Corresponding author: Dirk Ahrens (
Academic editor: A. Frolov
The species of the
Liu W-G, Fabrizi S, Bai M, Yang X, Ahrens D (2016) A taxonomic revision of
In the course of the revision of the species-rich genus
As shown earlier (Ahrens 2003,
The terminology and methods used for measurements, specimen dissection and genital preparation follow
Abbreviations used in the text for the collection depositories are as follows:
Bernice P.
collection M. Nikodým, Prague, Czech Republic
collection P. Pacholátko, Brno, Czech Republic
collection G. Sabatinelli, Prevessin, France
Museum of Hebei University, Baoding, Hebei Province, China
Muséum d’Histoire Naturelle, Genève, Switzerland
Museum national d’Histoire naturelle, Paris, France
Naturhistorisches Museum Wien, Austria
Northwest A & F University, Yangling, Shaanxi Province, China
Senckenbergmuseum, Frankfurt Main, Germany
Shanghai Normal University, Department of Biology, China
Sun Yat-Sen University, Guangzhou, China
Zoologisches Forschungsmuseum A. Koenig, Bonn, Germany
1 | Hypomeron not carinate |
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– | Hypomeron carinate |
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2 | Antennal club in female composed of 3 antennomeres |
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– | Antennal club in female composed of more than 3 antennomeres |
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3 | Posterior margin of metafemur serrate ventrally and dorsally |
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– | Posterior margin of metafemur smooth ventrally |
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4 | Anterior angles of pronotum obsolete |
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– | Anterior angles of pronotum acute and moderately produced |
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5 | Dorsal surface nearly glabrous |
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– | Dorsal surface densely setose |
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6 | Metatibia beside dorsal margin with a serrated longitudinal line or carina |
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– | Metatibia beside dorsal margin without a serrated longitudinal line or carina |
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7 | Metatibia beside dorsal margin with a serrated longitudinal line or carina |
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– | Metatibia beside dorsal margin without a serrated longitudinal line or carina |
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8 | Metatibia with one group of robust spines |
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– | Metatibia with two groups of robust spines |
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9 | Antennal club in males long and reflexed |
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– | Antennal club in males short or moderately long and straight |
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10 | Protibia bidentate |
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– | Protibia tridentate |
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11 | Elytra bicolored, yellowish or reddish brown and black |
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– | Elytra unicolored |
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12 | Parameres symmetrical |
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– | Parameres asymmetrical |
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13 | Apex of metatibia shallowly truncate at interior apex near tarsal articulation |
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– | Apex of metatibia sharply truncate at interior apex near tarsal articulation |
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14 | Dorsal surface yellowish brown to reddish brown, strongly and simply shiny |
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– | Dorsal surface dull or iridescent shiny |
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15 | Pronotum and elytra always nearly glabrous | |
– | Pronotum and elytra always distinctly setose |
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16 | Labrum without a transverse rim of very dense, short and robust setae |
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– | Labrum short, with a transverse rim of very dense, short and robust setae. Dorsal surface densely setose |
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17 | Metatibia slender and long |
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– | Metatibia short and wide |
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18 | Body smaller 8.5 mm |
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– | Body larger 9 mm | |
19 | Antennal club of males with 7 antennomeres |
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– | Antennal club of males with 6 or less antennomeres |
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20 | Metafemur with a continuously serrated line adjacent to the anterior margin of metafemur. Protibia more or less distinctly tridentate |
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– | Metafemur without a continuously serrated line adjacent to the anterior margin of metafemur. Protibia always distinctly bidentate |
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21 | Basis of labroclypeus dull. Antennal club of males with 6 antennomeres |
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– | Antennal club of males with 5 or 4 antennomeres |
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22 | Angle between basis of hypomeron and that of pronotum strongly rounded, angle between surfaces of hypomeron and pronotum basally blunt. Hypomeron basally strongly produced ventrally and transversely sulcate |
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– | Angle between basis of hypomeron and that of pronotum sharp, angle between surfaces of hypomeron and pronotum sharp. Hypomeron basally not produced ventrally and not sulcate |
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23 | Apex of metatibia shallowly truncate at interior apex near tarsal articulation |
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– | Apex of metatibia deeply truncate at interior apex near tarsal articulation | |
24 | Body surface strongly shiny. Body smaller (5.7–6.6 mm) |
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– | Body surface dull. Body larger (8 mm) |
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1 | Labrum without densely setose carina |
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– | Labrum with densely setose carina |
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2 | Distal hook of left paramere nearly half as long as paramere itself |
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– | Distal hook of left paramere shorter than one quarter of length of paramere itself |
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3 | Distal hook of left paramere strongly curved and at apex bent backwards |
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– | Distal hook of left paramere moderately curved and at apex bent externally only |
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Holotype: ♂ “China (Yunnan) Nujiang Lisu Aut. Pref., Salween side valley, 5 km S Fugong, road SS228, km 223 (creek bank, litter sifted) 8.VI.2007 leg. D. Wrase/ X-DA1554” (
The new species has the genitalia similar in shape to
Body length: 6.7 mm, length of elytra: 4.5 mm, body width: 3.7 mm. Body oval, yellowish brown, dorsal surface strongly shiny and glabrous.
Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins nearly straight, convergent anteriorly, anterior angles strongly rounded, anterior margin very shallowly sinuate medially, margins moderately reflexed; surface convexly elevated at centre, shiny, finely and sparsely punctate, with a few single setae anteriorly; frontoclypeal suture distinctly incised, slightly elevated and weakly curved; smooth area anterior to eye weakly convex, approximately 1.5 times as wide as long; ocular canthus short and narrow (1/3 of ocular diameter), impunctate, with one terminal seta. Frons with fine and sparse punctures, with a few long erect setae beside eyes and on posterior half of frons. Eyes large, ratio diameter/ interocular width: 0.69. Antenna with ten antennomeres, club with five antennomeres and straight, slightly longer than remaining antennomeres combined. Mentum elevated and convex anteriorly. Labrum short, lens-shaped in anterior view, not produced medially, with shallow median sinuation and without densely setose anterior margin.
Pronotum moderately transverse, widest shortly behind middle, lateral margins evenly convex and weakly convergent towards base, more strongly convergent anteriorly; anterior angles distinctly produced and sharp, posterior angles blunt and weakly rounded at tip; anterior margin straight, with a fine complete marginal line; surface moderately densely and finely punctate, glabrous; lateral and anterior border sparsely setose; hypomeron distinctly carinate basally, not produced ventrally. Scutellum narrow, triangular, with fine, dense punctures, impunctate on basal midline, glabrous.
