pronotal base extended in a rounded median lobe. Furrow variable. Microsculpture strong, slightly transverse.
Intervals 1, 3, 5 and 7 with subtle, irregular setiferous punctures bearing minute, barely visible setae. Interval 7 narrow and strongly convex. All intervals convex near base and conspicuously convex elsewhere. Striae deep from base to apex, slightly punctate. Microsculpture strong, polygonal. Apices slightly sinuate.
Legs brownish yellow. Tarsomere V with 3 pairs of ventral setae. Mes and metepisterna slightly punctate. Claws with 5 teeth. Venter brown, abdominal sterna blackish. All abdominal sterna with short pubescence, no longer than metatarsomere IV.
Aedeagus –
. Habitus of
Varies in body size, in the extent of pale area at elytral base (may be reduced), and depth and length of the pronotal furrow.
Type locality: Athens, Greece (
Bulgaria, Greece, Turkey, Syria.
ISRAEL: Palestina, Gebatha, 2.IX.1927, collectto Paganetti (1 ex. NHMW); Israel, Golan Mach, Pappelrinde 16.7.1985 (1♂, NHMW); Galil, 8.6.1999, Yoqne'am, I. Trojan (1♂, DWWC). SYRIA: Syria occ., distr. Tartus, Safita env., 400 m a.s.l., 30 km SE of Tartus, 1.V.2000, steppe, S. Benedikt leg. (1♂, DWWC).
This species is most similar to
Length 6.8–7.0 mm. Head, pronotum, and legs brown, elytra piceous to black, sometimes slightly paler around scutellum. Venter brown, with darker abdomen (
Head deeply microsculptured, very deeply and densely punctate, less or no more than one diameter distance from each other. Clypeus punctate in posterior half. Eyes big and prominent, with no short setae at posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples short and smooth. Scape with a very long seta at 2/3 of its length and a few short thin setae towards the apex; pedicel with numerous setae in apical half; antennomere III pubescent towards apex. Antennae pubescent almost from the base of antennomere IV.
Pronotum 1.30–1.37 times as wide as head, 1.57 times as wide as long, widest just behind marginal setae. Anterior margin emarginate, anterior angles slightly prominent, sides very broadly rounded, not sinuate at posterior angles, which are rectangular and protrude as minute denticle. Punctation very coarse, as dense or denser than on head. Lateral explanate margin rapidly widened from the apex, broad and flat at base. Posterior pore right in front of angle. Basal grooves shallow, rugose, confluently punctate. Pronotal base extended in a rounded median lobe. Furrow usually shallow, fine, and incomplete. Microsculpture same as on head.
Elytra piceous to black, sometimes with base narrowly paler. Intervals 1 and 7 with a single row, 3 and 5 with two rows of irregular setiferous pores. Setae minute, barely visible. Interval 7 convex and narrow in basal half. Microsculpture polygonal to almost isodiametric. Apices slightly sinuate. Striae deep, punctate. All intervals convex at base, slightly convex at apex; outer intervals convex throughout.
Legs brownish yellow. Tarsomere V with 4–5 pairs of ventral setae. Claws with 4 long teeth. Epimera slightly rugose. Mes and metepisterna smooth. Venter brownish yellow, with 3–4 apical sterna darker. Abdomen with short pubescence.
Aedeagus –
Originally described as a subspecies of
Israel, Lebanon, Syria.
Type: ♂, Syrie (MNHN). JORDAN: Petra, Taybeh, 21.V.1994, W.G. Ullrich (1♀, DWWC). LEBANON: Appl, Beirut, 1878 (2♂, NHMW); Libanon, O v. Saida, 9–16.V.1963 Kasy & Vartian; “Phloeozetaeus apicalis Jedl. det. Ing. Jedlička" (1♂, NHMW).
Among the species consisting of beetles with strongly transverse and very densely punctate pronotum, easily diagnosable by bicoloured elytra, with apical half black.
Length 5.5–6.2 mm. Head, pronotum and ventral segments ferrugineous, apical half of elytra piceous (
Head very coarsely and deeply punctate on the sides and towards base, more sparsely on the frons, and very distinctly microsculptured. Punctures near eyes often confluent. Clypeus smooth, with distinctly microsculpture. Eyes large and bulging, with no short setae at the posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples very short, smooth. Scape with several setae besides the very long subapical one; pedicel with a band of apical setae; antennomere III in apical half with several setae besides the usual apical ones. Antennae pubescent from the basal fourth of antennomere IV.
