Corresponding author: Jakub Straka (
Academic editor: M. Ohl
Straka J (2016)
Material from the following institutions and private collections was examined:
British Museum, (Natural History), London, Great Britain (David Notton)
Christian Schmid-Egger, Berlin, Germany
Institute of Biodiversity and Ecosystem Research, Sofia, Bulgaria (Toshko Ljubomirov)
Jakub Straka, Praha, Czech Republic
Hungarian Natural History Museum, Budapest, Hungary (Sandor Csősz)
Martin Říha, Brno, Czech Republic
Naturhistorisches Museum Wien, Wien, Austria (Manuela Vizek)
Niklas Johansson, Habo, Sweden
National Museum, (Natural History), Praha, Czech Republic (Jan Macek)
Oberösterreichisches Landesmuseum, Linz, Austria (Fritz Gusenleitner)
Piotr Olszewski, Toruń, Poland
Pavel Tyrner, Litvínov, Czech Republic
Upper Silesian Museum, Bytom, Poland (Waldemar Żyła)
Werner Arens, Bad Hersfeld, Germany
Wolfgang Schlaefle, Kaiseraugst, Switzerland
Zoological Institute, Russian Academy of Sciences, St. Petersburg, Russia (Yulia Astafurova)
Zdeněk Karas, Zliv, Czech Republic
Staatliche Naturwissenschaftliche Sammlungen Bayerns, Zoologische Staatssammlung, Munich, Germany (Stefan Schmidt)
Morphological terms are used according to
clypeus median lobe width
clypeus maximum length
clypeus width
length of antennal article III, dorsally
width of antennal article III, dorsoapically
length of antennal article V, dorsally
width of antennal article V, dorsoapically
vertex width
vertex length
diameter of median ocellus
forefemur length
forefemur width, laterally
All of the newly described species were labeled in the following manner: “HOLOTYPUS ♂ or ♀, name of taxon sp. nov., J. Straka det. 2015” on a red card; neotypes were designated on red cards, and paratypes were designated on yellow cards. Exact label data are cited only for the holotypes. Separate lines on the labels are indicated with a slash “/”, and separate labels are indicated with double slashes “//”.
From the other species of the
Austria, Czech Republic, Germany, Hungary, Kazakhstan, Mongolia, Poland and Slovakia.
The holotype was redescribed by
The female of
Body length: 8.2 mm.
Head. Clypeus conspicuously convex, highly elevated, its nearly angular top located slightly dorsally from clypeal midlength; basomedian area well developed, as densely and finely punctate as lateral section, with intermixed large punctures; bevel nearly flat, conspicuously declining, with few large punctures, much longer medially than laterally, but not reaching base of clypeus, bright shiny; lip very slightly arcuate to nearly straight, with median emargination and well developed lateral incisions, separated from bevel by distinct punctate groove,
Mesosoma. Scutum without distinct anterior impression; scutum and scutellum with well defined punctures variable in size, less than one diameter apart, interspaces variable in size, shiny, setae gold-brown, about 0.5 ×
Metasoma. Terga I-III with slightly developed, but distinct silvery apical fasciae, densely, very finely micropunctate, punctures ill-defined, evanescent in microsculpture, interspaces aciculate, slightly shiny; apical depressions shallow with indistinct micropunctures; sculpture of terga VI-V distinctly coarser than on previous terga, punctures denser, ill-defined, but large, one to less than half diameter apart, interspaces microsculptured, apical depression of tergum V coarsely microsculptured, with few scattered punctures. Pygidium of usual size, sparsely punctate, large and minute punctures intermixed, ill defined, but large punctures deep, interspaces nearly unsculptured, shiny. Central part of sternum II with several larger punctures, interspaces microsculptured, shiny; lateral part slightly shiny, densely micropunctate; remaining sterna with uniform sculpture similar to that on sternum II, but more or less reduced laterally.
Coloration. Large area in center of mandibles, terga and sterna I-III and base of tergum and sternum IV red. Distal tarsomeres, apex of pronotal lobe, ventral part of distal antennal segments, apex of clypeal bevel and tip of pygidium dark red. Tegulae reddish translucent. Apical parts of terga I-III slightly translucent. Remaining body parts all black.
Known only from Hungary.
Male unknown.
The species is named after the country of origin.
Kazakhstan, Kyrgyzstan and Turkey.
Full description and all material is given in
Body length: 6.7 mm.
Head. Clypeus markedly convex, medially uniformly curved, top at clypeal midlength; basomedian area large, densely punctate; bevel well developed, nearly flat and declining, bright shiny, with several larger punctures; lip conspicuously arcuate, well developed medially, with well-developed, nearly rectangular, lateral corner, lip separated from bevel by distinct groove with large punctures;
Mesosoma. Scutum without distinct anterior impression; scutum and scutellum densely punctate, punctures well defined, most punctures less than half diameter apart, interspaces distinct, unsculptured, slightly shiny, setae about 1.0 ×
Metasoma. Terga I-III with silvery apical fasciae. Apical depressions of all terga shallow, with micropunctures. Terga I-III densely and distinctly micropunctate, punctures ill defined, interspaces microsculptured, slightly shiny to dull; sculpture of tergum IV-VII coarser than on other terga, also punctures denser. Sterna uniformly punctate, nearly like terga.