Elytra oval, widest at posterior third, striae finely impressed, finely and densely punctate, intervals nearly flat, with sparse, fine punctures concentrated along striae, glabrous; epipleural edge fine, ending at widely rounded external apical angle of elytra, epipleura densely setose, apical border with a fine fringe of microtrichomes (visible at 100× magnification).
Ventral surface shiny, finely and densely punctate, metasternum glabrous; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures, each bearing a short robust seta. Mesosternum between mesocoxae as wide as the mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 1.55. Pygidium moderately convex and shiny, finely and sparsely punctate, without smooth midline, with a few long setae along apical margin.
Legs short; femora shiny, with two rudimentary longitudinal rows of setae, superficially and sparsely punctate, glabrous; metafemur with anterior margin acute, without serrated line behind anterior edge, posterior margin smooth ventrally in apical half only weakly widened, posterior margin smooth dorsally. Metatibia moderately wide and short, widest at middle, ratio of width/ length: 1/ 2.8; dorsal margin sharply carinate, with two groups of spines, basal group at one third, apical group at two thirds of metatibial length, basally with a few short single setae; lateral face weakly convex, finely and very sparsely punctate, smooth along the middle; ventral edge finely serrated, with three robust nearly equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, short setae, smooth, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally, smooth; first metatarsomere slightly shorter than following two tarsomeres combined and distinctly longer than dorsal tibial spur. Protibia moderately long, bidentate, distal tooth sharply pointed at apex; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex.
Aedeagus: Fig.
This new species is named with reference to its type locality, Fugong.
Holotype: ♂ “CHINE - Yunnan Tongbinguan
The new species is very similar to
Body length: 6.8 mm, length of elytra: 4.5 mm, body width: 3.7 mm. Body oval, yellowish brown, dorsal surface strongly shiny and glabrous.
Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins nearly straight, convergent anteriorly, anterior angles strongly rounded, anterior margin very shallowly sinuate medially, margins moderately reflexed; surface convexly elevated at centre, shiny, finely and sparsely punctate, with a few single setae anteriorly; frontoclypeal suture distinctly incised, slightly elevated and weakly curved; smooth area anterior to eye weakly convex, approximately 1.5 times as wide as long; ocular canthus short and narrow (1/3 of ocular diameter), impunctate, with one terminal seta. Frons with fine and sparse punctures, with a few long erect setae beside eyes and on posterior half of frons. Eyes large, ratio diameter/ interocular width: 0.69. Antenna with ten antennomeres, club with five antennomeres and straight, slightly longer than remaining antennomeres combined. Mentum elevated and convex anteriorly. Labrum short, lens-shaped in anterior view, not produced medially, with shallow median sinuation and without densely setose anterior margin.
Pronotum moderately transverse, widest shortly behind middle, lateral margins evenly convex and weakly convergent towards base, more strongly convergent anteriorly; anterior angles distinctly produced and sharp, posterior angles blunt and weakly rounded at tip; anterior margin weakly convex, with a fine complete marginal line; surface moderately densely and finely punctate, glabrous; lateral and anterior border sparsely setose; hypomeron distinctly carinate basally, not produced ventrally. Scutellum narrow, triangular, with fine, dense punctures, impunctate on basal midline, glabrous.
Elytra oval, widest at posterior third, striae finely impressed, finely and densely punctate, intervals nearly flat, with sparse, fine punctures concentrated along striae, glabrous; epipleural edge fine, ending at widely rounded external apical angle of elytra, epipleura densely setose, apical border with a fine fringe of microtrichomes (visible at 100× magnification).
Ventral surface shiny, finely and densely punctate, metasternum glabrous; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures, each bearing a short robust seta. Mesosternum between mesocoxae as wide as the mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 1.42. Pygidium moderately convex and shiny, finely and sparsely punctate, without smooth midline, with a few long setae along apical margin.
Legs short; femora shiny, with two rudimentary longitudinal rows of setae, superficially and sparsely punctate, glabrous; metafemur with anterior margin acute, without serrated line behind anterior edge, posterior margin smooth ventrally in apical half only weakly widened, posterior margin smooth dorsally. Metatibia moderately wide and short, widest at middle, ratio of width/ length: 1/ 2.8, dorsal margin sharply carinate, with two groups of spines, basal group at one third, apical group at three quarters of metatibial length, basally with a few short single setae; lateral face weakly convex, finely and sparsely punctate, smooth along the middle; ventral edge finely serrated, with three robust nearly equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, short setae, smooth, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally, smooth; first metatarsomere slightly shorter than following two tarsomeres combined and distinctly longer than dorsal tibial spur. Protibia moderately long, bidentate, distal tooth sharply pointed at apex; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex.
Aedeagus: Fig.
This new species is named after one of its collectors, Mr. Mantilleri, who provided us with a series of unidentified specimens from his expedition to China.
Holotype: ♂ “Dongjiafen, Jingdong, Yunnan, 16.VI.1956, leg. Zaguljaev” (
The new species is in shape of genitalia and in external appearance very similar to
Body length: 6.4 mm, length of elytra: 4.5 mm, body width: 3.5 mm. Body oval, yellowish brown, dorsal surface strongly shiny and glabrous.
Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins nearly straight, convergent anteriorly, anterior angles strongly rounded, anterior margin very shallowly sinuate medially, margins moderately reflexed; surface convexly elevated at centre, shiny, finely and sparsely punctate, with a few single setae anteriorly; frontoclypeal suture distinctly incised, slightly elevated and weakly curved; smooth area anterior to eye weakly convex, approximately 1.5 times as wide as long; ocular canthus short and narrow (1/3 of ocular diameter), impunctate, with one terminal seta. Frons with fine and sparse punctures, with a few long erect setae beside eyes and on posterior half of frons. Eyes moderately large, ratio diameter/ interocular width: 0.6. Antenna with ten antennomeres, club with five antennomeres and straight, as long as remaining antennomeres combined. Mentum elevated and convex anteriorly. Labrum short, lens-shaped in anterior view, not produced medially, with shallow median sinuation and without densely setose anterior margin.
Pronotum moderately transverse, widest at base, lateral margins evenly convex and weakly convergent anteriorly; anterior angles distinctly produced and sharp, posterior angles blunt and weakly rounded at tip; anterior margin weakly convex, with a fine complete marginal line; surface moderately densely and finely punctate, glabrous; lateral and anterior border sparsely setose; hypomeron distinctly carinate basally, not produced ventrally. Scutellum narrow, triangular, with fine, moderately dense punctures, impunctate on basal midline, glabrous.
