Corresponding author: Dirk Ahrens (
Academic editor: A. Frolov
A review on the Chinese species of
Liu W-G, Fabrizi S, Bai M, Yang X-K, Ahrens D (2014) A taxonomic review on the species of
The genus
In this study, we examined the material collected in China mainland and deposited in Chinese institutional collections as well as various European and American collections. We found twenty-nine new taxa, which are described herein. Additionally, non-Chinese records are added to the species recorded from China, while the taxa occurring exclusively out of China are not revised herein. A key to the Chinese
The terminology and methods used for measurements, specimen dissection and genital preparation follow
Type specimens and other examined material are deposited in the following institutions or collections:
Body moderately large to large (6–12 mm), mostly dark brown; ventral surface reddish brown; dorsal surface dull and glabrous.
Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles, anterior margin weakly sinuate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate; frontoclypeal suture indistinctly incised, flat and weakly curved medially; ocular canthus short and triangular, impunctate, with a single terminal seta. Frons dull, with sparse, fine punctures, with single erect setae beside each eye. Antenna yellowish, with 10 antennomeres; club composed of 4 antennomeres in male, straight, rarely longer than 1.5 times as the remaining antennomeres combined; club in female composed of 3 antennomeres, as long as the remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly.
Pronotum moderately wide and strongly convex, lateral margins evenly convex, more strongly narrowed anteriorly towards sharp and slightly produced anterior angles. Anterior margin of pronotum slightly convex, with fine complete marginal line. Posterior angles blunt or strongly rounded. Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate.
Elytra oblong, widest just behind middle, striae distinctly impressed, finely and moderately densely punctate, intervals distinctly convex, with coarse and dense punctures concentrated along striae, with very minute setae in punctures; epipleural edge robust, ending at weakly curved and slightly blunt external apical angle of elytra, epipleura densely setose, apical border with a broad fringe of microtrichomes (100×).
Ventral surface weakly shiny, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae, metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Pygidium weakly convex and dull, densely punctate, without smooth midline, almost glabrous, but with a few longer setae along apical margin; pygidium without strong sexual dimorphism.
Legs moderately wide; femora finely and sparsely punctate; metafemur wide and moderately shiny or dull, anterior margin acute, posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, with a few short setae basally. Metatibia moderately wide to wide and moderately long, widest at half of metatibial length, dorsal margin sharply carinate, with two groups of spines; lateral face finely and sparsely punctate; ventral edge finely serrated, with four robust equidistant setae, medial face smooth, apex interiorly near tarsal articulation with a shallow sinuation. Tarsomeres with fine, very dense setae ventrally on distal half, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres with a strongly serrated ridge ventrally and glabrous; first metatarsomere slightly shorter than two following tarsomeres combined, one third of its length longer than dorsal tibial spine. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.
Aedoeagus. Phallobasis with a more or less long median ventral extension.