Coloration. Apical part of mandibles and terga I and II red. Distal tarsomeres, forefemoral notch, sternum I and base of sternum II dark red. Tegulae reddish translucent. Apical parts of all terga slightly translucent. Remaining body parts black.
Variation of males. Body length: 6.5–7.0 mm. Clypeus slightly to conspicuously convex medially. Forefemoral notch and all metasoma dark in some specimens.
Body length: 7.6–8.2 mm.
Head. Clypeus conspicuously convex, highly elevated, its top slightly dorsally from clypeal midlength; basomedian area large, densely and finely punctate as in lateral section, with intermixed large punctures; bevel nearly flat, markedly declining, with few large, sparse punctures, much longer medially than laterally, not reaching base of clypeus, bright shiny; lip very slightly arcuate to nearly straight, with small median emargination and well developed lateral incisions, separate from bevel by distinctly punctate groove,
Mesosoma. Scutum without distinct anterior impression; scutum and scutellum densely punctate, punctures well defined, uniform in size, about half diameter apart (up to one diameter apart in some specimens), interspaces uniform in size, slightly shiny, setae about 0.5 ×
Metasoma. Terga I-III with silvery apical fasciae; densely and very finely micropunctate, punctures ill defined, evanescent in microsculpture, interspaces very finelly microsculptured, slightly shiny; apical depressions shallow, with indistinct micropunctures; sculpture of terga IV-V distinctly coarser than on previous terga, also punctures denser; terga IV-V coarsely punctate, punctures ill-defined, but large, one to less than half diameter apart, interspaces microsculptured, apical depression of terga IV and V coarsely microsculptured, apical depression of tergum IV very finely punctate, apical depression of tergum V with a few scattered punctures, slightly shiny, but coarsely microsculptured and dull in some specimens. Pygidium relatively wide, sparsely punctate, with large and minute punctures intermixed, large punctures well defined, deep, interspaces nearly unsculptured, shiny. Central part of sternum II with several larger punctures, interspaces microsculptured, shiny; lateral part slightly shiny, densely micropunctate; remaining sterna with uniform sculpture similar to that on sternum II, but more or less reduced laterally.
Coloration. Central part of mandibles and terga and sterna I-II red. Distal tarsomeres dark reddish. Tegulae brown translucent. Apical parts of terga I-III slightly translucent. Remaining body parts black.
France, Spain, Turkey.
The species is named in honor of the outstanding entomologist and collector of most of the type specimens, Jane van der Smissen.
Body length: 7.3 mm.
Head. Clypeus uniformly convex, distinctly elevated in midlength; basomedian area large, densely, distinctly punctate; bevel small, decreasing toward clypeal lip, shiny, sparsely punctate; lip uniformly, distinctly arcuate, not longer medially than laterally, with distinct lateral corner, lip separated from bevel by a groove;
Mesosoma. Scutum without distinct anterior impression; scutum and scutellum densely punctate, punctures well defined, most punctures less than half diameter apart, interspaces distinct, anteriorly microsculptured, slightly shiny to dull, setae less than 1.0 ×
Metasoma. Silvery apical fasciae developed faintly on terga I-IV. Apical depressions of terga distinct, sparsely micropunctate, nearly like more anterior parts. Terga I-IV sparsely micropunctate, punctures ill defined, interspaces coarsely microsculptured, slightly shiny to dull; sculpture of terga IV-VII slightly more conspicuous than that of other terga, also densely punctate. Sterna uniformly punctate, nearly like terga. Volsella light brown, ventral setae pointing in various directions in medial part, but almost in one direction at apical process; apical process relatively sparsely setose; dorsal process large, wider than corpus; corpus lined with distinct ledge along base of setae on inner side of corpus.
Coloration. Apical part of mandibles, distal tarsomeres and terga I-III red. Forefemoral notch black. Tegulae reddish translucent. Apical parts of all terga slightly translucent. Remaining body parts black.
Body length: 10.3 mm.
Head. Clypeus distinctly convex, top at clypeal midlength; basomedian area relatively large, more sparsely punctate and punctures larger than on lateral section, interspaces between punctures well developed, shiny, larger medially than laterally, punctures mostly less than one diameter apart; bevel large, distinctly convex, medially longer than basomedian area, laterally much shorter, obtusely triangular, with numerous large, sparse punctures, not reaching base of clypeus, shiny; lip distinctly arcuate, with small lateral incisions, without medial emargination, separate from bevel by punctate groove,
Mesosoma. Scutum without distinct anterior impression; scutum and scutellum densely punctate, punctures well defined, most punctures less than half diameter apart, interspaces distinct, slightly shiny to shiny, setae less than 1.0 ×
Metasoma. Terga I-III with slightly developed but distinct silvery apical fasciae. Apical depressions of all terga well developed, very sparsely micropunctate. Terga I-III sparsely micropunctate, punctures ill-defined but distinct, interspaces microsculptured, slightly shiny; sculpture of tergum IV-V distinctly coarser than on previous terga, also slightly more densely punctate. Pygidium sparsely punctate, punctures variable, some ill defined, some well defined, but shallow, interspaces microsculptured medially, unsculptured laterally, slightly shiny to shiny. Central part of sternum II with several larger punctures, interspaces microsculptured, shiny; lateral part slightly shiny, densely micropunctate; remaining sterna with uniform sculpture similar to that on sternum II, but more or less reduced laterally.