Elytra oval, widest at posterior third, striae finely impressed, finely and densely punctate, intervals nearly flat, with sparse, fine punctures concentrated along striae, glabrous except a few long setae on penultimate lateral interval; epipleural edge fine, ending at widely rounded external apical angle of elytra, epipleura densely setose, apical border with a fine fringe of microtrichomes (visible at 100× magnification).
Ventral surface shiny, finely and densely punctate, metasternum glabrous; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures, each bearing a short robust seta. Mesosternum between mesocoxae as wide as the mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 1.64. Pygidium moderately convex and shiny, finely and sparsely punctate, without smooth midline, with a few long setae along apical margin.
Legs short; femora shiny, with two rudimentary longitudinal rows of setae, superficially and sparsely punctate, glabrous; metafemur with anterior margin acute, without serrated line behind anterior edge, posterior margin smooth ventrally in apical half only weakly widened, posterior margin smooth dorsally. Metatibia moderately wide and short, widest at middle, ratio of width/ length: 1/ 2.8, dorsal margin sharply carinate, with two groups of spines, basal group at one third, apical group at three quarters of metatibial length, basally with a few short single setae; lateral face weakly convex, finely and sparsely punctate, smooth along the middle; ventral edge finely serrated, with three robust nearly equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, short setae, smooth, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally, smooth; first metatarsomere distinctly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate, distal tooth sharply pointed at apex; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex.
Aedeagus: Fig.
The name of the new species is derived from the type locality, Dongjiafen.
Body length: 5.3–6.4 mm, length of elytra: 4.2–4.5 mm, body width: 3.4–3.5 mm.
Holotype: ♂ “[China] Menglun, Yunnan, 19.V.1991, leg. Wang Yinglun, Tian Binggang” (
Body length: 5.5 mm, length of elytra: 4.0 mm, body width: 3.7 mm. Body oval, yellowish brown, dorsal surface strongly shiny and glabrous.
Labroclypeus short and subtrapezoidal, distinctly wider than long, widest at base, lateral margins nearly straight, convergent anteriorly, anterior angles moderately rounded, anterior margin broadly sinuate medially, margins moderately reflexed; surface nearly flat, shiny, finely and sparsely punctate, with a few single setae anteriorly; frontoclypeal suture distinctly incised, slightly elevated and weakly curved; smooth area anterior to eye weakly convex, approximately 1.5 times as wide as long; ocular canthus short and narrow (1/3 of ocular diameter), impunctate, with one terminal seta. Frons with fine and moderately dense punctures, with a few long erect setae beside eyes and behind frontoclypeal suture. Eyes moderately large, ratio diameter/ interocular width: 0.64. Antenna with ten antennomeres, club with five antennomeres and straight, slightly shorter than remaining antennomeres combined. Mentum elevated and convex anteriorly. Labrum short, nearly lens-shaped in anterior view, not produced medially, with shallow median sinuation and with a rim of dense setae near anterior margin.
Pronotum moderately transverse, widest at base, lateral margins in basal half nearly straight and moderately convergent to middle, evenly convex and weakly convergent anteriorly; anterior angles distinctly produced and sharp, posterior angles blunt and weakly rounded at tip; anterior margin weakly convex, with a fine complete marginal line; surface moderately densely and finely punctate, glabrous; lateral and anterior border sparsely setose; hypomeron distinctly carinate basally, not produced ventrally. Scutellum narrow, triangular, with fine, moderately dense punctures, impunctate on basal midline, glabrous.
Elytra oval, widest at posterior third, striae finely impressed, finely and densely punctate, intervals nearly flat, with sparse, fine punctures concentrated along striae, glabrous except a few long setae on penultimate lateral interval; epipleural edge fine, ending at widely rounded external apical angle of elytra, epipleura densely setose, apical border without a fine fringe of microtrichomes (visible at 100× magnification).
Ventral surface shiny, finely and densely punctate, metasternum glabrous; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures, each bearing a short robust seta. Mesosternum between mesocoxae as wide as the mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 1.61. Pygidium moderately convex and moderately shiny, finely and densely punctate, without smooth midline, with a few long setae along apical margin.
Legs short; femora shiny, with two rudimentary longitudinal rows of setae, superficially and sparsely punctate, glabrous; metafemur with anterior margin acute, without serrated line behind anterior edge, posterior margin smooth ventrally in apical half only weakly widened, posterior margin smooth dorsally. Metatibia wide and short, widest at middle, ratio of width/ length: 1/ 2.4, dorsal margin sharply carinate, with two groups of spines, basal group at one third, apical group at three quarters of metatibial length, basally with a few short single setae; lateral face weakly convex, finely and sparsely punctate, smooth along the middle; ventral edge finely serrated, with three robust nearly equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, short setae, smooth, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally, smooth; first metatarsomere distinctly shorter than following two tarsomeres combined and only slightly longer than dorsal tibial spur. Protibia moderately long, bidentate, distal tooth sharply pointed at apex; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex.
Aedeagus: Fig.
Body length: 5.4–5.5 mm, length of elytra: 3.9–4.0 mm, body width: 3.3–3.7 mm.
The new species is named after the type locality, Menglun.
1 | Phallobase with a strong ventral lamina on the right side. |
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– | Phallobase without a ventral lamina. |
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2 | Phallobase at apex with a pair of distinct dorsal protuberances |
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– | Phallobase at apex without a pair of distinct dorsal protuberances |
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3 | Phallobase at apex with a pair of distinct dorsal protuberances |
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– | Phallobase at apex without a pair of distinct dorsal protuberances |
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4 | Parameres distinctly less than half as long as phallobase |
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– | Parameres half as long as phallobase |
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5 | Right paramere nearly straight |
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– | Right paramere strongly bent ventrally behind middle |
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Lectotype (
8 ex. “China Fujian prov. Sangang env. 3.-5./.1991 M. Nikodým leg.” (
Body length: 5.6 mm, length of elytra: 3.9 mm, body width: 3.6 mm. Body short-oval, black, elytra reddish brown, dorsal surface except anterior labroclypeus dull, pronotum and elytra glabrous.
Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex, convergent anteriorly; anterior angles strongly rounded; anterior margin shallowly sinuate medially, margins moderately reflexed; surface weakly convex, shiny, base dull, coarsely and densely punctate, with numerous erect setae; frontoclypeal suture indistinctly incised, vanishing under dull toment; smooth area in front of eye convex, nearly as long as wide; ocular canthus short and triangular (1/3 of ocular diameter), sparsely punctate, with one or more terminal setae. Frons with fine and moderately dense punctures, with a few long erect setae beside eyes and behind frontoclypeal suture. Eyes small, ratio diameter/ interocular width: 0.41. Antenna with ten antennomeres, club (♂) with four antennomeres and straight, as long as remaining antennomeres combined. Mentum convexly elevated and flattened anteriorly.