So far seven species from Himalaya and the type species from Myanmar have been formally assigned to
The genus is distributed almost in the entire Oriental region; we know described species so far assigned to “
1 | Labroclypeus completely glabrous. Basal group of dorsal spines of metatibia before middle. Ratio ocular diameter/interocular distance <0.75 |
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– | Labroclypeus with few fine setae. Basal group of dorsal spines of metatibia behind middle. Anterior margin of metafemur with continuously serrated adjacent line. Ratio ocular diameter/interocular distance >0.8 |
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2 | Metatibia more robust, ratio length/width: < 3.3. Metacoxa shorter, ratio length of metepisternum/metacoxa: 1/1.5 |
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– | Metatibia more slender, ratio length/width: >3.6. Metacoxa longer, ratio length of metepisternum/metacoxa: 1/1.68 |
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3 | Left paramere with long interior basal lobe (its length almost half of paramere length) |
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– | Left paramere with short interior basal lobe (its length less than quarter of paramere length) |
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4 | Posterior margin of metafemur straight or slightly convex |
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– | Posterior margin of metafemur with blunt tooth or sharp hook |
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5 | Posterior margin of metafemur with blunt tooth |
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– | Posterior margin of metafemur with sharp hook |
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6 | Left paramere long and narrow |
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– | Left paramere short and strout |
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7 | Dorsal lobe of right paramere short, not exceeding length of ventral one |
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– | Dorsal lobe of right paramere longer, exceeding length of ventral one |
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8 | Left paramere more narrow, dorsal margin weakly and evenly curved |
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– | Left paramere stout, dorsal margin bluntly angulate |
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9 | Eyes smaller, ratio diameter/interocular distance: 0.59. Dorsal lobe of right paramere large and directed distally, exceeding ventral lobe by far |
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– | Eyes larger, ratio diameter/interocular distance: 0.72. Dorsal lobe of right paramere very small and bent basally |
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10 | Ventral process of phallobasis short, distinctly shorter than phallobasis |
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– | Ventral process of phallobasis long, subequal to length of phallobasis |
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11 | Ventral process of phallobasis short, subequal to at maximum half of length of phallobasis |
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– | Ventral process of phallobasis medium in length, about three quarter of phallobasis length |
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12 | Metatibia more slender, ratio length/width ca. 3.4. Right paramere lacking basal lobe |
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– | Metatibia more robust, ratio length/width ca. 3.0. Right paramere with long filiform and curved basal lobe |
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13 | Eyes of medium size, ratio diameter/interocular distance ≥ 0.6 |
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– | Eyes small, ratio diameter/interocular distance ≤ 0.5 |
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14 | Phallobasis in dorsal view only slightly asymmetric. Left and right parameres simple, without two lobes. Posterior angles of pronotum strongly rounded |
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– | Phallobasis in dorsal view strongly asymmetric. Right paramere simple, left paramere with ventral lobe shorter than dorsal one. Posterior angles of pronotum moderately rounded |
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15 | Both parameres with dorsal and ventral lobe |
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– | One or both parameres simple, without two lobes |
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16 | Both parameres simple, without two lobes |
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– | One of parameres complex, with two lobes |
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17 | Right paramere simple, left one with two lobes; right paramere basiventrally strongly widened towards apex |
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– | Left paramere simple, right one with two lobes |
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18 | Left paramere more slender, strongly evenly bent externally |
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– | Left paramere more stout, almost straight |
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19 | Right paramere basally with brush of robust trichome-like spines |
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– | Right paramere without brush of spines |
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20 | Left paramere composed of two lobes. Ventral lobe of right paramere abruptly and strongly widened at apex |
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– | Left paramere simple |
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21 | Left paramere with small lateral basal tooth |
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– | Left paramere without small lateral basal tooth |
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22 | Left paramere split in two filiform branches behind middle |
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– | Left paramere simply filiform |
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23 | Left paramere bent twice |
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– | Left paramere evenly curved, without being clearly bent, before apex with tiny lateral tooth |
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24 | Dorsal lobe of right paramere very small |
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– | Dorsal lobe of right paramere large, nearly as long as ventral lobe |
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25 | Right paramere simple, not composed of two lobes |
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– | Right paramere composed of two lobes |
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26 | Left paramere simple |
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– | Left paramere composed of two lobes |
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27 | Left paramere simple |
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– | Left paramere composed of two lobes |
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28 | Dorsal lobe of right paramere wide, with sickle-shaped, large apical hook |
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– | Dorsal lobe of right paramere narrow, evenly curved and sharply pointed |
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29 | Dorsal lobe of right paramere triangular and short, sharply pointed |
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– | Dorsal lobe of right paramere convexly widened and elongate |
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Holotype: ♂ [China] “Mt. Daqingshan, Longzhou, Guangxi, 24.IV.1963, 600–700m, leg. Shi Yongshan” (IZAS). Paratype: 1 ♂ [China] “Mt. Daqingshan, Longzhou, Guangxi, 13.IV.1963, 360m, leg. Wang Shuyong” (ZFMK).
Body length: 9.2 mm, length of elytra: 7 mm, width: 5.5 mm.
Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Antennal club 1.1 times as long as remaining antennomeres combined. Eyes small; ratio of diameter/interocular width: 0.53. Ratio of length of metepisternum/metacoxa: 1/1.63. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin with a blunt tooth. Metatibia short and wide, ratio width/length: 1/3.1; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
Body length: 9.2 mm, length of elytra: 7.0–7.3 mm, width: 5.5–5.9 mm.
The new species differs from all other so far known
The new species is named after the type locality, Mt. Daqingshan.
Holotype: ♂ “China: Hubei; Dahongshan 1700m, Shuizhou VI-2003 leg. Ying et al.” (ZFMK). Paratypes. 1 ♂ [China] “Hekou, Southeast of Yunnan, 9.VI.1956, 1200m, leg. Panfilov” (IZAS), 1 ♂ [China] “Luxi, Yunnan, 22.V.1980, leg. Li Hongxing” (IZAS).
Body length: 9.1 mm, length of elytra: 7.3 mm, width: 5.4 mm.
Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.6. Antennal club 1.2 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.7. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin with a blunt tooth. Metatibia short and wide, ratio width/length: 1/3.2; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
Body length: 9.1–9.8 mm, length of elytra: 7.3–7.4 mm, width: 5.4–6.0 mm.
From the Latin word
Holotype: ♂ [China] “Wang Tong, to light, 29/4/07” (ZFMK). Paratypes: 1 ♂ [China] “Huangniushi, Mt. Jiulianshan, Jiangxi, 16.VI.1975, leg. Zhang Youwei” (IZAS), 1 ♂ [China] “Luoyang, Lianxian County, Guangdong, 22.VI.1965, leg. Zhang Youwei” (IZAS).
Body length: 9.1 mm, length of elytra: 6.9 mm, width: 5.8 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.6. Antennal club 1.3 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.7. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin straight, without blunt tooth. Metatibia short and wide, ratio width/length: 1/3.2; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
Body length: 8.2–9.1 mm, length of elytra: 6.5–6.9 mm, width: 5.5–5.8 mm.
Holotype: ♂ “Guangxi, Maoershan, 2011-VI-4/ LW-1066” (ZFMK).
Body length: 8.3 mm, length of elytra: 6.8 mm, width: 3.1 mm.
Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.61. Antennal club 1.2 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.5.
Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin with a blunt tooth. Metatibia short and wide, ratio width/length: 1/2.84; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
The new species is named after its type locality, Maoershan.
Holotype: ♂ “China, Hainan Isl., 4-6.v.2011 Limushan Mts. Frst administr. Centre (at light)
Body length: 9.7 mm, length of elytra: 7 mm, width: 6 mm.
Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.6. Antennal club 1.2 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.8. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin straight, without blunt tooth. Metatibia short and wide, ratio width/length: 1/2.7; basal group of dorsal spines of metatibia at first third of metatibial length. Aedeagus. Fig.
Body length: 8.0–9.7 mm, length of elytra: 6.0–7.0 mm, width: 4.8–6 mm. Head of the female paratype is missing.
The new species is named after one of the collectors of the type series, Martin Fikáček (Prague).
Holotype: ♂ “Bawangzhen, Changjiang, Hainan, 5–7.VI.2008, leg. Ba Yibin, Lang Juntong” (HBUM). Paratype: 1 ♂ “Mt. Jianfengling, Hainan, 10.VI.1965” (IZAS).
Body length: 8.4 mm, length of elytra: 6.5 mm, width: 5.4 mm.
Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.63. Antennal club 1.2 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.67.
Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin straight, without blunt tooth. Metatibia short and wide, ratio width/length: 1/2.74; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
Body length: 7.7–8.4 mm, length of elytra: 6.1–6.5 mm, width: 5.0–5.4 mm.
The new species is named after its type locality, Changjiang.
Holotype: ♂ “China: Sichuan; Wolong Reserve, Sigulian Shan,
Body length: 7.6 mm, length of elytra: 5.9 mm, width: 4.8 mm.
Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.6. Antennal club 1.2 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.4.
Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin straight, without blunt tooth. Metatibia short and wide, ratio width/length: 1/3; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
Body length: 6.6–7.6 mm, length of elytra: 5.1–5.9 mm, width: 3.9–4.8 mm.
The new species is named after its type locality, Sigulian Shan.
Body length: 7.6 mm, length of elytra: 5.8 mm, width: 4.6 mm.
Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.67. Antennal club 1.3 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.5. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin straight, without blunt tooth. Metatibia short and wide, ratio width/length: 1/3.21; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Body length: 6.9–7.6 mm, length of elytra: 5.1–5.8 mm, width: 4.0–4.6 mm. Female has the antennal club composed of three lamellae, short, as long as the remaining antennomeres combined; eyes as large as those in male.
The new species differs from the other known
The new species is named after its type locality, Damaidi.
Holotype: ♂ “Shunbi, Yangbi, Yunnan, 16.VIII.2009, leg. Shi Fuming” (HBUM).
Body length: 8.9 mm, length of elytra: 6.3 mm, width: 5 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.66. Antenna missing in holotype. Ratio of length of metepisternum/metacoxa: 1/1.5. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin straight, without blunt tooth. Metatibia short and wide, ratio width/length: 1/3.4; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
The new species differs from
The new species is named after its type locality, Shunbi.
Holotype: ♂ [China] “Nonggang, Longzhou, Guangxi, 15.VI.2000, 330m, leg. Chen Jun” (IZAS).
Body length: 7.5 mm, length of elytra: 5.5 mm, width: 4.6 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes small; ratio of diameter/interocular width: 0.48. Antennal club 1.1 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.48. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin straight, without blunt tooth. Metatibia short and wide, ratio width/length: 1/3.3; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
The new species is named after its type locality, Longzhou.
Holotype: ♂ [China] “Yao District, Mengla, Yunnan, 11.V.1991, leg. Liu Guangchun, Cai Wanzhi” (NUYS).
Body length: 8.3 mm, length of elytra: 6.5 mm, width: 5.1 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.62. Antennal club 1.6 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.5. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin straight, without blunt tooth. Metatibia short and wide, ratio width/length: 1/3.2; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
The new species is named after its type locality, Yaoqu.
Holotype: ♂ “China: E-Yunnan; Damaidi 2500m, Guangnan near Vietnam VII-2003 leg. Li et al.” (ZFMK). Paratypes: 1 ♂ “China, SE-Yunnan Xichou-E env. 1400–1700m, 13.–18.5.95/ 23°11–16'/ 104°41–49' L.+R. Businský lgt.” (CPPB), 1 ♂ [China] “Yunnan 2000–2500m
Body length: 7 mm, length of elytra: 5.3 mm, width: 4.6 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.7. Antennal club 1.2 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.4. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin straight, without blunt tooth. Metatibia short and wide, ratio width/length: 1/3.3; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
Body length: 7.0–7.8 mm, length of elytra: 5.3–5.6 mm, width: 4.4–4.6 mm.
The species name is derived from the combined Latin words,
Holotype: ♂ “China, S-Yunnan Prov. Xishuangbanna 23km NW Jinghong Na Ban village 680m
Body length: 7.6 mm, length of elytra: 5.8 mm, width: 5 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.63. Antenna missing in holotype. Ratio of length of metepisternum/metacoxa: 1/1.69. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin straight, without blunt tooth. Metatibia short and wide, ratio width/length: 1/3.36; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
Body length: 6.8–7.6 mm, length of elytra: 5.2–5.8 mm, width: 4.1–5.0 mm.
The species name is named after its type locality, Jinghong Na Ban village.
Holotype: ♂ [China] “Menglong, Yunnan, 22.IV.1982, light trap, leg. Jiang Shengqiao” (IZAS).
Body length: 7.6 mm, length of elytra: 5.8 mm, width: 4.3 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.59. Antennal club 1.2 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.64. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin straight, without blunt tooth. Metatibia short and wide, ratio width/length: 1/3.23; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
The new species is named after its type locality, Menglong.
