Corresponding author: Roger A. Burks (
Academic editor: M. Buffington
The endemic Australian parasitic wasp genus
The
The first Australian scelionids were described by
The current study focuses on one of the few genera,
The contributions of the individual authors are as follows; R.A. Burks: character definition, species concept development; key development, imaging, capture of specimen data, manuscript preparation, phylogenetic analysis and illustration; L. Masner: specimen acquisition, and generic overview; N.F. Johnson: generic concept development and manuscript preparation ; A.D. Austin: initial species concept development, manuscript preparation, and taxonomic overview.
Specimens examined were provided by the following collections: The American Entomological Institute, Gainesville, Florida, USA (AEIC)1; Australian Museum, Sydney, Australia (AMSA)2; Australian National Insect Collection, Canberra, Australia (ANIC)3; The Natural History Museum, London, United Kingdom (BMNH)4; Canadian National Collection of Insects, Arachnids and Nematodes, Ottawa, Canada (CNCI)5; Museum of Victoria, Entomology, Melbourne, Australia (MVMA)6; Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, USA (MCZC)7; C.A. Triplehorn Insect Collection, Ohio State University, Columbus, Ohio (OSUC)8; Queensland Primary Industries and Fisheries Insect Collection, Indooroopilly, Australia (QDPC)9; Queensland Museum, Brisbane, Australia (QMBA)10; R.M. Bohart Museum, University of California, Davis, USA (UCDC)12; Western Australian Museum, Perth, Australia (WAMP)13; Waite Insect and Nematode Collection, Adelaide, Australia (WINC)14. South Australian and Northern Territory holotypes for species newly described in this manuscript are deposited in the South Australian Museum, Adelaide, Australia (SAMA)11.
This revision is a product of the
Eye setose; frontal depression present; antenna 12-segmented; netrion present; postmarginal vein of fore wing present, longer than marginal and stigmal veins; mesotibia and metatibia each with 1 spur; metascutellum setose dorsally and ventrally; ovipositor
Body length: 2.75–6.88 mm (n=78).
Dodd did not specify the source of the name for this genus, but presumably it is derived the name of the Queensland town of Bracalba. (Dodd also used the names of Merriwa, NSW and Nyleta, QLD for other genera.) He originally combined this name with two species epithets of variable gender, both coined in feminine form:
Collection events for each
The hosts of
The implicit enumeration search found two optimum trees of 166 steps (strict consensus:
Strict consensus phylogram of two most parsimonious trees for species of
1 | Mandibles bidentate with no hint of a third tooth ( |
2 ( |
– | Mandibles either tridentate with a small middle tooth ( |
4 ( |
2 | Metascutellum flat ( |
|
– | Metascutellum projecting dorsally ( |
3 |
3 | Mesoscutellum sparsely foveolate, with smooth interspaces ( |
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– | Mesoscutellum densely foveolate, interspaces with irregular longitudinal rugae ( |
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4 | Propodeum with sharp protrusions (from lateral propodeal area) protruding dorsally immediately posterior to metascutellum, reaching at least the same height as the dorsal surface of metascutellum ( |
|
– | Propodeum without such protrusions | 5 |
5 | Metascutellum strongly narrowed apically ( |
6 |
– | Metascutellum with apex broadly truncate, as broad or broader than total metascutellar length ( |
10 |
6 | Mesosoma truncate anteriorly ( |
7 |
– | Mesosoma smoothly rounded anteriorly ( |
8 |
7 | Mesoscutellum without median carina ( |
|
– | Mesoscutellum with median carina ( |
|
8 | Antennal scrobes crossed by transverse carinae that meet at torular triangle ( |
|
– | Antennal scrobes foveolate or smooth. Females with hump near midlength T4 ( |
9 |
9 | Antennal scrobes smooth and distinct. Clypeus with blunt median carina ( |
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– | Antennal scrobes foveolate and indistinct ( |
|
10 | Frontal depression with strong longitudinal carina that is crossed by a strong transverse carina dorsally. Clypeus with median carina and transverse dorsal carina ( |
|