Pronotum transverse, 1.17 times as wide as head, 1.4–1.48 times as wide as long, widest just behind marginal setae. Anterior margin straight, anterior angles effaced or faintly marked, sides broadly and evenly rounded, sinuate towards posterior angles, which are acute. Punctation coarse and irregular, punctures (especially on disc) generally smaller and sparser than on head, sometimes confluent (especially on the sides of disc). Lateral explanate margin uniform near apex, rapidly widened, wide and flat towards base. Posterior pore right in front of angle. Basal grooves shallow. Pronotal base extended in a rounded median lobe. Furrow variable. Microsculpture less distinct than on head.
Elytra shinier than head and pronotum. Each interval with 1–2 rows of minute, inconspicuous setae. Microsculpture more delicate than on head and pronotum. Apices weakly obliquely sinuate, rounded at suture. Striae slightly punctate, deep at base and at shoulders, shallower on disc. Interval 7 not much more convex and narrow than interval 6. Intervals slightly convex at base, flat at apex.
Legs brownish yellow. Protarsomere V with 3 pairs of ventral setae, metatarsomere V with 4 pairs. Claws with 4 teeth. All abdominal sterna pubescent, pubescence as long as on tarsi.
Aedeagus –
Varies in body size, density and size of pronotal punctures, and length of furrow.
Type locality per original description is Beirut (
Jordan, Lebanon. (Iraq, Israel, and Saudi Arabia records need confirmation.)
Type: ♂, Sinai, ouadi Sa'al, 25.II.1902; Sinai, O. Saal (1♀, MNHN); ISRAEL: Negev 12 km NW Elat, Oberh. En Natafim, 17.II.1987, 600m, Schawaller & Schmalfuss leg. (1♀, SMNS); Negev, MesharEbene, 14.II.1987, 300 m, Schawaller & Schmalfuss leg. (1♀, SMNS); Negev, 12 km NW Elat, Oberh. En Natafim, 17.II.1987, 600m, Schawaller & Schmalfuss leg. (1♀, DWWC).
This species is most similar to the allopatric
Length 6.8–7.0 mm. Reddish brown, with apical third of elytra black (
Head shiny, densely minutely punctate, punctures sometimes almost confluent in frontal depressions, separated by 1–2 diameters on the frons. Microsculpture scarcely visible. Clypeus with a few punctures near lateral margins. Eyes large and bulging, with 3–4 setae at posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples smooth, 2.6 times as short as length of eye. Scape with a very long seta at 2/3 of its length and a few short thin setae towards the apex; pedicel with a band of apical setae; antennomere III with two bands of setae (at midlength and at apex). Antennae pubescent from midlength of antennomere IV.
Pronotum shiny, without microsculpture, 1.12–1.14 times as wide as head, 1.33–1.4 times as wide as long, widest just behind the marginal setae. Anterior margin straight, anterior angles effaced, sides evenly rounded, considerably to slightly sinuate towards posterior angles, which are acute and protrude as minute denticle. Punctation, especially on disc, sparse, irregular, somewhat sparser than on head, denser towards lateral margins, where may be confluent and rugose. Lateral explanate margin widened basad lateral setae, broad and flat at base. Posterior pore right in front of angle. Basal grooves small and shallow. Pronotal base extended in a rounded median lobe. Furrow fine.
Intervals 1, 3, 5 and 7 each from base to apex with an irregular row of sparse pores bearing extremely short, barely visible setae. At humerus, interval 7 flat or weakly convex and as wide as the adjacent intervals. Microsculpture delicate, polygonal. Apices slightly sinuate. Striae finely punctate. Intervals slightly convex at base and almost flat at apex.
Legs brownish yellow. Tarsomere V with 4 pairs of ventral setae. Propleuron, mes and metepisterna smooth, shiny. Claws with 3 short teeth and one minute denticle at base. Venter uniformly testaceous. Abdominal sterna smooth, shiny, with sparse long pubescence.
Aedeagus –
Habitus of
Egypt, Israel. Iraq and Saudi Arabia records need confirmation.