Pronotum transverse, widest at base, lateral margins in basal half nearly straight and moderately convergent to middle, evenly convex and weakly convergent anteriorly; anterior angles distinctly produced and sharp, posterior angles blunt and weakly rounded at tip; anterior margin straight, with a fine complete marginal line; surface densely and finely punctate, glabrous, with minute setae in punctures (100× magnification); lateral border densely setose; hypomeron distinctly carinate basally, not produced ventrally. Scutellum triangular, with fine, dense punctures, glabrous.
Elytra short-oval, widest shortly behind middle, striae finely impressed, finely and densely punctate, intervals weakly convex, with sparse, fine punctures concentrated along striae, glabrous except a few single, short setae on odd intervals; epipleural edge robust, ending at nearly blunt external apical angle of elytra, epipleura densely setose; apical border without a fine fringe of microtrichomes (visible at 100× magnification).
Ventral surface dull, finely and densely punctate; metasternum nearly glabrous except a few long robust setae on disc, punctures with minute setae (100× magnification); metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures, each bearing a short robust seta, last sternite half as long as penultimate one. Mesosternum between mesocoxae as wide as the mesofemur, with a semi-circular ridge bearing long setae. Ratio of length of metepisternum/ metacoxa: 1/ 1.9. Pygidium dull, moderately convex, finely and densely punctate, without smooth midline, with a few long setae along apical margin.
Legs short; femora moderately shiny, with two rudimentary longitudinal rows of setae, finely and sparsely punctate, glabrous; metafemur with anterior margin acute, without serrated line behind anterior edge, posterior margin smooth ventrally, in apical half only weakly widened, posterior margin smooth dorsally. Metatibia wide and short, widest at middle, ratio of width/ length: 1/ 2.7; dorsal margin sharply carinate, with two groups of spines, basal group at one third, apical group at three quarters of metatibial length, basally with a few short single setae; lateral face weakly convex, finely and sparsely punctate, smooth along middle; ventral edge finely serrated, with three robust nearly equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate and slightly concavely sinuate. Tarsomeres ventrally with sparse, short setae, smooth, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally, glabrous; first metatarsomere as long as following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate, distal tooth sharply pointed at apex; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex.
Aedeagus: Fig.
The colour may vary from being totally black to reddish brown. Female: antennal club also composed of 4 antennomeres, however, the club is slightly shorter than in males and the first joint of the club is slightly shorter than the club; pygidium moderately convex, at middle strongly shiny and finely punctate.
The parameres of the lectotype of
Holotype: ♂ “China coll. Chev./
Body length: 6.9 mm, length of elytra: 3.9 mm, body width: 3.6 mm. Body short-oval, dark brown, elytra black, dorsal surface except anterior labroclypeus dull, pronotum and elytra glabrous.
Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex, convergent anteriorly; anterior angles strongly rounded; anterior margin shallowly sinuate medially, margins moderately reflexed; surface weakly convex, shiny, base dull, densely punctate, coarse punctures mixed with minute ones, with numerous erect setae; frontoclypeal suture indistinctly incised, weakly curved medially; smooth area in front of eye convex, nearly as long as wide; ocular canthus short and triangular (1/3 of ocular diameter), sparsely punctate, with one or more terminal setae. Frons with fine and moderately dense punctures, with a few long erect setae beside eyes and behind frontoclypeal suture. Eyes small, ratio diameter/ interocular width: 0.42. Antenna with ten antennomeres, club (♂) with four antennomeres and straight, as long as remaining antennomeres combined. Mentum convexly elevated and flattened anteriorly.
Pronotum transverse, widest at base, lateral margins evenly convex and moderately convergent anteriorly; anterior angles distinctly produced and sharp, posterior angles blunt and weakly rounded at tip; anterior margin straight, with a fine complete marginal line; surface densely and finely punctate, glabrous, with minute setae in punctures (100× magnification); lateral border densely setose; hypomeron distinctly carinate basally, not produced ventrally. Scutellum triangular, with fine, dense punctures, glabrous.
Elytra short-oval, widest shortly behind middle, striae finely impressed, finely and densely punctate, intervals weakly convex, with sparse, fine punctures concentrated along striae, glabrous; epipleural edge robust, ending at nearly blunt external apical angle of elytra, epipleura densely setose; apical border without a fine fringe of microtrichomes (visible at 100× magnification).
Ventral surface dull, finely and densely punctate; metasternum nearly glabrous except a few long robust setae on disc, punctures with minute setae (100× magnification); metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures, each bearing a short robust seta, last sternite half as long as penultimate one. Mesosternum between mesocoxae as wide as the mesofemur, with a semi-circular ridge bearing long setae. Ratio of length of metepisternum/ metacoxa: 1/ 2.2. Pygidium dull, moderately convex, coarsely and densely punctate, without smooth midline, with a few long setae along apical margin.
Legs short; femora moderately shiny, with two rudimentary longitudinal rows of setae, finely and sparsely punctate, glabrous; metafemur with anterior margin acute, without serrated line behind anterior edge, posterior margin smooth ventrally, in apical half only weakly widened, posterior margin smooth dorsally. Metatibia wide and short, widest at middle, ratio of width/ length: 1/ 2.7; dorsal margin sharply carinate, with two groups of spines, basal group at one third, apical group at three quarters of metatibial length, basally with a few short single setae; lateral face weakly convex, finely and sparsely punctate, smooth along middle; ventral edge finely serrated, with three robust nearly equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate and slightly concavely sinuate. Tarsomeres ventrally with sparse, short setae, smooth, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally, glabrous; first metatarsomere as long as following two tarsomeres combined and as long as dorsal tibial spur. Protibia short, bidentate, distal tooth sharply pointed at apex; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex.
Aedeagus: Fig.
The name of the new species is derived from the Greek prefix “allo-” (other) and the Latin adjective “obscurus” (dark) with reference to the name and the similarity to
Lectotype (here designated): ♂ “Hainan Schmack/
1 ♂ “Qiongzhong, Hainan, Guangdong, 17.VII.1960, 400m, leg. Zhang Xuezhong” (
Body length: 6.8 mm, length of elytra: 4.8 mm, body width: 4.2 mm. Body short-oval, black, elytra reddish brown, dorsal surface except anterior labroclypeus dull, pronotum and elytra glabrous.
Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex, convergent anteriorly; anterior angles strongly rounded; anterior margin shallowly sinuate medially, margins moderately reflexed; surface weakly convex, shiny, base dull, coarsely and densely punctate, with numerous erect setae; frontoclypeal suture indistinctly incised, nearly vanishing under dull toment; smooth area in front of eye convex, nearly as long as wide; ocular canthus short and triangular (1/3 of ocular diameter), sparsely punctate, terminal setae in lectotype lacking. Frons with fine and moderately dense punctures, without erect setae. Eyes small, ratio diameter/ interocular width: 0.4. Antenna with ten antennomeres, yellowish, club (♂) with four antennomeres and straight, as long as remaining antennomeres combined. Mentum convexly elevated and flattened anteriorly.
Pronotum transverse, widest shortly before base, lateral margins in basal half nearly straight and moderately convergent to middle, evenly convex and weakly convergent anteriorly; anterior angles distinctly produced and sharp, posterior angles blunt and weakly rounded at tip; anterior margin straight, with a fine complete marginal line; surface densely and finely punctate, glabrous, with minute setae in punctures (100× magnification); lateral border densely setose; hypomeron distinctly carinate basally, not produced ventrally. Scutellum triangular, with fine, dense punctures, glabrous.
Elytra short-oval, widest shortly behind middle, striae finely impressed, finely and densely punctate, intervals weakly convex, with sparse, fine punctures concentrated along striae, glabrous except a few single, short setae on odd intervals; epipleural edge robust, ending at nearly blunt external apical angle of elytra, epipleura densely setose; apical border without a fine fringe of microtrichomes (visible at 100× magnification).
Ventral surface dull, finely and densely punctate; metasternum nearly glabrous except a few long robust setae on disc, punctures with minute setae (100× magnification); metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures, each bearing a short robust seta, last sternite half as long as penultimate one. Mesosternum between mesocoxae as wide as the mesofemur, with a semi-circular ridge bearing long setae. Ratio of length of metepisternum/ metacoxa: 1/ 2.0. Pygidium dull, moderately convex, finely and densely punctate, without smooth midline, with a few long setae along apical margin.
Legs short; femora moderately shiny, with two rudimentary longitudinal rows of setae, finely and sparsely punctate, glabrous; metafemur with anterior margin acute, without serrated line behind anterior edge, posterior margin smooth ventrally, in apical half only weakly widened, posterior margin smooth dorsally. Metatibia wide and short, widest at middle, ratio of width/ length: 1/ 2.8; dorsal margin sharply carinate, with two groups of spines, basal group at one third, apical group at three quarters of metatibial length, basally with a few short single setae; lateral face weakly convex, finely and sparsely punctate, smooth along middle; ventral edge finely serrated, with three robust nearly equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate and slightly concavely sinuate. Tarsomeres ventrally with sparse, short setae, smooth, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally, glabrous; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate, distal tooth sharply pointed at apex; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex.
Aedeagus: Fig.
The colour varies from an entirely black body, or a blackish anterior body (head and pronotum) with reddish elytra, to a nearly entirely reddish body with dark head and anterior pronotum, dorsal surface sometimes with greenish shine. Female: pygidium moderately convex, at middle strongly shiny and finely punctate; antennal club slightly shorter than the remaining antennomeres combined, composed of 4 antennomeres.
Holotype: ♂ “Tianchi, Mt. Jianfengling, Hainan, 25.IV.1980, 750m, leg. Wang Shuyong” (
Body length: 5.9 mm, length of elytra: 3.9 mm, body width: 3.6 mm. Body short-oval, dark brown partly reddish, dorsal surface except anterior labroclypeus moderately dull, pronotum and elytra glabrous.
Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex, convergent anteriorly; anterior angles strongly rounded; anterior margin shallowly sinuate medially, margins distinctly reflexed; surface nearly flat, shiny including base, coarsely and densely punctate, behind anterior margin with a few even coarser punctures each bearing an erect seta; frontoclypeal suture distinctly incised, weakly curved medially; smooth area in front of eye convex, nearly as long as wide; ocular canthus moderately short and triangular (1/3 of ocular diameter), sparsely punctate, with one terminal seta. Frons with moderately coarse and dense punctures, with a few long erect setae beside eyes and behind frontoclypeal suture. Eyes small, ratio diameter/ interocular width: 0.47. Antenna with nine antennomeres, club (♂) with four antennomeres and straight, 1.2 times as long as remaining antennomeres combined. Mentum convexly elevated and flattened anteriorly.
Pronotum transverse, widest at base, lateral margins evenly convex and moderately convergent anteriorly; anterior angles distinctly produced and sharp, posterior angles blunt and weakly rounded at tip; anterior margin straight, with a fine complete marginal line; surface densely and moderately coarsely punctate, glabrous, with minute setae in punctures (100× magnification); lateral border densely setose; hypomeron distinctly carinate basally, not produced ventrally. Scutellum triangular, with moderately coarse and dense punctures, glabrous.
Elytra short-oval, widest shortly behind middle, striae finely impressed, finely and densely punctate, intervals moderately convex, with sparse, fine punctures concentrated along striae, glabrous except a few short setae on odd intervals; epipleural edge robust, ending at rounded external apical angle of elytra, epipleura densely setose; apical border without a fine fringe of microtrichomes (visible at 100× magnification).
Ventral surface dull, finely and densely punctate; metasternum with a few short setae and long robust setae on metasternal disc; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures, each bearing a short robust seta, last sternite half as long as penultimate one. Mesosternum between mesocoxae as wide as the mesofemur, with a semi-circular ridge bearing long setae. Ratio of length of metepisternum/ metacoxa: 1/ 1.72. Pygidium dull, moderately convex, coarsely and densely punctate, without smooth midline, with a few long setae along apical margin.
Legs short; femora dull, with two rudimentary longitudinal rows of setae, finely and sparsely punctate, glabrous; metafemur shiny, with anterior margin acute, without serrated line behind anterior edge, posterior margin apically serrate ventrally, in apical half only weakly widened, posterior margin distinctly serrate dorsally. Metatibia wide and short, widest at middle, ratio of width/ length: 1/ 3.1; dorsal margin only in posterior quarter carinate, otherwise longitudinally convex, with two groups of spines, basal group at one third, apical group at three quarters of metatibial length, basally with a few short single setae; lateral face convex, finely and sparsely punctate, smooth along middle in posterior half; ventral edge finely serrated, with three robust nearly equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate and slightly concavely sinuate. Tarsomeres ventrally with sparse, short setae, smooth, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally, glabrous; first metatarsomere slightly longer than following two tarsomeres combined and distinctly longer than dorsal tibial spur. Protibia short, bidentate, distal tooth sharply pointed at apex; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex.