– | Frontal depression without such a set of carinae. Clypeus with weaker or no carinae ( |
11 |
11 | Metascutellum elongate-trapezoidal with concave lateral margins ( |
|
– | Metascutellum broadly trapezoidal with straight lateral margins ( |
12 |
12 | Mesoscutellum with a set of incomplete median longitudinal grooves ( |
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– | Mesoscutellum without median grooves ( |
13 |
13 | Mesoscutellum sparsely foveolate, with broad interspaces ( |
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– | Mesoscutellum densely foveolate (interspaces very narrow) except sometimes a small smooth antero-median area ( |
14 |
14 | Metascutellum trapezoidal in dorsal view, with straight lateral margins ( |
|
– | Metascutellum in dorsal view irregularly subrectangular or cushion-like, with irregular margins ( |
15 |
15 | Mesoscutellum with fine wrinkles or carinae in interspaces between foveolae ( |
|
– | Mesoscutellum with smooth interspaces between foveolae ( |
Ocular setae: short and dense. Frontal depression: smooth dorsally, ventrally with longitudinal carina and with additional carinae arising from medial margins of antennal foramina. Smooth depression extending dorsolaterally from antennal foramen: present. Dorsal clypeal margin: but without other sculpture. Clypeal median carina: absent. Ventral clypeal margin: with a small median point. Mandibular teeth: two, separated by narrow incision. Smooth area obliquely posterior to lateral ocellus: present. Genal sculpture: reticulate-rugose with strong dorsoventral carinae. Mandibular color: dark basally and at teeth, becoming lighter reddish brown between these areas.
Dorsal pronotal area: not set off by carina ventrally. Anterolateral corner of dorsal pronotal area: truncate anteriorly. Sculpture of posteromedian area of mesoscutum: foveolate with broad longitudinal septa. Lateral margin of dorsal axillar area: triangularly expanded or with posterior tooth, broadest posteriorly. Mesoscutellar sculpture: densely foveolate. Metascutellum in dorsal view: trapezoidal with broad apex. Dorsal surface of metascutellum: apex protruding dorsally. Femoral depression: crossed by rounded carinae. Anterior corner of lateral propodeal carina: flat, without tooth. Posteromedial corner of lateral propodeal area: protruding posteriorly. Leg color: yellowish-brown except for dorsal external part of coxae and sometimes tarsomeres 2-5.
Metasoma color: black to dark reddish brown. Median lobe of T1: with 7 or more longitudinal carinae, or with median smooth area interrupting carinae. Metasoma at middle of T4: without bend. T5 median carina: absent. Longitudinal sculptural septa on T5: weak, blunt and hardly raised. Transverse sculptural septa on T5: about as strong as the longitudinal septa. T5 setae: directed posteriorly, arising from anterior edge of sculptural mesh. T6: as broad or broader than long. T6 laterotergite: overlapping S6. S4 median carina: absent. Transverse sculptural septa on S5: weak or absent, much weaker than the longitudinal septa. S5 setae: directed posteriorly, arising from anterior edge of sculptural mesh. Lateral carinae of S6: absent. Apex of S6: without notch.
Holotype, female: AUSTRALIA: QLD, Chinchilla, 8.II.1928, A. P. Dodd, QMBA HY4733 (deposited in QMBA). Paratypes: AUSTRALIA: 2 females, OSUC 238448-238449 (ANIC). Other material: AUSTRALIA: 32 females, 20 males, OSUC 238516 (BMNH); OSUC 238120, 238123-238128, 238130-238153, 238159-238166, 238171-238173, 238176, 238180-238181, 238188, 238201-238202 (CNCI); OSUC 376915 (MCZC); OSUC 238451, QM Reg. No. T35161 (QDPC); OSUC 231779 (WINC).
Ocular setae: short and dense. Frontal depression: smooth dorsally, torular triangle foveolate with 1 transverse carina extending laterally from inner margin of antennal foramen. Smooth depression extending dorsolaterally from antennal foramen: present. Dorsal clypeal margin: but without other sculpture. Clypeal median carina: absent. Ventral clypeal margin: with a small median point. Mandibular color: dark basally and at teeth, becoming lighter reddish brown between these areas. Mandibular teeth: two, separated by narrow incision. Smooth area obliquely posterior to lateral ocellus: present. Genal sculpture: reticulate-rugose without any strong carinae.
