Corresponding author: Hannes Baur (
Academic editor: Michael Engel
Two new species,
Baur H (2015) Pushing the limits – two new species of
In Western Europe the vast majority of newly discovered insect species usually belong to complexes of cryptic species. Morphologically, such new species are therefore quite similar to known species and in many cases even rather difficult to separate from those (e.g.,
The genus
Specimens are deposited in the following collections (acronyms mostly according to
Geographical coordinates on data labels of type specimens are indicated as WGS 84 latitude and longitude.
Nomenclature and classification of
Abbreviation, name, definition, and magnification of the 41 measurements used in the description (see Material and methods for further information).
Abbreviation | Character name | Definition | Magnification in pixel/mm* |
---|---|---|---|
ant.l | Pedicel plus flagellum length | Combined length of pedicel plus flagellum, outer aspect ( |
1742 |
clv.b | Clava breadth | Greatest breadth of clava, outer aspect | 3910 |
clv.l | Clava length | Greatest length of clava, outer aspect | 3910 |
eye.b | Eye breadth | Greatest breadth of eye, lateral view | 3910 |
eye.d | Eye distance | Shortest distance between eyes, dorsal view | 1742 |
eye.h | Eye height | Greatest length of eye height, lateral view | 2549 |
eye.l | Eye length | Length of eye, dorsal view ( |
888 |
fl3.b | First funicular segment breadth | Greatest breadth of first funicular segment (= third flagellar segment), outer aspect | 3910 |
fl3.l | First funicular segment length | Greatest length of first funicular segment (= third flagellar segment), outer aspect | 3910 |
fl8.b | Sixth funicular segment breadth | Greatest breadth of sixth funicular segment (= eighth flagellar segment), outer aspect | 3910 |
fl8.l | Sixth funicular segment length | Greatest length of sixth funicular segment (= eighth flagellar segment), outer aspect | 3910 |
fm3.b | Metafemur breadth | Greatest breadth of metafemur, outer aspect | 3910 |
fm3.l | Metafemur length | Length of metafemur, from distal end of trochanter to tip of metafemur, measured along midline, outer aspect | 1742 |
fwi.b | Fore wing breadth | Greatest breadth of fore wing, measured at about right angle to marginal and postmarginal veins | 1742/1089 |
fwi.l | Fore wing length | Greatest length of fore wing, measured from end of humeral plate to tip of wing | 1089 |
gst.b | Gaster breadth | Greatest breadth of gaster, distance between the outermost lateral edges of the gaster, dorsal view | 1742/2549 |
gst.l | Gaster length | Length of gaster along median line from posterior edge of nucha to tip of ovipositor sheath, dorsal view | 1089 |
hea.b | Head breadth | Greatest breadth of head, dorsal view | 1742 |
hea.h | Head height | Distance between anterior margin of clypeus and anterior edge of anterior ocellus, frontal view | 1742 |
hea.l | Head length | Length of head, dorsal view ( |
888 |
lof.h | Lower face height | Distance between anterior margin of clypeus and lower margin of torulus | 2549 |
mav.l | Marginal vein length | Length of marginal vein, distance between the point at which the submarginal vein touches the leading edge of the wing and the point at which stigmal vein and postmarginal vein unite ( |
2549 |
msc.b | Mesoscutum breadth | Greatest breadth of mesoscutum just in front of level of tegula, dorsal view | 1742 |
msc.l | Mesoscutum length | Length of mesoscutum along median line from posterior edge of pronotum to posterior edge of mesoscutum, dorsal view | 2549 |
msp.l | Malar space | Distance between the point where malar sulcus enters mouth margin and malar sulcus enters lower edge of eye, lateral view ( |
3910 |
mss.l | Mesosoma length | Length of mesosoma along median line from anterior edge of pronotum collar to posterior edge of nucha, dorsal view | 1089 |
ool.l | OOL | Shortest distance between posterior ocellus and eye margin, dorsal view ( |
3910 |
pdl.b | Pedicel breadth | Greatest breadth of pedicel, outer aspect | 3910 |
pdl.l | Pedicel length | Length of pedicel, outer aspect | 3910 |
plc.d | Plica distance | Greatest distance between upper edge of anterior plica | 2549 |
