Corresponding author: Mingyi Tian (
Academic editor: B. Guéorguiev
The
Tian M, Deuve T (2016) A review of the
As part of the series works on the dominant genus
The members of the
All of the four species of this group are distributed in Southeast Asia. The first species of this group is
All specimens for this study are dry and mounted material. Dissection and observation of the specimens were made using a WILD M32 binocular microscope. Detailed descriptions are provided for the new species, while only diagnostic character states are given for the known species. Habitus and male genital illustrations for all species are also presented. Digital photographs were taken and processed as in
Body length was measured from apex of right mandibles to apex of elytra; body width = width of elytra.
Abbreviations of measurements used in the text are as followings:
head length (from apex of right mandible to base of vertex)
head width (maximum distance across head, including eyes)
length of pronotum (measured from front to basal margins, through midline)
width of pronotum (greatest width of pronotum)
length of elytra (measured from base to apex of elytra, through suture)
width of elytra (greatest transverse distance across both elytra)
The depository abbreviations used in the text are as following:
Collection of Dr. Ingo Brunk, Dresden, Germany
Collection of Mr. Pierre Morvan (Carentoir, France)
Large-sized and robust; body brown or yellowish brown, moderately convex; glabrous, impunctate on head and pronotum; head wide, eyes moderate large and prominent; mentum bisetose in most species (except for
Protarsi and abdominal ventrite VII in male of
1 | Median and posterior setiferous pores of the 3rd elytral interval located on median portion, never close to the 2nd elytra stria |
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– | Median and posterior setiferous pores of the 3rd elytral interval close to the 2nd elytra stria |
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2 | Mentum asetose, the 3rd elytral interval with only median and posterior setiferous pores, anterior one wanted |
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– | Mentum bisetose, the 3rd elytral interval with three setiferous pores |
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3 | Prosternal process unbordered at apex, inner angle of elytra sharp |
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– | Prosternal process well-bordered at apex, inner angle of elytra blunt |
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4 | Labrum slightly emarginated at frontal margin, the 4th hind tarsomere with a deeper emargination, outer lobe half as long as the joint |
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– | Labrum strongly emarginated at frontal margin, 4th hind tarsomere with a shallower emargination, outer lobe shallower, about 1/3 as long as the joint |
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Length: 15.0 mm; width: 7.0 mm. Habitus as in Fig.
Head dark brown, other parts of the body yellowish brown; head stout, as long as wide; labrum slightly emarginated at frontal margin; clypeus even near base, with two longitudinal furrows along both side, joining to frontal impressions respectively; mentum bisetose; pronotum transverse,
Male genitalia (Fig.
Female: Unknown.
Easily separated from its congeners,
1 male, the holotype, “Nord Borneo, Mont Kina Balu, 5-8, 1903, John Waterstradt”, in
Malaysia (Sarawak).
Length: 14.0 mm; width: 5.5 mm. Habitus as in Fig.
Elytra and sides of pronotum yellowish brown, head dark brown, pronotum light dark brown on disc; head as long as wide, clypeus smooth, front margin of labrum slightly emarginated; mentum bisetose; pronotum strongly transverse,
Male genitalia (Fig.
Allied to
1 male, the holotype, “Perak”, in the Collection of Bates,
Malaysia (Perak).
Male, “Thailand South, near Khao Sok NP, 3-6. II. 1997, A. Kudrna lgt”, in
Length: 14.0 mm; width: 5.5 mm. Habitus as in Fig.
Yellowish brown, with head dark brown, the 2nd–11th antennomeres, disc of pronotum, tibiae and tarsi of legs brown; surface glabrous, impuctate, head and pronotum finely striate, the 3rd, 5th and 7th elytral intervals each with a few tiny sparse punctures; surface moderately shiny; microsculptural engraved meshes densely isodiametric on the surface of head, pronotum and elytra; body rather flat.
Head stout, as long as wide; eyes very large and prominent; frons flat, vertex slightly convex, smooth; frontal impressions very small, foveate; clypeus bisetose, rather smooth near base; labrum 6-setose, evidently emarginate at apical margin; ligula very small and narrow, bisetose at apex; palps slender or elongate, subcylindrical, normally setose; palpiger asetose, mentum edentate, mentum and submentum each with one pair of setae, mental ones much shorter; antennae slender and long, reaching 1/4 of elytra from base, pubescent from apical 2/3 of the 4th antennomere; the 3rd antennomere as long as the 4th, both shorter than the 1st; the 1st–3rd antennomeres glabrous; evidently expanded laterally in the 4th and 5th antennomere; the 1st antennomere 1.6 times as long as the 2nd.
Pronotum strongly transverse,
Elytra elongate ovate, much longer than wide,
Legs moderately developed; hind femur long and elongate, with 2 setae posteriorly; middle and hind coxae smooth and glabrous; fore tibia deeply sinuate at apical margin, outer angle sharp, outer margin distinctly serrate; middle tibia slender, not dilated medially in male; hind tibia slender, apex slightly dilated, apical spurs very long and sharp, the 1st tarsomere slightly longer than the 2nd, the 3rd tarsomere about 1.2 times longer than the 4th which is bilobed at apex, with outer lobe slightly shorter than half as long as the joint, outer lobe evidently longer than the inner; all tarsal claws moderately pectinate.
Male genitalia (Fig.
Prosternal process bordered at apex; apical margin of abdominal ventrite VII feebly emarginated in male.
Female: Unknown.
Close to
To be dedicated to Mr. Pierre Morvan (Carentoir, France), a good expert of
Southern Thailand.
male, “Malaisie, Sabah, Crocker Range, Avril 94, Chew”, in
Length: 16.0–17.0 mm; width: 6.5–6.6 mm. Habitus as in Fig.
Head dark brown, other parts reddish brown; smooth and glabrous, impunctate, elytral intervals covered with fine and sparse punctures; head and pronotum faintly wrinkled; microsculptural engraved meshes isodiametric on head, pronotum and elytra.
Head stout, slightly longer than wide,
Pronotum transverse, notably wider than long,
Elytra elongate-ovate, distinctly longer than wide,
Femora moderately expanded, hind femur bisetose posteriorly; for tibia dilated at apex, apical margin obtusely truncate or sinuate towards outer angle, outer margin notably serrate; middle tibia slightly curved; hind legs slender, apical spurs long and sharp, the 1st and 3rd tarsomeres much longer than the 2nd and 4th respectively, the 4th tarsomere deeply emarginated at apex, with lobes about half as long as the joint; fore and middle tarsal claws weakly pectinate, hind claws simple.
Prosternal process bordered at apex; abdominal ventrite VII finely and shallowly emarginate at apical margin in male.
Male genitalia (Fig.
It is a very peculiar species, differing from other congeners by its mentum asetose, elytral base incompletely bordered, anterior dorsal pores missed on the 3rd interval, and hind tarsal claws simple.
To refer to the type locality.
Malaysia (Sabah).
We would like to thank Azadeh Taghavian (