Holotype: ♂ “Tianchi, Jianfeng, Hainan, 13.IV.1980, 900m, leg. Wang Shuyong” (IZAS). Paratype: 1 ♂ “Tianchi, Jianfeng, Hainan, 18.IV.1980, 750m, leg. Wang Shuyong” (ZFMK).
Body length: 9.1 mm, length of elytra: 6.9 mm, width: 5.7 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.63. Antennal club 1.3 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.56. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin straight, without blunt tooth. Metatibia short and wide, ratio width/length: 1/3.24; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
Body length: 9.1–10.5 mm, length of elytra: 6.9–8.4 mm, width: 5.7–6.1 mm.
The new species is named after its type locality, Tianchi.
Holotype: ♂ [China] “Yunnan, Lianghe, 2011-V-4,
Body length: 9.7 mm, length of elytra: 7.3 mm, width: 5.5 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.59. Antennal club 1.1 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.56. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin with a large sharp hook. Metatibia short and wide, ratio width/length: 1/3.1; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
The new species is named after its type locality, Lianghe.
Holotype: ♂ “China, W Yunnan prov., mts. 60km E Tengchong, 2200m, 19.–22.v.2006, S. Murzin & I. Shokhin” (CPPB). Paratype: 1 ♂ “Yunnan 2200-2500m
Body length: 8.9 mm, length of elytra: 7.2 mm, width: 5.7 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes small; ratio of diameter/interocular width: 0.5. Antennal club 1.2 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.6. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin with a large sharp hook. Metatibia short and wide, ratio width/length: 1/3.2; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
Body length: 8.9–9.8 mm, length of elytra: 7.1–7.2 mm, width: 5.5–5.7 mm.
The species name is derived from the combined Latin words,
Holotype: ♂ “China: Hunan; Mupu Mt. 1600m, Pingjiang VIII-2003, leg. Li et al.” (ZFMK). Paratypes:
Body length: 10.0 mm, length of elytra: 7.5 mm, width: 6.0 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.6. Antennal club 1.2 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.6. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin with a blunt tooth. Metatibia short and wide, ratio width/length: 1/2.9; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
The new species is named after its type locality, Pingjiang.
Body length: 8.8–10.0 mm, length of elytra: 6.9–7.5 mm, width: 5.4–6.0 mm. Female has the antennal club composed of three lamellae, short, as long as remaining antennomeres combined; eyes are as large as those in male.
Holotype: ♂ [China] “Yunnan, Wanding, 2011-IV-29,
Body length: 9.3 mm, length of elytra: 6.9 mm, width: 5.4 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes large; ratio of diameter/interocular width: 0.72. Antennal club 1.5 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.56. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin with a large sharp hook. Metatibia short and wide, ratio width/length: 1/3.13; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
Body length: 8.8–9.3 mm, length of elytra: 6.9–7.1 mm, width: 5.4–5.6 mm.
The new species is named after the type locality, Wanding.
Lectotype (here designated): ♂ [Vietnam] “Tonkin Montes Mauson April, Mai 2-3000' H. Fruhstorfer/ tonkinensis Mos.” (ZMHB). Paralectotypes: 1 ♀ [Vietnam] “Tonkin Montes Mauson April, Mai 2-3000' H. Fruhstorfer/ tonkinensis Mos.” (ZMHB), 5 ♀♀ [Vietnam] “Tonkin Montes Mauson April, Mai 2-3000' H. Fruhstorfer” (ZMHB).
1 ♂ [China] “Luoxiang, Jinxiu, Guangxi, 15.V.1999, 400m, leg. Li Wenzhu” (IZAS).