pmv.l | Postmarginal vein length | Length of postmarginal vein ( |
2549 |
pol.l | POL | Shortest distance between posterior ocelli, dorsal view ( |
3910 |
ppd.l | Propodeum length | Length of propodeum measured along median line from anterior edge to posterior edge of nucha, dorsal view | 2549 |
scp.b | Scape breadth | Greatest breadth of scape, outer aspect | 3910 |
scp.l | Scape length | Length of scape exclusive of radicle, outer aspect ( |
2549 |
sct.l | Scutellum length | Length of scutellum along median line from posterior edge of mesoscutum to posterior edge of scutellum, dorsal view | 2549 |
stv.l | Stigmal vein length | Length of stigmal vein, distance between the point at which stigmal vein and postmarginal vein unite apically, and the distal end of the stigma ( |
2549 |
ta3.l | Metatarsus length | Length of metatarsus, including pretarsus | 2549 |
tb3.b | Metatibia breadth | Apical breadth of metatibia, outer aspect | 3910 |
tb3.l | Metatibia length | Length of metatibia, measured along midline, outer aspect | 1742 |
tmp.l | Temple length | Length of temple, dorsal view ( |
888 |
Where more than one number is present, the first refers to the magnification used for
Most characters were measured on photographs taken by Lisa Wilmsmeier with a Leica DFC425 camera mounted on a Leica M16 stereomicroscope. Photographs were taken at different magnifications depending on the size of the character, in order to reduce measurement error for the smaller ones. For all measurements, it was ensured that the points of reference were in perfect focus and that the diaphragm of the lens was fully open. The distances were finally measured using ImageJ, version 1.46v (
I measured three characters, eye length, head length, and temple length, on a single stack photograph taken with a Keyence VHX 2000 digital photomicroscope and a VH-Z20R/W zoom lens at a magnification of 200× (i.e., 1000 µm corresponded to 888 pixels, see Table
I also used the Keyence microscope for making stack-images of qualitative character states. A 4-digit individual code including the notion “Baur” (e.g., “Baur 2410”) was provided for specimens that have been measured or photographed, or used as reference specimens for comparison with newly described species.
I compiled all morphological data in a FileMaker Pro 12®, version 12.0v5, database, of which natural language descriptions as well as ranges of body ratios were generated using the FileMaker script language. Because this is commercial software, a qualitative and a quantitative data matrix (raw values in µm) were exported as comma separated values (CSV) files made available at the BMNH data portal at DOI:
Holotype ♀ Switzerland, Canton Bern, Köniz, Niederwangen, 570 m,
Scape: testaceous; pedicel: testaceous, slightly infuscate dorsally; flagellum: brown.
Fore wing: hyaline; fore wing venation: testaceous; setae on fore wing: fuscous; hind wing: hyaline.
Coxae: green; trochanters: testaceous; femora: testaceous; tibiae: testaceous; tarsi: testaceous with fifth segment slightly infuscate; pretarsi: slightly infuscate.
Petiole: green with purplish tinge; gaster: green; gastral terga: one to five with strong purplish tinge.
Mesoscutum: finely reticulate, meshes rather high, areoles small and only moderately enlarged in posterior part of sclerite (Fig.
Pro- and mesocoxa: finely alutaceous, metacoxa: finely reticulate.
Median area of propodeum: evenly reticulate, as strong as on mesoscutum (Fig.
Petiole in dorsal view: smooth; gastral terga: smooth and shining, sixth tergum and syntergum alutaceous (Fig.
Antenna (Fig.
Mesosoma in lateral view: moderately strongly bent; propodeum in lateral view sloping with respect to dorsal plane of mesoscutum and scutellum at an angle of: 45 degrees; pronotum breadth with respect to mesoscutum breadth: distinctly narrower; pronotum collar: horizontal, well defined, its length with respect to mesoscutum length: one sixth, its anterior margin: rounded edge; pronotum posterior margin: thin, shiny strip; notaulus: extremely superficial, hardly traceable, reaching: about half along mesoscutum (Fig.
Fore wing (Fig.
Femora: moderately slender; metatibia: quite abruptly expanded before the middle (Fig.