Coloration. Central part of mandibles, distal tarsomeres, terga and sterna I-III red. Tegulae brown translucent. Apical parts of terga I-III slightly translucent. Remaining body parts black.
Known only from Czech Republic.
The species is named after the Bohemian part of the Czech Republic, where the type specimens were collected.
Body length: 8.1 mm.
Head. Mandibular inner margin shallowly emarginated distally from inner tooth, with no furrow. Clypeus uniformly convex, slightly elevated, top at clypeal midlength; basomedian area large, sparsely punctate, with nearly impunctate line medially; bevel convex, decreasing to clypeal lip, shiny, with several larger punctures; lip distinctly sinuate, conspicuously produced medially, with distinct lateral corner, lip separated from bevel by punctate groove;
Mesosoma. Scutum without anterior impression; scutum and scutellum densely punctate, punctures well defined, most punctures less than half diameter apart, interspaces distinct, microsculptured, slightly shiny, setae about 1.0 ×
Metasoma. Silvery apical faciae of terga I-IV distinctly developed. Apical depressions of all terga shallow, nearly as densely micropunctate as more anterior parts. Terga I-III densely and distinctly micropunctate, punctures ill defined, interspaces microsculptured, slightly shiny; sculpture of terga IV-VII coarser than on previous terga, also puncture denser. Sterna uniformly punctate, nearly like terga. Volsella brown, ventral setae pointing in various directions; apex with numerous setae; dorsal process about as wide as corpus.
Coloration. All body black, except central part of mandibles and terminal tarsomeres dark red. Tegulae brown translucent. Apical parts of all terga slightly translucent.
Known only from the island of Crete, Greece.
Female unknown. Only two males have been collected so far.
The species is named after the island of Crete, Greece, where the specimens were collected.
Body length: 5.0–8.0 mm.
Head. Mandibular inner margin with one rectangular tooth and distinct furrow next to tooth distally. Labrum flat. Clypeus convex, conspicuously elevated in basal half and continuously declining apically, top at clypeal midlength or in basal half of clypeus; basomedian area large, densely punctate, sparsely punctate medially in some specimens; bevel slightly convex to nearly flat, variable in size, shiny with several larger punctures; lip slightly arcuate, short, with small lateral incisions, lip separated from bevel by variable fine groove;
Mesosoma. Scutum without distinct anterior impression; scutum and scutellum densely punctate, punctures well defined, most punctures less than one diameter apart, interspaces distinct, unsculptured, shiny, setae about 1.0 ×
Metasoma. Terga I-III with silvery apical faciae. Apical depressions of all terga shallow, micropunctate. Terga I-III sparsely, finely micropunctate, punctures evanescent in microsculpture, slightly shiny; sculpture of tergum IV-VII coarser, than on previous terga, also punctation slightly denser. Sterna uniformly punctate nearly like terga. Volsella light brown, ventral setae pointing in various directions; dorsal process slightly wider than corpus in most specimens.
Coloration. Apical part of mandibles, tarsi, forefemoral notch, tegulae, terga and sterna I, II, and partly or all tergum III red. Apical parts of all terga and tegula translucent. Remaining body parts all black. Forefemoral notch very rarely black in specimens from high altitude mountains.
(neotype). Body length: 7.4 mm.
Head. Clypeus distinctly convex, top at clypeal midlength; basomedian area relatively large, densely punctate as lateral section; bevel large, distinctly convex, triangular, with sparse, large punctures, reaching base of clypeus, shiny; lip slightly arcuate, with lateral incisions, separate from bevel by punctate groove,
Mesosoma. Scutum without distinct anterior impression; scutum and scutellum densely punctate, punctures well defined, half to one diameter apart, interspaces slightly microsculptured, shiny, setae about 1.0 ×
Metasoma. Terga I-III with silvery apical fasciae. Apical depressions of all terga shallow, with micropunctures evanescent in microsculpture. Terga I-III sparsely and finely micropunctate, punctures ill defined, interspaces microsculptured, shiny to slightly shiny; sculpture of tergum IV-V slightly coarser than on previous terga, also slightly more densely punctate. Pygidium sparsely punctate, punctures ill defined, interspaces microsculptured, slightly shiny. Central part of sternum II with several larger punctures, interspaces microsculptured, shiny; lateral part slightly shiny, densely micropunctate; remaining sterna with uniform sculpture similar to that on sternum II, but sculpture more or less reduced laterally.
Coloration. Central part of mandibles, three distal tarsomeres, terga and sterna I-III red. Tegulae brown translucent. Apical parts of terga I-III slightly translucent. Remaining body parts all black.
Variation of females. Extremely variable in most of characters. The clypeus is typically conspicuously elevated basally, with the top in the basal third; the bevel can be well separated from the clypeal base in some specimens, althouth infrequently. Vertex wider than long in large specimen and slightly wider than long in small specimens,
Austria, Czech Republic, Bulgaria, France, Germany, Italy, Sweden and Switzerland. Specimens from mountains of Greece, Morocco and Spain probably also belong to this species.