Holotype: ♂, Afganistan, Kaboul //
Resembles
Length 6.8 mm. Yellowish redbrown, apical third of elytra pitchbrown (
Head densely finely punctate, slightly isodiametrically microsculptured; punctures confluent and rugose near eyes, separated by 1 to 3 diameters on the frons. Clypeus smooth. Eyes large and bulging, with 3–4 short setae at the posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples smooth, 2.6 times as short as eye length. Scape with a very long seta at 2/3 of its length and a few short thin setae towards the apex; pedicel with the usual band of apical setae; antennomere III with two bands of setae, at the middle and apex. Antennae pubescent from midlength of antennomere IV.
Pronotum 1.13 times as wide as head, 1.28 times as wide as long, widest just behind the marginal setae. Anterior margin slightly concave, sides behind marginal setae straight, sinuate towards posterior angles, which are rectangular and protrude as minute denticle. Punctation sparse, irregular, punctures (especially on disc) sparser than on head, transversely rugose and confluent along lateral margins. Lateral explanate margin widened from the marginal setae, broad and flat at base. Posterior pore right in front of angle. Basal grooves small and shallow. Pronotal base extended in a rounded median lobe. Furrow thin. Microsculpture faint, isodiametric.
Elytra elongate, in middle 1.46 times as long as wide. Intervals 1, 3, 5 and 7 with conspicuous pores bearing minute, barely visible setae. Microsculpture polygonal. Apices slightly sinuate, rounded at suture. Striae deep and minutely punctate. Intervals 3–7 convex at humerus, gradually flattened towards apices; other intervals slightly convex at base and almost flat at apex.
Legs pale brownish yellow. Tarsomere V with 4 pairs of ventral setae. Claws with three long and one small basal teeth. Venter entirely brownish yellow. Propleuron faintly rugose, episterna smooth. All abdominal sterna with short pubescence.
Aedeagus –
Named after the eminent Czech coleopterist Arnost Jedlička.
Known from a single male from Kaboul, Afghanistan.
OMAN: Oman bor., Prov. Batinah, AlJabal alAhdar mts., SE Rustaq, W Awabi, 430 m, Wadi Bani Awi,
Extremely similar to
Length 7.0–8.6 mm. Pale yellowish redbrown, elytra with large piceous preapical spot (
Head deeply irregularly punctate, microsculptured, punctures sometimes almost confluent in frontal depressions, separated by over three diameters on the frons. Clypeus with few punctures near lateral margins. Eyes large and bulging, with a few short setae at posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples short and smooth. Scape with a very long seta at 2/3 of its length and a few short thin setae towards apex; pedicel with the usual band of apical setae; antennomere III with two bands of setae at midlength and at apex. Antennae pubescent from midlength of antennomere IV.
Pronotum shinier than head and elytra, 1.16–1.18 times as wide as head, 1.39–1.42 times as wide as long, widest just behind marginal setae. Anterior margin straight, anterior angles effaced, sides very broadly and evenly rounded, slightly sinuate towards posterior angles, which are acute and protrude as minute denticle. Punctation coarse, irregular, deep, somewhat sparser than on head, especially on disc; rugose and confluent at apical and basal margins. Lateral explanate margin rapidly widened from the apex, broad and flat at base. Posterior pore right in front of angle. Basal grooves shallow, rugose, confluently punctate. Pronotal base extended in a rounded median lobe. Furrow deep and complete. Microsculpture faint.
Elytra pale yellowish redbrown, with red brown apices and piceous preapical spot reaching lateral margins (may be reduced to five inner intervals). Intervals 1, 3, 5 and 7 each with minute setae and a row of pores from base to apex. Interval 7 flat, as wide as adjacent intervals. Microsculpture deep, irregular, polygonal, same as on head. Apices slightly sinuate. Striae slightly punctate, shallower on disc and at apex. Intervals slightly convex near base, flat at apex.