Petiole in dorsal view: conical, in ventral view: open; gaster in dorsal view: ovate, obtusely pointed (Fig.
Head breadth to height: 1.2–1.41; head breadth to length: 2.02–2.08; head breadth to mesoscutum breadth: 1.26–1.34; lower face height to head height: 0.5–0.58; POL to OOL: 0.76–0.87; eye height to breadth: 1.3–1.36; eye distance to height: 1.74–1.88; temple length to eye length: 0.35–0.44; malar space to eye height: 0.68–0.76.
Pedicel plus flagellum length to head breadth: 0.72–0.87; scape length to eye height: 0.99–1.04; scape length to breadth: 5.24–5.82; pedicel length to breadth: 1.22–1.54; pedicel length to first funicular segment length: 0.84–1.13; first funicular segment length to breadth: 0.91–1.33; sixth funicular segment length to breadth: 0.85–1.04; first funicular segment breadth to clava breadth: 0.85–1.06; clava length to breadth: 2.01–2.57.
Mesosoma length to mesoscutum breadth: 1.5–1.6; mesoscutum breadth to length: 1.57–1.76; mesoscutum length to scutellum length: 1.03–1.1; propodeum length to scutellum length: 0.57–0.62; plica distance to propodeum length: 1.21–1.31.
Fore wing length to breadth: 2–2.18; marginal vein length to stigmal vein length: 1.51–1.68; postmarginal vein length to stigmal vein length: 0.78–1.01.
Metafemur length to breadth: 3.27–4.47; metatibia length to breadth: 5.61–7.82; metatarsus length to metatibia length: 0.65–0.89.
Gaster length to breadth: 1.17–1.62; gaster length to mesosoma length: 0.82–1.11.
Scape: testaceous; pedicel: testaceous, slightly infuscate dorsally; flagellum: testaceous, slightly infuscate dorsally.
Fore wing: hyaline; fore wing venation: testaceous; setae on fore wing: fuscous; hind wing: hyaline.
Coxae: green; trochanters: testaceous; femora: testaceous; tibiae: testaceous; tarsi: testaceous with fifth segment slightly infuscate; pretarsi: slightly infuscate.
Petiole: green with purplish tinge; gaster: green; gastral terga: one to three with an indistinct yellowish spot.
Mesoscutum: finely reticulate, meshes rather high, areoles small and only moderately enlarged in posterior part of sclerite; scutellum: reticulate, meshes about as strong and coarse as on posterior part of mesoscutum, but with a narrow band of smaller areoles in anterior half of median longitudinal line; frenum: reticulate, meshes of similar size to those on scutellum; axilla: reticulate, about as strong as on central part of scutellum; prepectus upper triangular area: reticulate; upper mesepimeron: anteriorly smooth, posterior corner distinctly alutaceous; upper mesepisternum: reticulate, about as strong as on mesoscutum; metapleuron: reticulate, about as strong as on mesepisternum.
Pro- and mesocoxa: finely alutaceous, metacoxa: finely reticulate.
Median area of propodeum: evenly reticulate, as strong as on mesoscutum; inner corner of anterior plica: with a depression, weakly reticulate; nucha: reticulate, as strong as on median area of propodeum; callus of propodeum: reticulate; paraspiracular sulcus: reticulate with few transverse costulae.
Petiole in dorsal view: smooth; gastral terga: smooth and shining, sixth tergum and syntergum alutaceous.
Antenna (Fig.
Mesosoma in lateral view: moderately strongly bent; propodeum in lateral view sloping with respect to dorsal plane of mesoscutum and scutellum at an angle of: 50 degrees; pronotum breadth with respect to mesoscutum breadth: distinctly narrower; pronotum collar: horizontal, well defined, its length with respect to mesoscutum length: one sixth, its anterior margin: rounded edge; pronotum posterior margin: thin, shiny strip; notaulus: extremely superficial, hardly traceable, reaching: about half along mesoscutum; scutellum in lateral view: moderately convex; scutellum in posterior view: moderately convex; scutellum posterior margin projection: level of anterior margin of dorsellum; scutellum posterior margin in posterior view: narrowly emarginate in the middle; frenal line: finely indicated, especially on sides; prepectus upper triangular area: separated by a fine oblique carina; upper mesepimeron: strongly narrowing below, not reaching base of mesopleuron; anterior plica: bent inwards in anterior two fifths and strong; posterior plica: present, joining anterior plica; orientation of posterior plicae: almost parallel; median carina of propodeum: weakly indicated, irregular; nucha: elevated but not clearly differentiated from median area of propodeum; spiracle: oval, size: small, separated from anterior margin of propodeum by: shortest diameter; callus pilosity: numerous long setae; paraspiracular sulcus: narrow and deep.