Holotypus: ♂, Turkey: Trabzon: Boztepe,
Georgia, Russia (Caucasus) and Turkey.
Full description of the species is presented by
Body length: 7.2 mm.
Head. Clypeus distinctly convex, its top at clypeal midlength; basomedian area relatively large, as densely punctate as lateral section; bevel large, distinctly convex, as long as basomedian area laterally, slightly longer medially, obtusely triangular, with sparse large punctures, not reaching base of clypeus, shiny; lip slightly arcuate, with lateral incision, separated from bevel by punctate groove,
Mesosoma. Scutum without distinct anterior impression; scutum and scutellum densely punctate, punctures well defined, less than half to one diameter apart, interspaces well developed, variable in size, microsculptured, slightly shiny, setae about 1.0 ×
Metasoma. Terga I-III with ill defined silvery apical fasciae. Terga I-III densely, finely micropunctate, punctures ill defined, interspaces microsculptured, slightly shiny; apical depressions shallow, with distinct micropunctures; sculpture of terga IV-V distinctly coarser than on previous terga, also slightly more densely punctate; tergum V coarsely punctate, punctures ill-defined, but large, half to one diameter apart, interspaces microsculptured, apical depression of tergum V very sparsely and finely punctate. Pygidium large, sparsely punctate, punctures large, ill defined, interspaces nearly unsculptured, shiny. Central part of sternum II with several larger punctures, interspaces microsculptured, shiny; lateral part slightly shiny, densely micropunctate; remaining sterna with uniform sculpture similar to that on sternum II, but more or less reduced laterally.
Coloration. Central part of mandibles, terga and sterna I-III red. Distal tarsomeres dark red. Tegulae reddish translucent. Apical parts of terga I-III slightly translucent. Remaining body parts black.
Variation of females. Very variable in sculpture, but finely sculptured over all body in most specimens except apical terga (in comparison to other species). Clypeal bevel reaching base of clypeus in some specimens, but well separated from the base in most. Antenna short,
Body length: 5.6–8.7 mm.
Head. Mandibular inner margin with one distinct rectangular tooth and distinct furrow next to tooth distally. Clypeus slightly convex, uniformly curved, or more steeply declining ventrally than dorsally, top about in clypeal midlength; basomedian area very large, about two thirds of bevel or more, densely punctate; bevel small, slightly convex, shiny with several larger punctures; lip conspicuously arcuate or slightly sinuate, with small or large lateral corner, lip separated from bevel by distinct groove with large punctures;
Mesosoma. Scutum without distinct anterior impression; scutum and scutellum densely punctate, punctures well defined, most punctures less than half diameter apart, interspaces distinct, microsculptured, slightly shiny to dull, setae about 1.0 ×
Metasoma. Terga I-III with silvery, apical fasciae. Apical depressions of all terga distinct, with micropunctures similar as on central part of terga. Terga I-III densely and distinctly micropunctate, punctures ill defined, interspaces microsculptured, slightly shiny; sculpture of terga IV-VII slightly coarser and slightly more densely punctate than other terga. Sterna uniformly punctate nearly like terga.
Coloration. Apical part of mandibles, terga and sterna I and II and forefemoral notch red. Metasoma very variable in coloration; some specimens with only tergum I dark red, or terga I and II and sternum II basally red, or terga and sterna I-III equally red. Distal tarsomeres dark reddish. Tegulae brown to reddish. Apical parts of all terga and tegulae translucent. Remaining body parts black.
Austria, France, Germany, Hungary, Poland, Slovakia and Sweden.
Neotypus: ♀, Czech Republic: “CZ, Mor.mer.
(neotype). Body length: 8.9 mm.
Head. Clypeus distinctly convex, top at clypeal midlength; basomedian area relatively large, as sparsely punctate as lateral section, punctures small, ill-defined, interspaces variable, one to less than half diameter apart, shiny; bevel large, distinctly convex, triangular, with sparse large punctures, not reaching base of clypeus, shiny; lip distinctly sinuate, long medially, with poorly developed lateral incision, lip separate from bevel by punctate groove,
Mesosoma. Scutum without distinct anterior impression; scutum and scutellum densely punctate, punctures well defined, less than one diameter apart, interspaces variable in size, microsculptured, slightly shiny, setae about 1.0 ×
Metasoma. Terga I-III with distinct but sparse silvery, apical faciae. Terga I-III densely micropunctate, with very large, but superficial, sparse punctures, micropunctures ill-defined, evanescent in micosculpture, faint, interspaces microsculptured, slightly shiny; apical depressions well developed, with distinct micropunctures; sculpture of terga III-V distinctly coarser than other terga, also more densely punctate; tergum V coarsely punctate, punctures ill defined, but large, half to one diameter apart, interspaces microsculptured, apical depression of tergum V distinctly, densely finely punctate. Pygidium distinctly narrower than in other related species, integument sparsely punctate, large and small punctures intermixed, punctures ill defined, interspaces nearly unsculptured, bright shiny. Central part of sternum II with several large punctures, interspaces microsculptured, shiny; lateral part slightly shiny, densely micropunctate; remaining sterna with uniform sculpture similar to that on sternum II, but more or less reduced laterally.
Coloration. Central part of mandibles, terga I-II and distal tarsomeres dark red. Tegulae brown translucent. Apical parts of terga I-III slightly translucent. Remaining body parts all black.