Legs brownish yellow. Tarsomere V with 3 pairs of ventral setae. Mes and metepisterna slightly punctate. Claws with 5 teeth. Venter entirely light brownish yellow. All abdominal sterna with pubescence more than twice as long as protarsomere IV.
Aedeagus (
Varies in body size; elytral spot sometimes reduced. Pronotal basal angles usually acute and prominent but sometimes rectangular and less prominent.
Named after Ron Felix, my friend and collaborator who discovered this species.
Oman (country record), UAE.
HOLOTYPE: ♀, S.O. Iran, Djiroft, AnbarAbad, 1–18.V.1956, W. Richter leg. (SMNS).
This species is most similar to
Length 7.0 mm. Reddish yellow, with apical third of elytra dark (
Head microsculptured and rather coarsely punctate. Clypeus punctate in basal half. Eyes with a few small setae at the posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples short and smooth. Scape with a very long seta at 2/3 of its length and a few short thin setae towards the apex; pedicel with a band of apical setae; antennomere III with a few setae on apical half. Antennae pubescent from the basal fourth of antennomere IV.
Pronotum brownish yellow, shinier than head and elytra, 1.2 times as wide as head, 1.46 times as wide as long, widest behind the marginal setae. Anterior margin straight, anterior angles effaced, lateral margin very broadly and evenly rounded from anterior angle to marginal setae, almost straight behind that point, slightly sinuate towards posterior angles, which are acute and protrude as minute denticle. Punctation sparse, especially on disc, rugose and confluent at apical and basal margins. Lateral explanate margin widened behind marginal setae, broad and flat at base. Posterior pore right in front of angle. Basal grooves shallow, rugose, confluently punctate. Pronotal base extended in a rounded median lobe. Furrow deep and complete. Microsculpture faint.
Elytra: striae fine, finely punctate. Intervals 1, 3, 5 and 7 with extremely tiny and short setae and row of small pores from base to apex, very difficult to see. Interval 7 at shoulder slightly convex and narrower than the adjacent ones. Microsculpture deep, irregular, polygonal. Intervals slightly convex at base and almost flat at apex.
Legs brownish yellow. Tarsomere V with 3 pairs of ventral setae. Episterna of meso and metathorax slightly punctate. Claws with 3–4 obtuse teeth. Venter entirely light brownish yellow. All abdominal sterna with pubescence short, no longer than protarsomere IV.
Male unknown.
Iran.
Holotype: ♂, Iran, Khorasan prov., Gonobad area, 10 km SW Khezri, 1800m, 22.V.2009 Kolesnichenko K. leg.; Paratype: ♂, Kopetdag, 12 km SW KizylArvat, light trap, 5.VII.1953 Kryzhanovskii leg. (ZIN).
This new species shares with
Length 6.3–6.5 mm. Uniformly yellowish redbrown (
Head very coarsely and deeply irregularly punctate, with distinct isodiametric microsculpture, punctures confluent near eyes, separated by over three diameters on the frons. Clypeus smooth, with a few small punctures near base. Eyes very large and bulging. Second supraocular seta just before the level of the posterior margin of the eye. Temples very short, smooth. Scape smooth, with one very long and 3–4 short subapical setae; pedicel with the usual band of apical setae; antennomere III with two bands of apical setae. Antennae pubescent from the basal third of antennomere IV.
Pronotum 1.16 times as wide as head, 1.36–1.42 times as wide as long, widest just behind marginal setae. Anterior margin straight, anterior angles effaced, sides broadly and evenly rounded, slightly sinuate towards posterior angles, which are rectangular and protrude as minute denticle. Punctation irregular, punctures (especially on disc) shallower and sparser than on head, confluent at basal grooves. Lateral explanate margin even from apex to marginal setae, then rapidly widened and flat. Posterior pore right in front of angle. Basal grooves shallow. Pronotal base extended in a rounded median lobe. Furrow deep. Microsculpture as on head.