Body length: 9.0 mm, length of elytra: 7.0 mm, width: 5.9 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes large; ratio of diameter/interocular width: 0.66. Antennal club 1.2 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.45. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin with a large sharp hook. Metatibia short and wide, ratio width/length: 1/3.25; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Body length: 8.7–9.0 mm, length of elytra: 6.7–7.0 mm, width: 5.4–5.9 mm. Female has small eyes (ratio of diameter/interocular width: 0.6) and antennal club composed of 3 antennomeres being as long as the remaining antennomeres combined.
The species is recorded for the first time for China.
Holotype: ♂ “China, SE Yunnan, Xichou - E env. 1400–1700m, 13.18.5.95/ 23°22–26'N/ 104°41–49'E L.+R. Businský lgt.” (CPPB).
Body length: 9.0 mm, length of elytra: 6.7 mm, width: 5.8 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes large; ratio of diameter/interocular width: 0.7. Antennal club 1.2 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.6. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin with a large sharp hook. Metatibia short and wide, ratio width/length: 1/3.4; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
The new species is named after its type locality, Xichou.
Holotype: ♂ [China] “Meng'e, Xishuangbanna, Yunnan, 19.V.1958, 1050–1080m, leg. Hong Chunpei” (IZAS). Paratypes: 1 ♂ [China] “Menghai, Xishuangbanna, Yunnan, 19.VII.1958, 1200–1600m, leg. Wang Shuyong” (IZAS), 1 ♂ [China] “Menghun, Xishuangbanna, Yunnan, 18.V.1958, 1200–1400m, leg. Zhang Yiran” (IZAS), 1 ♂ [China] “Yunnan, Huanglianshan, 2012-V-9, 1800m/ LW-1277” (IZAS), 1 ♂ [China] “Meng'e, Xishuangbanna, Yunnan, 19.V.1958, 1050–1080m, leg. Hong Chunpei” (ZFMK).
Body length: 7.4 mm, length of elytra: 5.7 mm, width: 4.3 mm. Body reddish brown. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes large; ratio of diameter/interocular width: 0.7. Antennal club 1.2 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.5. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin straight. Metatibia short and wide, ratio width/length: 1/3.2; basal group of dorsal spines of metatibia at first third of metatibial length. Aedeagus. Fig.
Female unknown.
Body length: 7.4–8.6 mm, length of elytra: 5.7–7 mm, width: 4.3–5.9 mm.
The new species is named after its type locality, Meng'e.
Holotype: ♂ [China] “Nanmu Garden, Changshou, Sichuan, 9.VI.1994, 450m, leg. Zhang Youwei” (IZAS). Paratype: 1 ♂ [China] “Mt. Xingdoushan, Lichuan, Hubei, 22.VII.1989, light trap, 810m, leg. Wang Shuyong” (ZFMK).
Body length: 8.8 mm, length of elytra: 6.7 mm, width: 5.3 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.65. Antennal club as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.57. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin straight. Metatibia short and wide, ratio width/length: 1/3.35; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
Body length: 8.0–8.8 mm, length of elytra: 6.0–6.7 mm, width: 5.0–5.3 mm.
The new species is named after its type locality, Changshou.
Holotype: ♂ [China] “Hongqi Forestry Farm, Shangsi, Guangxi, 29.V.1999, 300m, leg. Ke Xin” (IZAS). Paratypes: 1 ♂ [China] “Nonggang, Longzhou, Guangxi, 21.V.1982, 240m, leg. Li Fasheng” (CAU), 1 ♂ [China] “Fu'ai, Pingxiang, Guangxi, 17.VI.1976, leg. Zhang Baolin” (ZFMK), 2 ♂♂ [China] “Hong Kong: Lantau I: San Shek Wan; v.1988” (BPBM), 1 ♂ [China] “Hongqi Forestry Farm, Shangsi, Guangxi, 29.V.1999, 300m, leg. Ke Xin” (IZAS), 1 ♂ [China] “Taojiang, Leishan, Guizhou, 5.VII.1988, 1000m, leg. Yang Longlong, light trap” (IZAS).