Variation of females: Body length: 7.0–10.1 mm.
Body length: 6.4–7.9 mm.
Head. Mandibular inner margin with one distinct rectangular tooth and distinct furrow next to tooth distally. Clypeus convex, slightly elevated, distinctly declining apically, top variably located between apical third and midlength; basomedian area large, sparsely punctate, more sparsely punctate medially in some specimens; bevel convex, decreasing toward clypeal lip, variable in size, reaching base of clypeus in most specimens, shiny, with several large punctures; lip distinctly sinuate, conspicuously produced medially, with more or less distinct lateral corner, lip separated from bevel by groove;
Mesosoma. Scutum with shallow anterior impression, densely punctate, punctures well defined, most punctures less than half diameter apart, interspaces distinct, microsculptured, slightly shiny to dull, setae about 1.0 ×
Metasoma. Silvery apical fasciae of terga I-III distinctly developed, but fasciae brown in specimens from eastern Turkey. Apical depressions of all terga shallow, densely micropunctate, nearly as densely as on tergal disk. Terga I-III densely and distinctly micropunctate, punctures ill defined, interspaces microsculptured, slightly shiny; terga IV-VII more coarsely and more densely punctate than previous terga. Sterna uniformly punctate nearly like terga.
Coloration. Apical part of mandibles red; terga I and II dark red; apex of sternum I and base of sternum II dark red in some specimens; distal tarsomeres dark red on ventral side; forefemoral notch dark red in most specimens, but black in some. Tegulae brown translucent. Apical parts of terga slightly translucent. Remaining body parts all black.
Czech Republic, France, Germany, Hungary, Italy, Kyrgyzstan, Poland, Slovakie and Turkey.
Body length: 8.9 mm.
Head. Clypeus slightly convex, top at clypeal midlength; basomedian area cover majority of space of clypeus, as densely punctate as lateral section; bevel very small, slightly convex to flat, only about 2–3 × as long as clypeal lip, slightly longer medially than laterally, with few large punctures, ending far from base of clypeus, shiny; lip insignificanly arcuate to nearly straight, with faint lateral incision, separate from bevel by punctate groove,
Mesosoma. Scutum without distinct anterior impression; scutum and scutellum densely punctate, punctures well defined, less than half diameter apart, interspaces well developed and relatively uniform in size, microsculptured, slightly shiny, setae about 1.0 ×
Metasoma. Terga I-III with distinct silvery, apical faciae; terga I-II very densely and finely micropunctate, punctures ill-defined but distinct, interspaces microsculptured, slightly shiny; apical depressions well developed, with distinct micropunctures; sculpture of terga III-V distinctly coarser than on previous terga, also more densely punctate; tergum V coarsely punctate, punctures ill-defined, but large, half to one diameter apart, interspaces microsculptured, apical depression of tergum V very sparsely, finely punctate. Pygidium of usual size, sparsely punctate, punctures large, ill defined, interspaces microsculptured basally, but unsculptured in apical half, slightly shiny to shiny. Central part of sternum II with several large punctures, interspaces microsculptured, shiny; lateral part slightly shiny, densely micropunctate; remaining sterna with uniform sculpture similar to that on sternum II, but more or less reduced laterally.
Coloration. Central part of mandibles, terga and sterna I-III red. Distal tarsomeres are dark red. Tegulae brown translucent. Apical parts of terga I-III slightly translucent. Remaining body parts all black.
Variation of females: Body length: 7.2–9.0 mm. Antenna relatively short,
Body length: 5.4–6.8 mm.
Head. Mandibular inner margin with one distinct rectangular tooth and distinct furrow next to tooth distally. Clypeus very slightly convex (appearing nearly flat), uniformly curved, top at about clypeal midlength; basomedian area dominant on clypeus, densely, uniformly punctate; bevel nearly or completely absent, reduced to shiny area ventromedially; lip arcuate or slightly sinuate, with small rectangular lateral corner, lip separated from bevel by distinct, punctate groove;
Mesosoma. Scutum without distinct anterior impression; scutum and scutellum densely punctate, punctures well defined, most punctures less than half diameter apart, interspaces distinct, microsculptured, slightly shiny to dull, setae about 1.0 ×
Metasoma. Terga I-III with silvery apical faciae. Apical depressions of terga distinct, micropunctate similarly to that on tergal disk. Terga I-III densely, distinctly micropunctate, punctures ill defined, interspaces microsculptured, slightly shiny; sculpture of terga IV-VII slightly coarser than on previous terga, also punctures slightly denser. Sterna uniformly punctate, nearly like terga.
Coloration. Apical part of mandible, terga and sterna I-III and forefemoral notch red. Distal tarsomeres reddish. Tegulae brown to reddish. Apical parts of all terga and tegulae translucent. Remaining body parts all black.
Bulgaria, Czech Republic, Greece, Hungary, Poland and Turkey.
The species is named after its uniform sculptures. The species appears as a “noble” among other species of the
China, Iran, Kazakhstan, Kyrgyzstan, Tajikistan, Turkey, and Uzbekistan. It is probably also distributed in Jordan, Lebanon and Syria.
Holotype came from China (Uighur region). Type material not examined. The species was redescribed by
Body length: 5.3–7.8 mm.