Fore wing apex with respect to apex of gaster when folded back: distinctly exceeding; basal cell number of setae: 6; basal setal line: complete, with: 8 setae; cubital setal line: incomplete, with: 4 setae; costal cell pilosity on dorsal side: bare; costal cell pilosity on lower side: numerous setae in distal half and a complete setal line extending to base; speculum on upper side: bare, widely open below; fore wing disc: rather thickly pilose; marginal setae: present, short; stigma: subcircular, small; uncus: short.
Femora: moderately slender; metatibia: quite abruptly expanded before the middle; metacoxa pilosity, dorsally: bare.
Petiole in dorsal view: conical, in ventral view: open; gaster in dorsal view: ovate; gastral terga: weakly sunken.
Head breadth to height: 1.46–1.47; head breadth to length: 2.02–2.03; head breadth to mesoscutum breadth: 1.3; lower face height to head height: 0.59–0.6; POL to OOL: 0.89–0.96; eye height to breadth: 1.29–1.3; eye distance to height: 1.78; temple length to eye length: 0.39–0.43; malar space to eye height: 0.58–0.61.
Pedicel plus flagellum length to head breadth: 0.84; scape length to eye height: 0.97–0.98; scape length to breadth: 4.89–5.15; pedicel length to breadth: 1.37; pedicel length to first funicular segment length: 0.85–0.96; first funicular segment length to breadth: 1.27–1.57; sixth funicular segment length to breadth: 1.02–1.05; first funicular segment breadth to clava breadth: 0.91–0.98; clava length to breadth: 2.44–3.13.
Mesosoma length to mesoscutum breadth: 1.63–1.64; mesoscutum breadth to length: 1.48–1.5; mesoscutum length to scutellum length: 1.08–1.12; propodeum length to scutellum length: 0.55–0.59; plica distance to propodeum length: 1.21–1.39.
Fore wing length to breadth: 2–2.02; marginal vein length to stigmal vein length: 1.39–1.56; postmarginal vein length to stigmal vein length: 0.84–0.93.
Metafemur length to breadth: 4.23–4.62; metatibia length to breadth: 7.16–7.35; metatarsus length to metatibia length: 0.72–0.74.
Gaster length to breadth: 1.68–1.71; gaster length to mesosoma length: 1–1.01.
Close examination of the expanded metatibia under a stereomicroscope did not reveal any distinctive characteristics compared to the “normal”, i.e. unexpanded, metatibia of the other
The female of
Below the most important differences are given for those species with which
Following the suggestion of the collector of the new species, Jacqueline Grosjean,
Holotype ♀ Switzerland, Canton Wallis, Kippel, Zend, 2100 m,
Scape: fuscous with basal third testaceous; pedicel: fuscous; flagellum: fuscous.
Fore wing: hyaline; fore wing venation: brownish; setae on fore wing: fuscous; hind wing: hyaline.
Coxae: green; trochanters: slightly greenish, testaceous at tips; pro- and mesofemur: green, testaceous in apical quarter, metafemur: green, testaceous in apical sixth; protibia: testaceous, meso- and metatibia: testaceous, medially slightly infuscate; tarsi: testaceous, apical segments slightly infuscate; pretarsi: slightly infuscate.
Petiole: dark purplish; gaster: green to blue-green with metallic luster; gastral terga: one to five with strong purplish tinge.