Elytra subparallel, 1.4 times as long as wide. Intervals 1, 3, 5 and 7 with small pores bearing minute inconspicuous setae. Microsculpture as on pronotum. Apical edge straight, rounded at suture. Striae deep, slightly punctate. Intervals 4–8 convex in basal half, gradually flattened towards apex; other intervals slightly convex at base and flat at apex.
Legs pale brownish yellow. Tarsomere V with 5 pairs of ventral setae. Claws with 5 teeth. Episterna all smooth. Abdominal sterna with long pubescence.
Aedeagus –
In the holotype, elytral intervals are more convex and striae more crenulate than in the paratype, whose striae are almost smooth. However the aedeagal configuration is identical, and so the two specimens are considered conspecific.
Named after the eminent Russian coleopterist Oleg Kryzhanovsky.
Iran, Turkmenistan.
Holotype: ♂, Iran, Fars, 20 km W Estahban, 2400m, 27–30.V.2008 Anichtchenko A. leg. (ZIN). Paratypes: same locality and date (2♂♂ 4♀♀, ZIN, AAC).
This new species shares with
Length 5.7–6.7 mm. Uniformly yellowish redbrown (
Head very coarsely and deeply punctate on sides and towards the base, more sparsely on the frons; punctures near eyes often confluent. Head with very distinct microsculpture. Clypeus smooth, with distinct microsculpture. Eyes large and bulging, with no short setae at the posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples very short, smooth. Scape with several setae besides the very long subapical one; pedicel with the usual band of apical setae; antennomere III in apical half with several setae besides the usual apical ones. Antennae pubescent from the basal fourth of antennomere IV.
Pronotum 1.17 times as wide as head, 1.4–1.48 times as wide as long, widest just behind marginal setae. Anterior margin straight, anterior angles effaced to faintly marked, sides broadly and evenly rounded, sinuate towards posterior angles, which are acute. Punctation coarse, irregular, punctures (especially on disc) generally smaller and sparser than on head, sometimes confluent (especially on the sides of disc). Lateral explanate margin uniform near apex, rapidly widened, wide and flat towards base. Posterior pore right in front of angle. Basal grooves shallow. Pronotal base extended in a rounded median lobe. Furrow variable. Microsculpture less distinct than on head.
Elytra shinier than head and pronotum. Each interval with 1–2 rows of minute, inconspicuous setae. Microsculpture more delicate than on head and pronotum. Apices weakly obliquely sinuate, rounded at suture. Striae slightly punctate, deep at base and at shoulders, shallower on disc. Interval 7 not much more convex and narrow than interval 6. Intervals slightly convex at base, flat at apex.
Legs brownish yellow. Protarsomere V with 3 pairs of ventral setae, metatarsomere V with 4 pairs. Claws with 4 teeth. Abdominal pubescence as long as tarsal, with no long setae.
Aedeagus –
Varies in body size, density and size of pronotal punctures, and length of furrow.
Named after my friend Saeed Mobarra.
Iran.
Syrie, Monts Amanus, Delagrande (in coll. Pic, MNHN). Not examined.
I failed to locate type specimens in MNHN, including the Pic collection. I have examined one female determined as
Holotype: ♂, Persien, Löffler (=Iran, Kuhrang) (NMPC); Iran, Kerman, Sirjan, 8 km N Balvard, 11–12.V.2007, Anichtchenko A. leg. (1♂, CAA).
In
Length 5.4 mm. redbrown with postmedian black band on elytra (
Head smooth, microsculptured, deeply irregularly punctate, punctures sometimes almost confluent near eyes, separated by 3 to 6 diameters on the front and by their diameter towards head base. Clypeus impunctate. Eyes large and bulging, with a few short setae at posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples short and smooth. Scape with a very long seta at 2/3 of its length and a few thin setae towards the apex; pedicel irregularly setose throughout; antennomere III with numerous setae in apical 2/3. Antennae pubescent from the basal fourth of antennomere IV.