Aedeagus: Fig.
The new species is named after its collector, Wang Shuyong.
Among the paratypes no apparent size variation was found; colour varied from entirely reddish brown to dark brown. Female: pygidium less convex, antennal club in the paratype missing.
Holotype: ♂ “China: Hainan I., No-dong nr. Sa ko lia 12.VII.1935/ L. & M. Gressitt Collectors BISHOP Mus.” (
Body length: 6.4 mm, length of elytra: 4.4 mm, body width: 4.2 mm. Body short-oval, dark brown, ventral face reddish brown, entire surface except anterior labroclypeus dull, head with some greenish shine, pronotum and elytra glabrous.
Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex, convergent anteriorly; anterior angles strongly rounded; anterior margin shallowly sinuate medially, margins moderately reflexed; surface weakly convex, shiny, base dull, finely and densely punctate, mixed with a few larger punctures bearing each an erect seta; frontoclypeal suture distinctly incised, weakly curved medially; smooth area in front of eye convex, nearly as long as wide; ocular canthus short and triangular (1/3 of ocular diameter), sparsely punctate, with a terminal seta. Frons with fine and moderately dense punctures, with a few long erect setae beside eyes and behind lateral frontoclypeal suture. Eyes small, ratio diameter/ interocular width: 0.4. Antenna with ten antennomeres, club (♂) with four antennomeres and straight, slightly longer than remaining antennomeres combined. Mentum convexly elevated and flattened anteriorly.
Pronotum transverse, widest at base, lateral margins evenly convex and moderately convergent anteriorly; anterior angles distinctly produced and sharp, posterior angles blunt and weakly rounded at tip; anterior margin straight, with a fine complete marginal line; surface densely and finely punctate, glabrous, with minute setae in punctures (100× magnification); lateral border densely setose; hypomeron distinctly carinate basally, not produced ventrally. Scutellum triangular, with fine, dense punctures, on midline impunctate, glabrous.
Elytra short-oval, widest shortly behind middle, striae finely impressed, finely and densely punctate, intervals weakly convex, with sparse, fine punctures concentrated along striae, except a few robust setae on penultimate external intervals glabrous; epipleural edge robust, ending at nearly blunt external apical angle of elytra, epipleura densely setose; apical border without a fine fringe of microtrichomes (visible at 100× magnification).
Ventral surface dull, finely and densely punctate; metasternum nearly glabrous except a few long robust setae on disc, punctures with minute setae (100× magnification); metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures, each bearing a short robust seta, last sternite half as long as penultimate one. Mesosternum between mesocoxae as wide as mesofemur, with a semi-circular ridge bearing long setae. Ratio of length of metepisternum/ metacoxa: 1/ 2.2. Pygidium dull, moderately convex, coarsely and densely punctate, without smooth midline, with a few long setae along apical margin.
Legs short; femora moderately shiny, with two rudimentary longitudinal rows of setae, finely and sparsely punctate, glabrous; metafemur with anterior margin acute, without serrated line behind anterior edge, posterior margin smooth ventrally, in apical half only weakly widened, posterior margin smooth dorsally. Metatibia wide and short, widest at middle, ratio of width/ length: 1/ 2.6; dorsal margin sharply carinate, with two groups of spines, basal group at one third, apical group at three quarters of metatibial length, basally with a few short single setae; lateral face weakly convex, finely and sparsely punctate; ventral edge finely serrated, with three robust nearly equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate and slightly concavely sinuate. Tarsomeres ventrally with sparse, short setae, smooth, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally, glabrous; first metatarsomere slightly shorter than following two tarsomeres combined and as long as dorsal tibial spur. Protibia short, bidentate, distal tooth sharply pointed at apex; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex.
Aedeagus: Fig.
The name of the new species is derived from the type locality, Sa ko lia.
Body length: 6.4–8.1 mm, length of elytra: 4.4–5.2 mm, body width: 4.2–5.3 mm. Colour varies from entirely dark reddish brown to nearly black, often with dark pronotum and brown elytra. Female: antennal club composed of 4 antennomeres, first joint of club slightly shorter than the club, club slightly shorter than remaining antennomeres combined.
Holotype: ♂ “Hainan I. (C.): Ta Hian (TaSianKwang) 600m. VI-10-35 J.L. Gressitt” (
Body length: 6.5 mm, length of elytra: 4.3 mm, body width: 4.3 mm. Body short-oval, dark brown, elytra black, abdomen dark brown, dorsal surface except anterior labroclypeus dull, head and pronotum with some greenish shine, pronotum and elytra glabrous.
Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex, convergent anteriorly; anterior angles strongly rounded; anterior margin shallowly sinuate medially, margins moderately reflexed; surface weakly convex, shiny, base dull, finely and densely punctate, mixed with a few larger punctures bearing each an erect seta; frontoclypeal suture distinctly incised, weakly curved medially; smooth area in front of eye convex, nearly as long as wide; ocular canthus short and triangular (1/3 of ocular diameter), sparsely punctate, without terminal seta. Frons with fine and moderately dense punctures, with a few long erect setae beside eyes and behind lateral frontoclypeal suture. Eyes small, ratio diameter/ interocular width: 0.42. Antenna with ten antennomeres, club (♂) with four antennomeres and straight, slightly longer than remaining antennomeres combined. Mentum convexly elevated and flattened anteriorly.
Pronotum transverse, widest at base, lateral margins evenly convex and moderately convergent anteriorly; anterior angles distinctly produced and sharp, posterior angles blunt and weakly rounded at tip; anterior margin straight, with a fine complete marginal line; surface densely and finely punctate, glabrous, with minute setae in punctures (100× magnification); lateral border densely setose; hypomeron distinctly carinate basally, not produced ventrally. Scutellum triangular, with fine, dense punctures, glabrous.
Elytra short-oval, widest shortly behind middle, striae finely impressed, finely and densely punctate, intervals weakly convex, with sparse, fine punctures concentrated along striae, except a few robust setae on sutural interval glabrous; epipleural edge robust, ending at nearly blunt external apical angle of elytra, epipleura densely setose; apical border without a fine fringe of microtrichomes (visible at 100× magnification).