Mesoscutum: finely reticulate, meshes moderately high, areoles small and not enlarged in posterior part of sclerite; scutellum: reticulate, meshes about as strong and coarse as on posterior part of mesoscutum; frenum: reticulate, meshes larger than those on scutellum; axilla: reticulate, about as strong as on central part of scutellum; prepectus upper triangular area: weakly reticulate; upper mesepimeron: anteriorly smooth, posterior corner distinctly alutaceous; upper mesepisternum: reticulate, about as strong than on mesoscutum; metapleuron: weakly reticulate, less strong as on mesepisternum.
Coxae: weakly reticulate.
Median area of propodeum: uniformly reticulate, as strong as on mesoscutum; inner corner of anterior plica: with a smooth depression and transverse carinae; nucha: reticulate, as strong as on median area of propodeum; callus of propodeum: weakly reticulate; paraspiracular sulcus: smooth with few transverse costulae.
Petiole in dorsal view: smooth; gastral terga: smooth and shining, third to fifth tergum anteriorly, sixth tergum and syntergum wholly alutaceous.
Antenna (Fig.
Mesosoma in lateral view: rather flattened; propodeum in lateral view sloping with respect to dorsal plane of mesoscutum and scutellum at an angle of: 20 degrees (Fig.
Fore wing (Fig.
Femora: slender; metatibia: gradually widening towards apex; metacoxa pilosity, dorsally: bare.
Petiole in dorsal view: conical, in ventral view: open; gaster in dorsal view: very elongate and acuminate; gastral terga: strongly convex; posterior margin of first gastral tergum: entire; first gastral tergum reaching: one fourth of gaster; tip of hypopygium reaching: almost three fifths of gaster (Fig.
Head breadth to height: 1.35–1.39; head breadth to length: 2.08–2.1; head breadth to mesoscutum breadth: 1.18; lower face height to head height: 0.42–0.44; POL to OOL: 1.19–1.2; eye height to breadth: 1.54–1.65; eye distance to height: 1.4–1.46; temple length to eye length: 0.36–0.45; malar space to eye height: 0.45–0.48.
Pedicel plus flagellum length to head breadth: 1.05–1.07; scape length to eye height: 0.81–0.86; scape length to breadth: 6.05–6.39; pedicel length to breadth: 1.51; pedicel length to first funicular segment length: 0.64–0.68; first funicular segment length to breadth: 1.77–1.83; sixth funicular segment length to breadth: 1.06–1.1; first funicular segment breadth to clava breadth: 0.8–0.84; clava length to breadth: 2.09–2.24.
Mesosoma length to mesoscutum breadth: 1.7–1.71; mesoscutum breadth to length: 1.44–1.52; mesoscutum length to scutellum length: 1.13–1.22; propodeum length to scutellum length: 0.59–0.61; plica distance to propodeum length: 1.2–1.36.
Fore wing length to breadth: 2.24–2.3; marginal vein length to stigmal vein length: 1.7–1.78; postmarginal vein length to stigmal vein length: 0.93–0.99.
Metafemur length to breadth: 3.88–4.47; metatibia length to breadth: 7.19–7.29; metatarsus length to metatibia length: 0.8.
Gaster length to breadth: 5.04–5.35; gaster length to mesosoma length: 1.51–1.52.
Scape: fuscous with basal two fifths testaceous; pedicel: fuscous; flagellum: brown.
Fore wing: hyaline; fore wing venation: brownish testaceous; setae on fore wing: fuscous; hind wing: hyaline.
Coxae: green; pro- and mesotrochanter: slightly infuscate, metatrochanter: fuscous; pro- and mesofemur: infuscate, testaceous in apical third, metafemur: green, testaceous on tips; tibiae: testaceous; protarsus: slightly infuscate, meso- and metatarsus: testaceous, apical segments slightly infuscate; pretarsi: slightly infuscate.
Petiole: dark purplish; gaster: green; gastral terga: basal terga with large dark yellow spot.
Mesoscutum: finely reticulate, meshes moderately high, areoles small and not enlarged in posterior part of sclerite; scutellum: weakly reticulate, meshes less strong and coarse than on posterior part of mesoscutum; frenum: weakly reticulate, meshes larger than those on scutellum; axilla: reticulate, about as strong as on lateral part of scutellum; prepectus upper triangular area: weakly reticulate; upper mesepimeron: anteriorly smooth, posterior corner distinctly alutaceous; upper mesepisternum: reticulate, about as strong as on mesoscutum; metapleuron: weakly reticulate, less strong than on mesepisternum.