Head. Mandibular inner margin with one rectangular tooth and distinct furrow next to tooth distally. Clypeus convex, slightly elevated and steeply declining ventrally, top in apical third or in midlength; basomedian area large, sparsely punctate, more sparsely punctate medially than laterally in some specimens; bevel convex, conspicuously decreasing toward clypeal lip, variable in size, shiny with several larger punctures; lip distinctly sinuate, conspicuously produced medially in most specimens, with distinct lateral corner, lip separated from bevel by variable groove;
Mesosoma. Scutum without distinct anterior impression; scutum and scutellum densely punctate, punctures well defined, most punctures less than half diameter apart, interspaces distinct, microsculptured, slightly shiny to dull, setae about 1.0 ×
Metasoma. Silvery apical faciae of terga I-III faintly developed, often absent. Apical depressions of all terga shallow, densely micropunctate, nearly as densely as more anterior parts. Terga I-III densely and distinctly micropunctate, punctures ill defined, interspaces microsculptured, slightly shiny; sculpture of terga IV-VII coarser than on previous terga, also punctation denser. Sterna uniformly punctate nearly like terga.
Coloration. Apical part of mandible red; terga I and II dark red, base of tergum II dark red in some specimens; apex of sternum I and base of sternum II dark red, infrequently specimens with three red metasomal segments also occur; distal tarsomeres dark red on ventral side; forefemoral notch black in most specimens, but dark red in a few. Tegulae reddish or brown translucent. Apical parts of all terga slightly translucent. Remaining body parts all black.
Body length: 7.8 mm.
Head. Clypeus distinctly convex, top at clypeal midlength; basomedian area relatively large, more sparsely punctate and punctures larger than on lateral section, interspaces between punctures basolaterally small, but getting sparser in mesoventral direction, interspaces well developed, but punctures mostly less than one diameter apart; bevel large, distinctly convex, as long as basomedian area both laterally and medially, not triangular, with sparse large punctures, not reaching base of clypeus, shiny; lip slightly arcuate, with lateral incision, separate from bevel by punctate groove,
Mesosoma. Scutum without distinct anterior impression; scutum and scutellum densely punctate, punctures well defined, less than half to one diameter apart, microsculptured, slightly shiny, setae about 1.0 ×
Metasoma. Terga I-III with silvery apical fasciae inconspicuously developed. Apical depressions of all terga shallow, with distinct micropunctures, on terga I-II evanescent in microsculpture. Terga I-III densely, finely micropunctate, punctures ill defined but distinct, interspaces microsculptured, slightly shiny; sculpture of tergum III-V distinctly coarser than on previous terga, also punctures slightly denser. Pygidium sparsely punctate, punctures ill defined, interspaces microsculptured, dull. Central part of sternum II with several large punctures, interspaces microsculptured, shiny; lateral part slightly shiny, densely micropunctate; remaining sterna with uniform sculpture similar to that of sternum II, but more or less reduced laterally.
Coloration. Central part of mandible, three distal tarsomeres, terga and sterna I-III red. Tegulae brown translucent. Apical parts of terga I-III slightly translucent. Remaining body parts all black.
Variation of females: Body length: 6.7–9.3 mm.
Austria, Czech Republic, Bulgaria, Germany, France, Greece, Hungary, Italy, Kyrgyzstan, Portugal, Slovakia, Switzerland and Turkey.
Body length: 6.4–9.2 mm.
Head. Clypeus distinctly convex, top at clypeal midlength; basomedian area relatively large, about two fifths of bevel in space, as densely, uniformly punctate as lateral section; bevel large, slightly convex, obtusely triangular, with sparse large punctures, reaching base of clypeus in some specimens, but distantly separated from it in others, shiny; lip slightly sinuate with small median emargination and with small, but well developed lateral incision with rectangular corner, lip separate from bevel by punctate groove,
Mesosoma. Scutum without distinct anterior impression; scutum and scutellum densely punctate, punctures well defined, half to one diameter apart, interspaces slightly microsculptured, shiny, setae about 1.0 ×
Metasoma. Terga I-III with silvery apical fasciae. Apical depressions of terga shallow, with micropunctures evanescent in microsculpture. Terga I-III sparsely, finely micropunctate, punctures ill defined, interspaces microsculptured, shiny to slightly shiny; sculpture of tergum IV-V slightly coarser than on previous terga, punctures slightly denser. Pygidium sparsely punctate, punctures ill defined, interspaces microsculptured, slightly shiny. Central part of sternum II with several large punctures, interspaces microsculptured, shiny; lateral part slightly shiny, densely micropunctate; remaining sterna with uniform sculpture similar to that of sternum II, but more or less reduced laterally.
Coloration. Central part of mandibles, three distal tarsomeres, terga and sterna I-III red. Tegulae brown translucent. Apical parts of terga I-III slightly translucent. Remaining body parts all black.
Body length: 6.7 mm.