Body length: 8.1 mm, length of elytra: 6.2 mm, width: 4.6 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes large; ratio of diameter/interocular width: 0.76. Antennal club as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.61. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin straight. Metatibia short and wide, ratio width/length: 1/3.13; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
Body length: 7.0–8.1 mm, length of elytra: 5.6–6.7 mm, width: 4.4–4.8 mm.
The new species differs from the similar
The new species is named after its type locality in Shangsi prefecture (Guangxi province).
Holotype: ♂ [China] “Yunnan, Ruili, 2011-IV-27,
Body length: 8.1 mm, length of elytra: 5.8 mm, width: 4.8 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.63. Antennal club 1.2 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.74. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin straight. Metatibia moderately long and wide, ratio width/length: 1/3.54; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
The new species is named after its type locality, Ruili.
Holotype: ♂ [China] “Mts. Linaoshan, Langping, Tianlin, Guangxi, 28.V.2002, 1400m, leg. Jiang Guofang” (IZAS).
Body length: 8.8 mm, length of elytra: 6.6 mm, width: 5.2 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.63. Antennal club as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.5. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin straight. Metatibia moderately long and wide, ratio width/length: 1/3.69; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
The new species is named after its type locality, Mt. Linaoshan.
Holotype: ♂ [China] “Huyu County, Ruili, Yunnan, 11.VI.1956, 1400m, leg. Zhou Benshou” (IZAS).
Body length: 9 mm, length of elytra: 7 mm, width: 5.3 mm. Surface of labroclypeus and disc of frons glabrous. Smooth area anterior to eye twice as wide as long. Eyes moderately large; ratio of diameter/interocular width: 0.63. Antennal club 1.3 times as long as remaining antennomeres combined. Ratio of length of metepisternum/metacoxa: 1/1.55. Metafemur dull, anterior margin acute, without submarginal serrated line; anterior row of setae-bearing punctures absent; posterior margin straight. Metatibia moderately long and wide, ratio width/length: 1/3.47; basal group of dorsal spines of metatibia at first third of metatibial length.
Aedeagus. Fig.
Female unknown.
The new species is named after its type locality, Ruili.
Holotype: ♂ [China] “Cangchang, Anhua, Hunan, 15.VII.2004, leg. Wang Jiliang” (HBUM). Paratypes: 2 ♂♂ [China] “Hong Kong: N.T. Taipokau 12.VII.1965 Hand Net/ Lee Kit Ming & Hai Wai Ming Malaise Trap Bishop Museum” (BPBM, ZFMK), 1 ♂ [China] “Hong Kong: N.T. Taipokau 27.VI.1964/ M.J. Voss & Wai Ming Hui Collectors Bishop Museum” (BPBM), 1 ♂ [China] “Hong Kong: N.T. Taipokau 20.VI.1964/ Lee Kit Ming & Hai Wai Ming Light Trap Bishop Museum” (BPBM), 1 ♂, 2 ♀♀ [China] “Shekou, Fu‘an, Fujian, 26.VII.1963, leg. Zhang Youwei” (IZAS), 1 ♀ [China] “Shekou, Fu‘an, Fujian, 27.VII.1963, leg. Zhang Youwei” (IZAS), 2 ♂♂ [China] “Mt. Jiulianshan, Longnan, Jiangxi, 6.VI.1975, leg. Zhang Youwei” (IZAS), 1 ♂ [China] “Sidu, Guidong county, Hunan, 10.VII.2008, 774m, leg. Yang Ganyan” (IZAS), 2 ♂♂, 2 ♀♀ [China] “Qiliqiao, Chong'anxingcun, Fujian, 7,12,13.VII.1963, 840m, leg. Zhang Youwei” (IZAS).
Body length: 7.2 mm, length of elytra: 5.2 mm, width: 4.2 mm.
Labroclypeus surface with a few erect setae. Disc of frons with a few single setae. Smooth area in front of eye approximately 4 times as wide as long. Eyes large; ratio of diameter/interocular width: 0.82. Antennal club 1.4 times as long as remaining antennomeres combined.