Pro- and mesocoxa: finely alutaceous, metacoxa: finely reticulate.
Median area of propodeum: uniformly reticulate, as strong as on mesoscutum but with smaller meshes; inner corner of anterior plica: with a smooth depression and transverse carinae; nucha: reticulate, as strong as on median area of propodeum; callus of propodeum: weakly reticulate; paraspiracular sulcus: smooth with few transverse costulae.
Petiole in dorsal view: smooth; gastral terga: smooth and shining, second to sixth tergum and syntergum alutaceous.
Antenna (Fig.
Mesosoma in lateral view: rather flattened; propodeum in lateral view sloping with respect to dorsal plane of mesoscutum and scutellum at an angle of: about 25 degrees; pronotum breadth with respect to mesoscutum breadth: distinctly narrower; pronotum collar: horizontal, well defined, its length with respect to mesoscutum length: one sixth, its anterior margin: slightly elevated edge, medially carinate; pronotum posterior margin: thin, shiny strip; notaulus: superficial, reaching: two thirds along mesoscutum; scutellum in lateral view: almost flat; scutellum in posterior view: almost flat medially; scutellum posterior margin projection: level of anterior margin of dorsellum; scutellum posterior margin in posterior view: narrowly emarginate in the middle; frenal line: finely indicated, especially on sides; prepectus upper triangular area: ? (the lower part of the prepectus is concealed in the single male, but the character state is likely to be the same as for the females); upper mesepimeron: strongly narrowing below, not reaching base of mesopleuron; anterior plica: present, almost straight in anterior part; posterior plica: present, joining anterior plica; orientation of posterior plicae: almost parallel; median carina of propodeum: anteriorly indicated, effaced posteriorly; nucha: elevated but not clearly differentiated from median area of propodeum; spiracle: oval, size: small, separated from anterior margin of propodeum by: shortest diameter; callus pilosity: relatively sparsely setose; paraspiracular sulcus: narrow and deep.
Fore wing apex with respect to apex of gaster when folded back: not exceeding; basal cell number of setae: with up to 10 setae in distal part; basal setal line: complete, with: 11 setae; cubital setal line: incomplete, with: 4 setae; costal cell pilosity on dorsal side: bare; costal cell pilosity on lower side: with numerous setae in distal half and a complete setal line extending to base; speculum on upper side: bare, widely open below; fore wing disc: moderately thickly pilose; marginal setae: present, short; stigma: subrectangular, small; uncus: short.
Femora: slender; metatibia: gradually widening towards apex; metacoxa pilosity, dorsally: bare.
Petiole in dorsal view: conical, in ventral view: open; gaster in dorsal view: elongate, obtuse; gastral terga: weakly sunken; posterior margin of first gastral tergum: entire; first gastral tergum reaching: slightly less than one third of gaster.
Head breadth to height: 1.44; head breadth to length: 2.06; head breadth to mesoscutum breadth: 1.17; lower face height to head height: 0.51; POL to OOL: 1.33; eye height to breadth: 1.39; eye distance to height: 1.46; temple length to eye length: 0.38; malar space to eye height: 0.44.
Pedicel plus flagellum length to head breadth: 1.3; scape length to eye height: 0.84; scape length to breadth: 5.42; pedicel length to breadth: 1.28; pedicel length to first funicular segment length: 0.54; first funicular segment length to breadth: 2.08; sixth funicular segment length to breadth: 1.41; first funicular segment breadth to clava breadth: 0.97; clava length to breadth: 3.25.
Mesosoma length to mesoscutum breadth: 1.65; mesoscutum breadth to length: 1.44; mesoscutum length to scutellum length: 1.23; propodeum length to scutellum length: 0.64; plica distance to propodeum length: 1.1.
Fore wing length to breadth: 2.11; marginal vein length to stigmal vein length: 1.75; postmarginal vein length to stigmal vein length: 0.91.
Metafemur length to breadth: 4.3; metatibia length to breadth: 7.04; metatarsus length to metatibia length: 0.79.