Pronotum redbrown, smooth, shinier than head, 1.21 times as wide as head, 1.34 times as wide as long, widest right behind marginal setae. Anterior margin straight, anterior angles slightly prominent, sides very broadly and regularly rounded, slightly to moderately sinuate towards posterior angles, which are rectangular to acute. Punctation irregular, sparser and shallower than on the head, sparse on the disc and denser towards base. Lateral explanate margin rapidly widened from apex, broad and elevated at base. Posterior pore right in front of angle. Basal grooves punctate. Pronotal base extended in a rounded median lobe. Furrow short and shallow. Microsculpture weak, slightly transversely polygonal.
Elytra redbrown, with piceous to black postmedian band reaching lateral margins. Intervals 1, 3, 5, 7 and 8 setose, with a single irregular row of pores all along. All intervals setose at base, convex on basal third and slightly convex beyond that. Interval 7 convex from base to the middle. Microsculpture deep, irregular, polygonal. Apical margin slightly sinuate. Striae deep, punctate, shallower towards the apices.
Legs brownish yellow. Tarsomere V with 3 pairs of ventral setae. Mes and metepisterna smooth. Claws with 4 teeth. Venter entirely light brownish yellow. Abdominal sterna pubescent throughout, pubescence as long as protarsomere II.
Aedeagus –
Elytral band may be reduced to a spot not reaching beyond interval 4. Pronotal and elytral punctures vary in size.
Jedlička (1963b: 177) erroneously referred to the holotype as female.
Iran.
Holotype: ♂, small green rectangle; white label in Russian: “coll. Khristofa" [‘Christoph Collection']; white label in Russian: “na osnovanii tetradi etiketka: Schahrud, Persia 1870–1873" [‘Location per log: Schahrud, Persia 1870–1873'] (ZIN); Paratype: ♀, Iran, Lorestan, 1–2.V.2007, 20 km N PoleDokhtar, Baba Zeyd, near Muruni, Anichtchenko A. leg. (AAC).
This new species can be confused with specimens of sympatric
Length 6.1–6.2 mm. Yellowish redbrown, with legs slightly paler and a dark postmedian elytral spot reaching interval 4 (
Head coarsely and deeply irregularly punctate, dull, very distinctly microsculptured. Punctures in the frons sometimes separated by over twice the diameter. Clypeus with some small irregular punctures. Eyes very large and bulging, with no small setae at posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples short, smooth. Scape with several rather long setae besides the very long subapical one; pedicel with the usual band of apical setae; antennomere III with two bands of apical setae. Antennae pubescent from the basal third of antennomere IV.
Pronotum brownish yellow, 1.24 times as wide as head, 1.4 times as wide as long, widest just behind marginal setae. Anterior margin straight, anterior angles slightly marked, sides broadly and evenly rounded, more or less sinuate towards posterior angles, which are rectangular and protrude as minute denticle. Punctation coarse, irregular and deep, as dense as on head, somewhat sparser on disc and sometimes confluent, especially at the sides of disc. Lateral explanate margin rather wide at apex, rapidly widened basally, broad and flat towards base. Posterior pore right in front of angle. Basal grooves shallow. Pronotal base extended in a rounded median lobe. Furrow variable. Microsculpture less distinct than on head.
Elytra shinier than head and pronotum, 1.4 times as long as wide. Intervals 1, 3, 5 and 7 each with a row of setiferous pores. Pores of intervals 5 and 7 deep, with rather long setae. Intervals 2 and 4 with several minute setae at base. Microsculpture rather isodiametric but irregular, more delicate than on head and pronotum. Apices weakly obliquely sinuate, rounded at suture. Striae deep and crenulate. Interval 7 narrow and convex in basal half, then gradually flattened; other intervals convex at base, almost flat at apex.
Legs pale brownish yellow. Tarsomere V with 4 pairs of ventral setae. Claws with 4 teeth. Venter uniformly testaceous. All abdominal sterna with long pubescence. All episterna smooth.
Aedeagus –
Posterior pronotal angles almost rectangular in the holotype, more prominent and acute in the paratype.
The name (Latin, adjective: timid) refers to the cryptic lifestyle.
Iran.
1  Body elongate ( 
2 
–  Body wide ( 
4 
2  Bicolored  3 
–  Body completely black. (“Siberia", unspecified)  
3  Pronotal sides sinuate in front of base, hind angles acute ( 