Head. Clypeus slightly convex, uniformly curved, top at clypeal midlength; basomedian area large, densely punctate; bevel small, slightly convex, shiny, with several large punctures; lip conspicuously arcuate, well developed medially, with small lateral corner, lip separated from bevel by distinct groove with large punctures;
Mesosoma. Scutum without distinct anterior impression; scutum and scutellum densely, relatively finely punctate, punctures well defined, most punctures less than one diameter apart, interspaces distinct, unsculptured, shiny, setae about 1.0 ×
Metasoma. Terga I-III with silvery apical faciae. Apical depressions of terga shallow, micropunctate. Terga I-III sparsely, distinctly micropunctate, punctures ill defined, interspaces microsculptured, shiny; sculpture of tergum IV-VII coarser than on previous terga, punctures slightly denser. Sterna uniformly punctate nearly like terga.
Coloration. Apical part of mandibles, tegulae, terga and sterna I-III red; distal tarsomeres and forefemoral notch in center dark reddish. Apical parts of terga and tegula translucent. Remaining body parts all black.
Variation of males: Mandibular inner margin with one distinct rectangular tooth and distinct furrow next to the tooth distally. Antenna relatively short,
Austria, Bulgaria, Germany, Hungary, Italy, Slovenia and Turkey.
The species is named after its punctate mesopleuron in most males.
♀♀
1a | Gena dorsally inconspicuous, converging behind compound eyes (Figs |
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1b | Gena dorsally more robust (Figs |
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2a | Scutum and scutellum sparsely punctate, with large interspaces, punctures well developed, usually up to 1.5 diameter apart (more so in some specimens) (Fig. |
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2b | Scutum and scutellum densely punctate, with small but distinct interspaces, punctures usually less than one diameter apart (only few interspaces larger in some specimens) (Figs cf. 39, 40). Ventral part of mesopleuron densely punctate, punctures small, ill defined (Fig. cf. 51) |
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3a | Wing membrane light yellowish (Fig. cf. 105). Clypeus only slightly convex, bevel small, basomedian area densely punctate, as long as 2/3 of clypeal length (Fig. |
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3b | Wing membrane infumate (Fig. |
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3c | Character combination different |
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4a | Vertex distinctly wider than long (Fig. |
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4b | Vertex about as long as wide or slightly wider than long (Fig. |
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5a | Pygidium, mid- and hindtibiae light reddish (Fig. |
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5b | Pygidium and all tibiae dark |
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6a | Clypeus conspicuously elevated, transition between basomedian area and bevel relatively sharp, appearing angulated (Figs |
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6b | Clypeus less elevated, basomedian area sharply separated from bevel in sculpture, but gradually changing into one anoher in convexity (Figs |
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7a | Several punctures on scutum and trochanters about two diameters apart |
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7b | Punctures on scutum and trochanters less than one diameter apart |
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8a | Mesopleuron finely punctate, punctures less distinct, evanescent in microsculpture (Fig. |
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8b | Mesopleuron coarsely punctate, punctures relatively large and well developed ventrally (Fig. |
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9a | Clypeal bevel distinctly triangular, reaching clypeal base at middle and separating basomedian area in two parts; when clypeal base punctate, punctures and interspaces larger than on lateral side of basomedian area (Figs |
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9b | Clypeal bevel distinctly separated from clypeal base by basomedian area, which is continuous from side to side, uniformly densely punctate, punctures much finer than on bevel (Figs |
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10a | Clypeus markedly convex, most elevated point located between clypeal midlength and basal third of clypeus (Fig. |
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10b | Clypeus less convex, most elevated point not well defined, but appearing located near clypeal midlength (Fig. |
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11a | Punctures of central part of terga IVand V small, ill defined; interspaces between punctures large, distinct, slightly shiny to shiny (Figs cf. 80, 83) |
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11b | Punctures of central part of terga IVand V larger, well defined, approximately 0.5–1.5 diameter apart, slightly shiny to dull (Figs |
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12 | Similar and variable species, difficult to distinguish: | |
12a | Free margin of clypeus in fresh specimens sinuate, with small emargination medially (Fig. |
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12b | Free margin of clypeus in fresh specimens arcuate or straight, without emargination medially (Fig. |
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13a | All mesopleuron coarsely sculptured (Figs cf. 44, cf. 46, cf. 48). Hypoepimeral area of mesopleuron, and propodeal dorsum longitudinally ridged (propodeal dorsum sometimes coarsely sculptured, with indistinct ridges) (Figs cf. 46, 58. Tergum V, including apical depression, densely, distinctly punctate (Fig. |
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13b | Mesopleuron, including hypoepimeral area, finely sculptured (Fig. |
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14a | All mesopleuron coarsely sculptured (Figs cf. 44, cf. 46, cf. 48). Hypoepimeral area and propodeal dorsum and side longitudinally ridged (propodeal dorsum sometimes coarsely sculptured, but ridges indistinct) (Figs cf. 46, 58. Tergum V, including apical depression, densely and distinctly punctate (Fig. |
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14b | Mesopleuron, including hypoepimeral area, punctate or finely sculptured (Fig. |
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15a | Supraantennal tubercle rounded, distinctly punctate on top, in between and nearly at all sides; small impunctate area occurs only laterally (Fig. |