Gaster length to breadth: 3.38; gaster length to mesosoma length: 1.19.
The dorsoventrally compressed mesosoma and the shape of the propodeum allowed an easy association of the females with the male even though they were collected in separate localities (about 160 km as the crow flies).
The female keys out in
Most similar are the following species but they differ – among many other characters mentioned in the description – by a rather more strongly bent mesosoma and a much less elongate female gaster:
Host unknown. The females of
Although the two new species are clearly placed within the genus
The expansion of the metatibia is a very rare phenomenon in parasitoid wasps.
The rearing of the host larva under protected condition suggests that
The other species,
I would specially like to thank Jacqueline Grosjean (JGC) who discovered the new species and presented the majority of specimens to the NMBE. I thank Rahel Schnidrig (Spiez, Switzerland) who provided the specimens of
Overview of 41 measurements (in µm) of
Character | MIN | MAX | MEAN | SD | MIN | MAX | MEAN | SD | MIN | MAX | MEAN | SD | VALUE |
---|---|---|---|---|---|---|---|---|---|---|---|---|---|
ant.l | 577 | 815 | 749.2 | 73.72 | 743 | 743 | 743.4 | 0.01 | 1033 | 1035 | 1033.9 | 0.93 | 1116 |
clv.b | 65 | 90 | 77.2 | 6.88 | 64 | 75 | 69.3 | 8.23 | 94 | 97 | 95.4 | 1.64 | 72 |
clv.l | 168 | 197 | 184 | 8.31 | 183 | 199 | 190.9 | 10.9 | 201 | 211 | 206.1 | 6.58 | 235 |
eye.b | 237 | 304 | 280.4 | 28.08 | 272 | 274 | 273.4 | 1.47 | 291 | 299 | 295.2 | 5.42 | 282 |
eye.d | 559 | 739 | 673.6 | 79.6 | 631 | 631 | 631.2 | 0.34 | 668 | 674 | 671.2 | 4.3 | 572 |
eye.h | 315 | 408 | 371.3 | 38.59 | 355 | 355 | 355 | 0.01 | 459 | 481 | 470 | 15.25 | 393 |
eye.l | 231 | 301 | 276 | 28.51 | 262 | 267 | 264.9 | 3.57 | 291 | 297 | 293.9 | 4.8 | 280 |
fl3.b | 68 | 84 | 74 | 4.86 | 62 | 69 | 65.4 | 4.82 | 77 | 79 | 78.4 | 1.5 | 70 |
fl3.l | 63 | 110 | 88.4 | 15.64 | 87 | 97 | 92.3 | 6.99 | 140 | 141 | 140.7 | 0.55 | 146 |
fl8.b | 65 | 83 | 75.5 | 5.85 | 64 | 65 | 64.7 | 1.03 | 81 | 85 | 83 | 3.15 | 71 |
fl8.l | 63 | 82 | 73.3 | 6.22 | 65 | 69 | 67 | 2.61 | 85 | 94 | 89.8 | 6.18 | 101 |
fm3.b | 115 | 155 | 138.6 | 17 | 128 | 137 | 132.6 | 5.85 | 151 | 175 | 163.1 | 17.35 | 134 |
fm3.l | 496 | 659 | 581.7 | 70.88 | 578 | 594 | 586.2 | 11.28 | 675 | 681 | 677.7 | 3.99 | 577 |
fwi.b | 794 | 1054 | 961 | 110.3 | 903 | 905 | 903.8 | 1.32 | 1118 | 1142 | 1130.1 | 17.32 | 928 |
fwi.l | 1721 | 2214 | 2002.6 | 208.5 | 1813 | 1825 | 1818.8 | 8.59 | 2564 | 2568 | 2565.9 | 2.79 | 1963 |