–  Pronotal sides less sinuate in front of base, hind angles rectangular. Femora and apical part of tibiae piceous to black ( 

4  Pronotum strongly transverse, very densely punctate, distance between punctures less than their diameter ( 
5 
–  Pronotum less transverse or subquadrate, distance between punctures more than their diameter  7 
5  Elytra uniformly brownish yellow or piceous  6 
–  Elytra bicoloured, with apical half black ( 

6  Body colour uniformly yellow pale. Intervals of elytra flat. Small size species 4.8–5.4 mm. ( 

–  Body piceous to black, pronotum and sometimes elytral base narrowly paler ( 

7  All elytral intervals with conspicuous setiferous pores. Pubescence long  8 
–  Setae short and thin, sometimes very short and inconspicuous  14 
8  Intervals 1–7 with a single uninterrupted row of setiferous pores, 8^{th} with two rows  9 
–  All intervals of elytra with 2–3 irregular and dense rows of big setiferous pores  12 
9  Elytra elongate, flat, with black postmedian transverse band and redbrown apices. 6.0–6.5 mm. ( 

–  Elytra subovate, convex. Smaller, 4.6–5.1 mm.  10 
10  Uniformly yellow brownish  11 
–  Elytra with weak, diffuse, grey postmedian band, sometimes reduced to sutural spot, intervals 2, 4 and 6 without setiferous pores ( 

11  All elytral intervals with a single uninterrupted row of widely spaced setiferous pores. Pronotum transverse. Head and disc of pronotum very sparsely and superficially punctate ( 

–  Rows of setiferous punctures on intervals 3–5 widely interrupted. Setiferous pores on all intervals large. Pronotum cordate. Head smooth, pronotum almost smooth. ( 

12  Propleura smooth and shiny  13 
–  Propleura with wavy rugae. Size bigger, 6.8–8 mm. Elytra longer, basal red spot almost reaching the middle of elytra ( 

13  Propleura smooth and shiny. Body 6 mm. Elytra shorter and wider, basal third redbrown ( 

–  Small, 5.1 mm. Basal half of elytra red. Elytra shiny, lightly microsculptured. Setiferous pores on intervals large, almost never situated on a par in the same interval. Head and pronotum sparsely punctate ( 

14  Body and elytra uniformly yellowbrown  15 
–  Body bicolored  16 
15  All elytral intervals sparsely and irregularly punctate along striae ( 

–  Only odd elytral intervals with irregular and weak setiferous pores all along striae. Setae very thin and short, barely visible ( 

16  Head and pronotum redbrown. Elytra redbrown with black band or spot, or with black posterior third of elytra  17 
–  Head and elytra piceous to black. Pronotum and narrow diffuse band on the base of elytra paler ( 

17  Smaller, up to 5 mm.  18 
–  Larger, at least 6–7 mm.  21 
18  Pronotum strongly transverse. Elytra with black sutural spot extended to interval 4 or 5 ( 

–  Pronotum subquadrate or slightly transverse  19 
19  Pronotum subquadrate, sparsely irregularly punctate, punctures 2–6 diameter distance from each other, lateral sides sinuate before acutely prominent posterior angles. Intervals of elytra convex on the base, setiferous pores on odd intervals conspicuous. Transverse band variable ( 

–  Pronotum slightly transverse, densely punctate, punctures 1–2 diameters distance from each other, same as in head  20 
20  Elytra short. Transverse black band of elytra narrow, often reduced to inner intervals or disappeared ( 

–  Elytra more elongate. Transverse black band of elytra broad, almost reaching apices of elytra. Odd intervals in apical part with 2 rows of setae, even intervals with 1 row of sparse setae ( 

21  Pronotum densely punctate, same as on the head. Black apical spot of elytra not reaching apices  22 
–  Pronotum sparsely punctate, shiny. Black apical spot of elytra reaching apices  23 
22  Black elytral spot narrow, occupying zone from suture to 3 or 4 interval, not reaching apices of elytra ( 

–  Transverse black band of elytra broad, almost reaching apices of elytra ( 

23  Pronotum conspicuously microsculptured  24 
–  Pronotum shiny, without microsculpture ( 

24  Odd intervals of elytra with easily visible short setae along striae  25 
–  Odd intervals of elytra with extremely tiny and short setae along striae. Hairs on abdominal sterna short, length no more than 1/3 of length of the apical setae. Pronotum more strongly transverse, sparsely and irregular punctate, less dense than on the head, disc almost without punctures, slightly rugose ( 

25  Hairs on abdominal sterna long, the same or at least 2/3 of length of the apical setae. Pronotum weakly transverse, regularly punctate. ( 

–  Hairs on abdominal sterna short, length no more than 1/3 of length of the apical setae. Apical third of elytra black ( 
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Drs. Boris Kataev, Nikolai Nikitsky, Thierry Deuve and Azadeh Taghavian facilitated my work with the collections of ZIN, ZMM, and MNHN; Drs. Andreas Pütz, David W. Wrase, Ron F.F.L. Felix, Erich Kirschenhofer, and Wolfgang Schawaller provided reference specimens; Vassili Belov prepared the English version; the anonymous reviewers and editor improved my manuscript. The author is grateful for their help. Special thanks to Dr. Kirill Makarov for providing advice and excellent images.
This paper has been prepared under the European Social Fund project No.2009/0206/1DP/1.1.1.2.0/09/APIA/VIAA/010.