Dorsal pronotal area: not set off by carina ventrally. Anterolateral corner of dorsal pronotal area: weakly rounded anteriorly. Sculpture of posteromedian area of mesoscutum: foveolate with slightly stronger longitudinal septa. Lateral margin of dorsal axillar area: triangularly expanded or with posterior tooth, broadest posteriorly. Mesoscutellar sculpture: densely foveolate with slightly stronger longitudinal septa. Metascutellum in dorsal view: trapezoidal with broad apex. Dorsal surface of metascutellum: apex protruding dorsally. Femoral depression: crossed by rounded carinae. Leg color: dark except for tibiae, 2nd trochanter, and tarsomeres 2-5. Anterior corner of lateral propodeal carina: flat, without tooth. Posteromedial corner of lateral propodeal area: protruding posteriorly.
Metasoma color: black to dark reddish brown. Median lobe of T1: with a set of rugae that merge with one another. Metasoma at middle of T4: without bend. Posterolateral margins of metasomal terga: without protrusions. T5 median carina: absent. Longitudinal sculptural septa on T5: weak, blunt and hardly raised. Transverse sculptural septa on T5: about as strong as the longitudinal septa. T5 setae: directed posteriorly, arising from anterior edge of sculptural mesh. T6: as broad or broader than long. T6 laterotergite: overlapping S6. S4 median carina: absent. Transverse sculptural septa on S5: weak or absent, much weaker than the longitudinal septa. S5 setae: directed posteriorly, arising from anterior edge of sculptural mesh. Lateral carinae of S6: absent. Apex of S6: without notch.
Latin adjective, named after the convex mesoscutum of most specimens in this species.
Holotype, female: AUSTRALIA: WA, Kalbarri National Park, 12.XII-19.XII.1986, malaise trap/pan trap, J. S. Noyes, OSUC 148701 (deposited in WAMP). Paratype: AUSTRALIA: 1 female, OSUC 148612 (CNCI).
Ocular setae: short and dense. Frontal depression: with many irregularly transverse rugae. Smooth depression extending dorsolaterally from antennal foramen: present. Dorsal clypeal margin: absent between antennal foramina. Clypeal median carina: absent. Ventral clypeal margin: with a small median point. Mandibular color: dark basally and at teeth, becoming lighter reddish brown between these areas. Mandibular teeth: two, separated by narrow incision. Smooth area obliquely posterior to lateral ocellus: present. Genal sculpture: deeply reticulate-rugose with some septa much stronger than others, forming distinct rows differing in height.
Dorsal pronotal area: not set off by carina ventrally. Anterolateral corner of dorsal pronotal area: weakly rounded anteriorly. Sculpture of posteromedian area of mesoscutum: densely foveolate. Lateral margin of dorsal axillar area: triangularly expanded or with posterior tooth, broadest posteriorly. Mesoscutellar sculpture: sparsely foveolate, with large smooth interspaces. Metascutellum in dorsal view: semicircular. Dorsal surface of metascutellum: flat. Femoral depression: centrally smooth, peripherally foveolate. Leg color: coxae dark, leg becoming gradually lighter apically. Anterior corner of lateral propodeal carina: flat, without tooth. Posteromedial corner of lateral propodeal area: protruding posteriorly.
Metasoma color: black to dark reddish brown. Median lobe of T1: with 7 or more longitudinal carinae. Metasoma at middle of T4: without bend. Posterolateral margins of metasomal terga: without protrusions. T5 median carina: absent. Longitudinal sculptural septa on T5: weak, blunt and hardly raised. Transverse sculptural septa on T5: about as strong as the longitudinal septa. T5 setae: directed posteriorly, arising from anterior edge of sculptural mesh. T6: as broad or broader than long. T6 laterotergite: overlapped by rim from S6. S4 median carina: absent. Transverse sculptural septa on S5: about as strong as the longitudinal septa. S5 setae: directed posteriorly, arising from anterior edge of sculptural mesh. Lateral carinae of S6: absent. Apex of S6: without notch.