Ratio of length of metepisternum/metacoxa: 1/1.5. Metafemur with a serrated continuous line beside anterior margin, with fine sparse punctures behind line each bearing a short seta.
Metatibia moderately long, ratio width/length: 1/3.25; basal group of dorsal spines of metatibia behind middle; beside dorsal margin in basal half with a blunt carina being partly finely serrate.
Aedeagus. Fig.
Female unknown.
Body length: 6.8–7.2 mm, length of elytra: 5.0–5.2 mm, width: 3.8–4.2 mm.
The new species is named after its type locality, Anhua.
Holotype: ♂ [China] “Leishan, Guizhou, 28.VI.1988, 800m, leg. Yin Huifen” (IZAS). Paratypes: 1 ♂ [China] “Mts. Fanjingshan, Jiangkou, Guizhou, 13.VIII.1988, 550m, leg. Yang Xingke” (ZFMK), 1 ♂ [China] “Mts. Fanjingshan, Jiangkou, Guizhou, 19.VIII.1988, 550m, leg. Yang Xingke” (IZAS), 1 ♂ [China] “Jiuniutang, Mao‘ershan, Guangxi, 13.VII.1985, 1100m, leg. Liao Subai” (IZAS).
Body length: 6.8 mm, length of elytra: 5 mm, width: 4 mm. Body reddish brown. Labroclypeus surface with a few erect setae. Disc of frons with a few single setae. Smooth area in front of eye approximately 4 times as wide as long. Eyes large; ratio of diameter/interocular width: 0.81. Antennal club 1.4 times as long as remaining antennomeres combined.
Ratio of length of metepisternum/metacoxa: 1/1.68. Metafemur with a serrated continuous line beside anterior margin, with fine sparse punctures behind line each bearing a short seta.
Metatibia moderately long, ratio width/length: 1/3.64; basal group of dorsal spines of metatibia behind middle; beside dorsal margin in basal half with a blunt carina being partly finely serrate.
Aedeagus. Fig.
Female unknown.
Body length: 6.8–7.7 mm, length of elytra: 5.0–5.6 mm, width: 4.0–4.6 mm.
The new species is named after its type locality, Leishan.
Holotype: ♂ [China] “Yaoan, Lianxian County, Guangdong, 28.VI.1965, leg. Zhang Youwei” (IZAS). Paratypes:
Body length: 7.2 mm, length of elytra: 5.4 mm, width: 4.3 mm. Body reddish brown. Labroclypeus surface with a few erect setae. Disc of frons with a few single setae. Smooth area in front of eye approximately 4 times as wide as long. Eyes large; ratio of diameter/interocular width: 0.86. Antennal club 1.3 times as long as remaining antennomeres combined.
Ratio of length of metepisternum/metacoxa: 1/1.68. Metafemur with a serrated continuous line beside anterior margin, with fine sparse punctures behind line each bearing a short seta.
Metatibia moderately long, ratio width/length: 1/3.82; basal group of dorsal spines of metatibia behind middle; beside dorsal margin in basal half with a blunt carina being partly finely serrate.
Aedeagus. Fig.
Female unknown.
Body length: 6.6–7.9 mm, length of elytra: 4.9–6.0 mm, width: 4.1–4.6 mm. Female has small eyes (ratio of diameter/interocular width: 0.51) and the antennal club composed of 3 antennomeres being as long as the remaining antennomeres combined.
The new species is named after its type locality, Yaoan.
Distribution of
Distribution of
We are grateful to L. Bartolozzi (MZUF), J. Hájek (NMPC), M. Hartmann (NME), A. Napolov (Riga), P. Pacholátko (Brno), B. Rattcliffe (USNM), Guo-Dong Ren (HBUM), G.A. Samuelson (BPBM), H. Schillhammer (NHMW), and Xin-Li Wang (CAU), for their kindly loaning to us unidentified specimens, to J. Frisch and M. Uhlig (both ZMHB) for help with the type specimen loans of the ZMHB, to M. Nikodým (Prague) and A. Weigel (Wernburg) for the donation of a part of their Asian