Ventral surface dull, finely and densely punctate; metasternum nearly glabrous except a few long robust setae on disc, punctures with minute setae (100× magnification); metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures, each bearing a short robust seta, last sternite half as long as penultimate one. Mesosternum between mesocoxae as wide as the mesofemur, with a semi-circular ridge bearing long setae. Ratio of length of metepisternum/ metacoxa: 1/ 2.2. Pygidium dull, moderately convex, coarsely and densely punctate, without smooth midline, with a few long setae along apical margin.
Legs short; femora moderately shiny, with two rudimentary longitudinal rows of setae, finely and sparsely punctate, glabrous; metafemur with anterior margin acute, without serrated line behind anterior edge, posterior margin smooth ventrally, in apical half only weakly widened, posterior margin smooth dorsally. Metatibia wide and short, widest at middle, ratio of width/ length: 1/ 2.7; dorsal margin sharply carinate, with two groups of spines, basal group at one third, apical group at three quarters of metatibial length, basally with a few short single setae; lateral face weakly convex, finely and sparsely punctate; ventral edge finely serrated, with three robust nearly equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate and slightly concavely sinuate. Tarsomeres ventrally with sparse, short setae, smooth, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally, glabrous; first metatarsomere slightly shorter than following two tarsomeres combined and slightly shorter than dorsal tibial spur. Protibia short, bidentate, distal tooth sharply pointed at apex; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex.
Aedeagus: Fig.
The name of the new species is derived from the type locality, Ta Hian.
Body length: 6.5–7.1 mm, length of elytra: 4.3–4.6 mm, body width: 4.3–4.7 mm. Colour varies from entirely reddish brown to nearly completely black, often reddish elytra and pronotum with a dark margin. Female: Antennal club composed of 4 antennomeres, first joint of club slightly shorter than the club, club slightly shorter than remaining antennomeres combined.
1 | Labroclypeus with a distinct transverse elevation. Antennal club only little longer than the remaining antennomeres combined. |
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– | Labroclypeus flat. Antennal club more than 1.5 times as long as the remaining antennomeres combined. |
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2 | Left paramere shorter and less widely sinuated, its tip is directed straight forward |
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– | Left paramere longer and more widely sinuated, its tip is not straight but curved interiorly. |
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Syntypes: 2 ♂♂ “Ho-chan/ coll. Thery” (
Body length: 8.1 mm, length of elytra: 4.0 mm, body width: 3.8 mm. Body short-oval, black to dark brown, antenna yellow, dorsal surface except labroclypeus dull or with iridescent or greenish shine, pronotum and elytra glabrous.
Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins strongly convex, convergent anteriorly; anterior angles strongly rounded; anterior margin distinctly sinuate medially, margins moderately reflexed; surface with a convex transverse ridge, moderately shiny, coarsely and very densely punctate, with a few erect setae anteriorly; frontoclypeal suture finely incised, evenly curved; smooth area in front of eye convex, 1.5 times as wide as long; ocular canthus short and triangular (1/3 of ocular diameter), densely and finely punctate, with one terminal seta. Frons with fine and sparse punctures, with two long erect setae beside eyes. Eyes small, ratio diameter/ interocular width: 0.51. Antenna with ten antennomeres, club (♂) with four antennomeres and straight, slightly longer than remaining antennomeres combined. Mentum convexly elevated and flattened anteriorly.
Pronotum transverse, widest at base, lateral margins evenly convex and weakly convergent anteriorly; anterior angles distinctly produced and sharp, posterior angles blunt and moderately rounded at tip; anterior margin convex, with a very fine but complete marginal line; surface densely and finely punctate, glabrous, with minute setae in punctures (100× magnification); lateral border densely setose; hypomeron distinctly carinate basally, not produced ventrally. Scutellum triangular, with fine, dense punctures, glabrous.
Elytra oval, widest in posterior third, striae finely impressed, finely and densely punctate, intervals weakly convex, with dense, fine punctures concentrated along striae, glabrous except a few single, short setae on penultimate lateral interval; epipleural edge robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border with a fine fringe of microtrichomes (visible at 100× magnification).
Ventral surface dull, finely and densely punctate; metasternum nearly glabrous except a few long robust setae on disc, punctures with minute setae (100× magnification); metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures, each bearing a short robust seta, last sternite half as long as penultimate one. Mesosternum between mesocoxae as wide as the mesofemur, with a semi-circular ridge bearing long setae. Ratio of length of metepisternum/ metacoxa: 1/ 1.49. Pygidium dull, strongly convex, finely and densely punctate, without smooth midline, with a few long setae along apical margin.
Legs short; femora moderately shiny, with two longitudinal rows of setae, finely and sparsely punctate, glabrous; metafemur with anterior margin acute, with a continuous, serrated line behind anterior edge, posterior margin smooth ventrally, in apical half only weakly widened, posterior margin smooth dorsally. Metatibia moderately wide and short, widest at middle, ratio of width/ length: 1/ 2.95; dorsal margin sharply carinate, with two groups of spines, basal group shortly behind middle, apical group at 4/5 of metatibial length, in basal half with a serrated line beside dorsal margin ending at basal group of spines, beside it with a few single short setae; lateral face longitudinally convex, finely and densely punctate, smooth along middle; ventral edge finely serrated, with three robust nearly equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate and slightly concavely sinuate. Tarsomeres sparsely and finely punctate dorsally, with sparse, short setae ventrally, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally, glabrous; first metatarsomere slightly shorter than following two tarsomeres combined and one third of its length longer than dorsal tibial spur. Protibia short, bidentate, distal tooth sharply pointed at apex, external margin bluntly widened at middle; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex.
Aedeagus: Fig.
The colour varies from totally black or reddish brown to black with reddish brown elytra. Female: Antennal club also composed of 3 antennomeres, however, the club is slightly shorter than in males, and the first joint of club is slightly shorter.
Syntype: ♂ “N. E. Burma Kambaiti 7000 ft. 24/5.1934/ N. E. Burma R. Malaise B. M. 1945-71/Co-Type/
1 ♂ “China, W Yunnan prov., mts. 60km E Tengchong, 2200m, 19.-22.v.2006, S. Murzin & I. Shokhin” (
Body length: 7.2 mm, length of elytra: 5.2 mm, body width: 4.4 mm. Body short-oval, black to dark brown, antenna yellow, dorsal surface except labroclypeus dull, pronotum and elytra glabrous.
Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex, convergent anteriorly; anterior angles moderately rounded; anterior margin distinctly sinuate medially, margins moderately reflexed; surface nearly flat, moderately shiny, coarsely and very densely punctate, with a few erect setae anteriorly; frontoclypeal suture finely incised, convexly bent at middle; smooth area in front of eye convex, 1.2 times as wide as long; ocular canthus short and triangular (1/3 of ocular diameter), densely and finely punctate, with one terminal seta. Frons with coarse and dense punctures, glabrous. Eyes small, ratio diameter/ interocular width: 0.51. Antenna with ten antennomeres, club (♂) with four antennomeres and straight, 1.7 times as long as remaining antennomeres combined. Mentum convexly elevated and flattened anteriorly.
Pronotum transverse, widest at base, lateral margins in basal half nearly straight and moderately convergent, in anterior half evenly convex and weakly convergent anteriorly; anterior angles distinctly produced and sharp, posterior angles blunt and moderately rounded at tip; anterior margin convex, with a very fine but complete marginal line; surface densely and finely punctate, glabrous, with minute setae in punctures (100× magnification); lateral border densely setose; hypomeron distinctly carinate basally, not produced ventrally. Scutellum triangular, with fine, dense punctures, glabrous.
Elytra oval, widest in posterior third, striae finely impressed, finely and densely punctate, intervals weakly convex, with dense, fine punctures concentrated along striae, glabrous except a few single, short setae on penultimate lateral interval; epipleural edge robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border with a fine fringe of microtrichomes (visible at 100× magnification).
Ventral surface dull, finely and densely punctate; metasternum nearly glabrous except a few long robust setae on disc, punctures with minute setae (100× magnification); metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures, each bearing a short robust seta, last sternite half as long as penultimate one. Mesosternum between mesocoxae as wide as the mesofemur, with a semi-circular ridge bearing long setae. Ratio of length of metepisternum/ metacoxa: 1/ 1.52. Pygidium dull, strongly convex, finely and densely punctate, without smooth midline, with a few long setae along apical margin.
Legs moderately long; femora moderately shiny, with two longitudinal rows of setae, finely and sparsely punctate, glabrous; metafemur with anterior margin acute, with a continuous, serrated line behind anterior edge, posterior margin smooth ventrally, in apical half only weakly widened, posterior margin smooth dorsally. Metatibia narrow and moderately long, widest at middle, ratio of width/ length: 1/ 3.35; dorsal margin sharply carinate, with two groups of spines, basal group shortly behind middle, apical group at 4/5 of metatibial length, in basal half with a serrated line beside dorsal margin ending at basal group of spines, beside it with a few single short setae; lateral face longitudinally convex, finely and densely punctate, smooth along middle; ventral edge finely serrated, with three robust nearly equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate and slightly concavely sinuate. Tarsomeres sparsely and finely punctate dorsally, with sparse, short setae ventrally, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally, glabrous; first metatarsomere slightly shorter than following two tarsomeres combined and one third of its length longer than dorsal tibial spur. Protibia short, bidentate, distal tooth sharply pointed at apex, external margin bluntly widened at middle; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex.
Aedeagus: Fig.
Holotype: ♂ “[China] Yunnan, Yakou, 2012-V-11/ LW-1319” (
Body length: 8.0 mm, length of elytra: 6.1 mm, body width: 5.1 mm. Body short-oval, black to dark brown, antenna yellow, dorsal surface except labroclypeus dull, pronotum and elytra glabrous.
Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins strongly convex, convergent anteriorly; anterior angles moderately rounded; anterior margin distinctly sinuate medially, margins moderately reflexed; surface nearly flat, shiny, finely and very densely punctate, with a few erect setae anteriorly; frontoclypeal suture finely incised, convexly bent at middle; smooth area in front of eye convex, 1.3 times as wide as long; ocular canthus long and subtriangular (nearly half of ocular diameter), densely and finely punctate, with one terminal seta. Frons with moderately coarse and dense punctures, with two single erect setae beside eyes. Eyes small, ratio diameter/ interocular width: 0.54. Antenna with ten antennomeres, club (♂) with four antennomeres and straight, 1.7 times as long as remaining antennomeres combined. Mentum convexly elevated and flattened anteriorly.
Pronotum transverse, widest at base, lateral margins evenly convex and weakly convergent anteriorly; anterior angles distinctly produced and sharp, posterior angles blunt and moderately rounded at tip; anterior margin convex, with a very fine but complete marginal line; surface densely and finely punctate, with minute setae in punctures (100× magnification); lateral border densely setose; hypomeron distinctly carinate basally, not produced ventrally. Scutellum triangular, with fine, dense punctures, punctures on basal midline less dense, with minute setae in punctures.
Elytra oval, widest in posterior third, striae finely impressed, finely and densely punctate, intervals weakly convex, with dense, fine punctures concentrated along striae, glabrous except a few single, short setae on penultimate lateral interval; epipleural edge robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border with a fine fringe of microtrichomes (visible at 100× magnification).
Ventral surface dull, finely and densely punctate; metasternum nearly glabrous except a few long robust setae on disc, punctures with minute setae (100× magnification); metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures, each bearing a short robust seta, last sternite half as long as penultimate one. Mesosternum between mesocoxae as wide as the mesofemur, with a semi-circular ridge bearing long setae. Ratio of length of metepisternum/ metacoxa: 1/ 1.5. Pygidium dull, strongly convex, coarsely and densely punctate, without smooth midline, with a few long setae on apical quarter, otherwise punctures with minute setae.
Legs moderately long; femora moderately shiny, with two longitudinal rows of setae, finely and sparsely punctate, glabrous; metafemur with anterior margin acute, with a continuous, serrated line behind anterior edge, posterior margin smooth ventrally, in apical half only weakly widened, posterior margin smooth dorsally. Metatibia narrow and moderately long, widest at middle, ratio of width/ length: 1/ 3.75; dorsal margin sharply carinate, with two groups of spines, basal group at 3/5, apical group at 4/5 of metatibial length, in basal half with a serrated line beside dorsal margin ending at basal group of spines, beside it with a few single short setae; lateral face longitudinally convex, finely and densely punctate, smooth along middle; ventral edge finely serrated, with three robust nearly equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate and slightly concavely sinuate. Tarsomeres sparsely and finely punctate dorsally, with sparse, short setae ventrally, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally, glabrous; first metatarsomere slightly shorter than following two tarsomeres combined and one third of its length longer than dorsal tibial spur. Protibia short, bidentate, distal tooth sharply pointed at apex, external margin bluntly widened at middle; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex.
Aedeagus: Fig.
The name of the new species is the combined Greek prefix “pseudo-” (false) and the species name “
Distribution of the species of the
Part of this research was supported by the National Natural Science Foundation of China (No. 31501889, 31672345), Research Equipment Development Project of