Latin adjective, referring to the flat mesoscutum, mesoscutellum, and metascutellum of most specimens of this species.
Holotype, female: AUSTRALIA: NT, Centre for Zone Research (CSIRO), Alice Springs, XI-1992, malaise trap, Austin & Dangerfield, OSUC 230804 (deposited in SAMA). Paratypes: AUSTRALIA: 5 females, 3 males, OSUC 148706, 238115, 238168, 238170 (CNCI); OSUC 230802-230803, 230805-230806 (WINC).
Most specimens have a flat mesoscutum and mesoscutellum, but these features were not constant within this species nor in others of the genus.
Ocular setae: short and dense. Frontal depression: with many irregularly transverse rugae, or smooth dorsally, ventrally with oblique carinae converging on a longitudinal ruga. Smooth depression extending dorsolaterally from antennal foramen: present. Dorsal clypeal margin: forming a complete connection between antennal foramina medially. Clypeal median carina: absent. Ventral clypeal margin: with a small median point. Mandibular color: dark basally and at teeth, becoming lighter reddish brown between these areas. Mandibular teeth: three, but middle tooth tiny. Smooth area obliquely posterior to lateral ocellus: present. Genal sculpture: deeply reticulate-rugose with some septa much stronger than others, forming distinct rows differing in height.
Dorsal pronotal area: not set off by carina ventrally. Anterolateral corner of dorsal pronotal area: weakly rounded anteriorly. Sculpture of posteromedian area of mesoscutum: foveolate with broad longitudinal septa. Lateral margin of dorsal axillar area: triangularly expanded or with posterior tooth, broadest posteriorly. Mesoscutellar sculpture: densely foveolate but with smooth central area. Metascutellum in dorsal view: strongly transverse, subrectangular. Dorsal surface of metascutellum: flat, or convex. Femoral depression: centrally smooth, peripherally foveolate. Leg color: coxae, femora (aside from their apices), and at least the last two tarsomeres dark, otherwise yellowish brown. Anterior corner of lateral propodeal carina: flat, without tooth. Posteromedial corner of lateral propodeal area: not protruding posteriorly.
Metasoma color: black to dark reddish brown. Median lobe of T1: with 7 or more longitudinal carinae. Metasoma at middle of T4: with very weak bend. Posterolateral margins of metasomal terga: with tooth-like protrusions. T5 median carina: absent. Longitudinal sculptural septa on T5: strong, sharply raised. Transverse sculptural septa on T5: about as strong as the longitudinal septa. T5 setae: directed posteriorly, arising from anterior edge of sculptural mesh. T6: as broad or broader than long. T6 laterotergite: overlapping S6. S4 median carina: present. Transverse sculptural septa on S5: weak or absent, much weaker than the longitudinal septa. S5 setae: directed posteriorly, arising from anterior edge of sculptural mesh. Lateral carinae of S6: absent. Apex of S6: without notch.
Latin adjective meaning “clavate,” named for the clavate antenna with its distinct separation between the 6-segmented club and the basal flagellomeres.
Holotype, female: AUSTRALIA: SA, Brookfield Conservation Park,
Ocular setae: short and sparse. Frontal depression: smooth dorsally, torular triangle foveolate with transverse carinae lateral to this area. Smooth depression extending dorsolaterally from antennal foramen: present. Dorsal clypeal margin: wrinkle-like with a median peak. Clypeal median carina: present. Ventral clypeal margin: with a small median point. Mandibular color: dark basally and at teeth, becoming lighter reddish brown between these areas. Mandibular teeth: three of roughly equal size. Smooth area obliquely posterior to lateral ocellus: present. Genal sculpture: reticulate-rugose without any strong carinae.
Dorsal pronotal area: not set off by carina ventrally. Anterolateral corner of dorsal pronotal area: weakly rounded anteriorly. Sculpture of posteromedian area of mesoscutum: densely foveolate. Lateral margin of dorsal axillar area: with a semicircular expansion, broadest near midlength. Mesoscutellar sculpture: densely foveolate with one or two median longitudinal channels. Metascutellum in dorsal view: trapezoidal with broad apex. Dorsal surface of metascutellum: convex. Femoral depression: centrally smooth, peripherally foveolate. Leg color: dark except for trochanters, tips of femora and tibiae, and tarsomeres 1-4. Anterior corner of lateral propodeal carina: flat, without tooth. Posteromedial corner of lateral propodeal area: protruding posteriorly.