gst.b | 622 | 875 | 765.9 | 91.68 | 650 | 673 | 661.4 | 16.26 | 403 | 415 | 409.1 | 8.2 | 424 |
gst.l | 932 | 1111 | 1045.7 | 46.68 | 1113 | 1133 | 1123.1 | 14.28 | 2090 | 2157 | 2123.4 | 47.45 | 1435 |
hea.b | 789 | 1036 | 946.4 | 102.7 | 886 | 889 | 887.8 | 1.96 | 965 | 981 | 972.8 | 11.33 | 859 |
hea.h | 575 | 740 | 688.1 | 60.65 | 604 | 610 | 607.1 | 3.76 | 694 | 727 | 710.4 | 23.5 | 598 |
hea.l | 380 | 507 | 461.5 | 53.02 | 436 | 440 | 437.8 | 2.81 | 459 | 472 | 465.5 | 9.54 | 416 |
lof.h | 333 | 423 | 388.8 | 36.03 | 355 | 367 | 361.1 | 8.45 | 303 | 303 | 303 | 0.5 | 304 |
mav.l | 349 | 475 | 419.8 | 48.08 | 362 | 381 | 371.4 | 12.89 | 533 | 557 | 545 | 16.8 | 433 |
msc.b | 591 | 806 | 731 | 87.75 | 682 | 684 | 683.3 | 1.43 | 815 | 829 | 822 | 9.83 | 732 |
msc.l | 363 | 511 | 451.8 | 61.49 | 456 | 462 | 458.9 | 4.39 | 536 | 577 | 556.7 | 29.11 | 509 |
msp.l | 222 | 299 | 262.4 | 27.02 | 205 | 217 | 211.1 | 8.98 | 214 | 219 | 216.4 | 3.43 | 171 |
mss.l | 940 | 1279 | 1140.1 | 136.82 | 1118 | 1118 | 1118.3 | 0.32 | 1384 | 1419 | 1401.7 | 24.4 | 1208 |
ool.l | 170 | 236 | 211.3 | 25.8 | 187 | 189 | 187.9 | 1.93 | 185 | 188 | 186.3 | 2.03 | 153 |
pdl.b | 53 | 67 | 62.5 | 5.33 | 61 | 61 | 60.9 | 0.52 | 59 | 63 | 61.1 | 2.45 | 61 |
pdl.l | 69 | 97 | 86.5 | 10.33 | 83 | 84 | 83.4 | 0.99 | 90 | 95 | 92.2 | 3.68 | 79 |
plc.d | 256 | 358 | 314.9 | 40.58 | 299 | 318 | 308.5 | 13.17 | 345 | 380 | 362.6 | 25.06 | 294 |
pmv.l | 301 | 414 | 364.9 | 38.59 | 320 | 339 | 329.3 | 13.34 | 518 | 530 | 524.2 | 8.42 | 392 |
pol.l | 142 | 186 | 167.2 | 17.67 | 168 | 179 | 173.5 | 7.83 | 222 | 224 | 222.9 | 0.97 | 202 |
ppd.l | 202 | 286 | 251.1 | 30.68 | 228 | 248 | 238.3 | 14.05 | 279 | 288 | 283.4 | 6.24 | 267 |
scp.b | 59 | 78 | 69.2 | 6.93 | 67 | 71 | 69 | 3.07 | 61 | 65 | 62.8 | 3.12 | 61 |
scp.l | 320 | 412 | 375.9 | 37.81 | 344 | 348 | 346 | 2.7 | 387 | 393 | 390.2 | 4.27 | 329 |
sct.l | 346 | 463 | 421.5 | 51.33 | 413 | 422 | 417.5 | 6.71 | 474 | 475 | 474.3 | 0.55 | 414 |
stv.l | 215 | 293 | 263.8 | 32.12 | 244 | 260 | 251.9 | 11.59 | 314 | 314 | 313.9 | 0.41 | 248 |
ta3.l | 400 | 529 | 485.5 | 48.7 | 468 | 478 | 473.1 | 7.13 | 672 | 675 | 673.4 | 2.51 | 572 |
tb3.b | 78 | 110 | 98.5 | 10.45 | 88 | 91 | 89.6 | 2.31 | 115 | 117 | 116.1 | 0.94 | 102 |
tb3.l | 586 | 778 | 696.5 | 80.08 | 646 | 653 | 649.8 | 4.77 | 840 | 842 | 840.9 | 0.83 | 721 |
tmp.l | 84 | 130 | 108.8 | 18.49 | 103 | 113 | 107.8 | 6.77 | 107 | 130 | 118.6 | 15.93 | 107 |