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15b | Supraantennal tubercle prominent with relatively sharply delimited, shiny, nearly impunctate area laterally and at top (Figs |
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16a | Apex of terga IV and V finely microsculptured, sparsely punctate to impunctate, contrasting with coarsely punctate central part of tergum IV, punctures of terga IV and V well developed (Fig. |
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16b | Apex of tergum IV coarsely microsculptured, sparsely punctate (Fig. |
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17 | Similar and variable species, difficult to distinguish: | |
17a | Clypeal free margin in fresh specimens sinuate, with small anterior emargination medially (Fig. |
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17b | Clypeal free margin in fresh specimens arcuate or straight, without anterior emargination medially (Fig. |
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♂♂
1a | Surface of forefemoral notch slightly shiny to shiny, without elevated plate and ledge on margins and with distinct small setae (Fig. |
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1b | Surface of forefemoral notch microsculptured (finely so in some species), slightly elevated, with distinct ledge on inner margin (also on outer margin in most species), without distinct setae (Figs |
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2a | Central part of scutum sparsely punctate, punctures more than one diameter apart (Fig. cf. 38) |
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2b | Scutum densely punctate, puncture less than one diameter apart (Fig. cf. 39) |
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3a | Gena narrow (Figs cf. 25, cf. 28). Trochanteral venters with large, glabrous spaces between punctures. Ventral and lower part of mesopleuron coarsely punctate, punctures well defined. Vertex, propodeum and metasoma finely sculptured |
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3b | Gena robust (Fig. cf. 27). Trochanters densely punctate. Ventral and lower part of mesopleuron coarsely punctate (Fig. cf. 43), punctures well defined. Vertex (Fig. cf. 37), propodeum (Fig. cf. 55) and metasoma coarsely sculptured |
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3c | Gena robust (Fig. cf. 27). Trochanters densely punctate. Ventral and lower part of mesopleuron finely and densely punctate, punctures ill defined (Fig. cf. 42). Vertex (Fig. |
Tentative position of unknown male of |
4a | Gena dorsally short, conspicuously converging behind compound eyes (Figs |
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4b | Gena behind compound eyes more robust (Figs |
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5a | All tibia and tergum VII reddish (Fig. cf. 105) |
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5b | All tibia and tergum VII dark |
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6a | All clypeus densely punctate, clypeal bevel indistinct (Fig. |
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6b | Clypeal bevel distinct, sparsely punctate (Figs |
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7a | Forefemoral notch indistinctly microsculptured, shiny (Fig. |
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7b | Forefemoral notch densely microsculptured, dull (Fig. |
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8a | Supraantennal tubercle rounded, distinctly punctate on top, in-between and nearly at all sides; small impunctate area only laterally (Fig. |
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8b | Supraantennal tubercle prominent, relatively sharply delimited, shiny, nearly impunctate area dorsally and laterally (Figs |
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9a | Mesopleuron coarsely irregularly ridged (Figs |
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9b | Mesopleuron finely sculptured (Figs cf. 45, cf. 47). Forefemoral notch light red (Figs |
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10a | Body black, silvery apical fasciae present on terga I-IV (Fig. |
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10b | At least basal metasomal terga red, silvery apical fasciae present on terga I-III |
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11a | Clypeus coarsely punctate, interspaces between punctures large, punctures about one diameter apart (Fig. |
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11b | Clypeus finely and densely punctate, interspaces between punctures small, punctures less than one diameter apart (Fig. |
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12a | Mesopleuron laterally and ventrally with well-defined punctures (punctures distinct also on hypoepimeral area in some specimens), interspaces distinct, light shiny to shiny on most parts (Fig. |
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12b | Mesopleuron laterally and ventrally with ill-defined punctures, when interspaces distinct, then microsculptured, slightly shiny to dull. Forefemoral notch larger and deeper, distinctly carinated on anterior and posterior margin of notch (Figs cf. 63, 64–65). Clypeal lip of usual form, slightly arcuate (Fig. cf. 9) |
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13a | Supraclypeal area forming groove, supraantennal tubercle distinctly elevated ventromedially (Fig. cf. 108). Forefemoral notch light red, large, its diameter usually longer than distance that separates it from forefemoral base (Fig. |
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13b | Supraclypeal area flat, supraantennal tubercle slightly elevated ventromedially (Fig. cf. 107). Forefemoral notch variable, usually smaller, light red to red (Fig. |
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Morphological variation in
Head of male, front view, and mandibles.
Head of female, front view.
Head of female, lateral view of clypeus.
Gena of male.
Gena of female.
Vertex of female, detail.
Scutum of female, dorsal view.
Mesopleuron, lateral view, sculptures.
Mesopleuron, ventral view, sculptures, female.
Propodeum, dorsal and posterior view, female.
Forefemoral notch of male, ventral and lateral view.
Tergum V, sculpture, female.
Pygidium, female.
Volsella, lateral view.
Habitus, dorsal and lateral view.
Supraantennal tubercles, frontal view.
I would like to thank all of the people who provided material for this study, namely Werner Arens, Yulia Astafurova, Petr Bogusch, Sandor Csősz, Fritz Gusenleitner, Jiri Halada, Marek Halada, Niklas Johansson, Zdeněk Karas, Toshko Ljubomirov, Jan Macek David Notton, Piotr Olszewski, Martin Říha, Wolfgang Schlaefle, Christian Schmid-Egger, Stefan Schmidt, Pavel Tyrner, Manuela Vizek and Waldemar Żyła. I also thank W. J. Pulawski for his comments to the manuscript.