Metasoma color: black to dark reddish brown. Median lobe of T1: with 7 or more longitudinal carinae. Metasoma at middle of T4: with metasomal bend and abrupt transition in sculpture. Posterolateral margins of metasomal terga: without protrusions. T5 median carina: absent. or present. Longitudinal sculptural septa on T5: weak, blunt and hardly raised. Transverse sculptural septa on T5: about as strong as the longitudinal septa. T5 setae: directed posteriorly, arising from anterior edge of sculptural mesh. T6: longer than broad. T6 laterotergite: overlapped by rim from S6. S4 median carina: absent. Transverse sculptural septa on S5: weak or absent, much weaker than the longitudinal septa. S5 setae: not directed posteriorly, arising from center of sculptural mesh. Lateral carinae of S6: absent. Apex of S6: without notch.
Latin adjective, referring to the geographic distribution of this species.
Holotype, female: AUSTRALIA: WA, 3km W Walpole, Keystone Road,
Some singleton specimens similar to
Ocular setae: short and dense. Frontal depression: medially smooth, torular triangle sparsely foveolate, or foveolate dorsally, torular triangle foveolate, areas lateral to it with transverse carinae. Smooth depression extending dorsolaterally from antennal foramen: absent. Dorsal clypeal margin: bordering antennal foramina, absent between them. Clypeal median carina: absent. Ventral clypeal margin: convex. Mandibular color: dark basally and at teeth, becoming lighter reddish brown between these areas. Mandibular teeth: three, but middle tooth tiny. Smooth area obliquely posterior to lateral ocellus: present. Genal sculpture: reticulate-rugose without any strong carinae.
Dorsal pronotal area: not set off by carina ventrally. Anterolateral corner of dorsal pronotal area: weakly rounded anteriorly. Sculpture of posteromedian area of mesoscutum: foveolate with slightly stronger longitudinal septa. Lateral margin of dorsal axillar area: triangularly expanded or with posterior tooth, broadest posteriorly. Mesoscutellar sculpture: densely foveolate with slightly stronger longitudinal septa. Metascutellum in dorsal view: strongly transverse, subrectangular. Dorsal surface of metascutellum: flat. Femoral depression: centrally smooth, peripherally foveolate. Leg color: coxae, femora (aside from their apices), and at least the last two tarsomeres dark, otherwise yellowish brown. Anterior corner of lateral propodeal carina: flat, without tooth. Posteromedial corner of lateral propodeal area: protruding posteriorly.
Metasoma color: black to dark reddish brown. Median lobe of T1: with a set of rugae that merge with one another. Metasoma at middle of T4: without bend. Posterolateral margins of metasomal terga: without protrusions. T5 median carina: absent. Longitudinal sculptural septa on T5: weak, blunt and hardly raised. Transverse sculptural septa on T5: about as strong as the longitudinal septa. T5 setae: directed posteriorly, arising from anterior edge of sculptural mesh. T6: longer than broad. T6 laterotergite: overlapping S6. S4 median carina: present. Transverse sculptural septa on S5: weak or absent, much weaker than the longitudinal septa. S5 setae: directed posteriorly, arising from anterior edge of sculptural mesh. Lateral carinae of S6: absent. Apex of S6: without notch.
Latin adjective, referring to its unusual mixture of characters.
Holotype, female: AUSTRALIA: WA, via Dwellingup, Yarragil 4P Catchment, 18.II-25.II.1981, malaise trap, A. Postle, OSUC 230811 (deposited in QMBA). Paratypes: AUSTRALIA: 8 females, 23 males, OSUC 148705, 148707-148708, 180713, 238105-238109, 238111, 238179, 238182, 238184-238186 (CNCI); OSUC 230809-230810, 230812, 230814, 238450 (QDPC); OSUC 227572-227578, 230801, 238454, 238457, 384567 (WINC).
Dorsal pronotal area: not set off by carina ventrally. Anterolateral corner of dorsal pronotal area: truncate anteriorly. Sculpture of posteromedian area of mesoscutum: foveolate with slightly stronger longitudinal septa. Lateral margin of dorsal axillar area: with a semicircular expansion, broadest near midlength. Mesoscutellar sculpture: densely foveolate with a longitudinal carina. Metascutellum in dorsal view: elongate-trapezoidal but with incised apex. Dorsal surface of metascutellum: apex protruding dorsally. Femoral depression: crossed by rounded carinae. Anterior corner of lateral propodeal carina: flat, without tooth. Posteromedial corner of lateral propodeal area: protruding posteriorly. T7: arched and posteriorly concave.
Males: Lateral margin of dorsal axillar area semicircularly expanded; mesosoma broadly truncate anteriorly; mesoscutellum with median carina; metascutellum elongate-trapezoidal with incised apex.
Holotype, female: AUSTRALIA: QLD, Gogango, 40mi W Rockhampton, III-1928, A. P. Dodd, QMBA HY4732H (deposited in QMBA). Allotype: AUSTRALIA: 1 male, QMBA HY4732A (QMBA). Other material: AUSTRALIA: 3 males, OSUC 365210-365212 (UQIC).
Only the antenna and a few legs of the female holotype of
Because the antenna and some legs of the holotype remains, they now represent the holotype. A neotype cannot be properly designated without first requesting that the existing holotype be set aside. This was considered unnecessary, as the antenna clearly belongs to
Ocular setae: long and sparse. Frontal depression: medially smooth, torular triangle sparsely foveolate. Smooth depression extending dorsolaterally from antennal foramen: present. Dorsal clypeal margin: interrupted by a median areole below interantennal process. Clypeal median carina: present. Ventral clypeal margin: with a small median point. Mandibular color: dark basally and at teeth, becoming lighter reddish brown between these areas. Mandibular teeth: three, but middle tooth tiny. Smooth area obliquely posterior to lateral ocellus: present. Genal sculpture: reticulate-rugose without any strong carinae.
Dorsal pronotal area: not set off by carina ventrally. Anterolateral corner of dorsal pronotal area: protruding anteriorly. Sculpture of posteromedian area of mesoscutum: densely foveolate. Lateral margin of dorsal axillar area: with a semicircular expansion, broadest near midlength. Mesoscutellar sculpture: densely foveolate. Metascutellum in dorsal view: elongate-trapezoidal but with incised apex. Dorsal surface of metascutellum: apex protruding dorsally, or convex. Femoral depression: crossed by rounded carinae. Leg color: reddish but with at least last two tarsomeres dark. Anterior corner of lateral propodeal carina: flat, without tooth. Posteromedial corner of lateral propodeal area: protruding posteriorly.
Metasoma color: reddish, with last segment variably dark. Median lobe of T1: with 7 or more longitudinal carinae. Metasoma at middle of T4: with metasomal bend and abrupt transition in sculpture. Posterolateral margins of metasomal terga: with tooth-like protrusions. T5 median carina: present. Longitudinal sculptural septa on T5: strong, sharply raised. Transverse sculptural septa on T5: weak or absent, much weaker than the longitudinal septa. T5 setae: not directed posteriorly, arising from center of sculptural mesh. T6: longer than broad. T6 laterotergite: overlapping S6. S4 median carina: present. Transverse sculptural septa on S5: weak or absent, much weaker than the longitudinal septa. S5 setae: not directed posteriorly, arising from center of sculptural mesh. Lateral carinae of S6: not meeting apically. Apex of S6: with notch.
Compound adjective using the Latin adjectives magnus and ruber.
Holotype, female: AUSTRALIA: SA, 32km N Renmark, Amalia Dam, xeric mallee scrub, MT 4, ROM 2000040, Bookmark Biosphere Reserve,
This is the largest-bodied species of
Dorsal pronotal area: not set off by carina ventrally. Anterolateral corner of dorsal pronotal area: weakly rounded anteriorly. Sculpture of posteromedian area of mesoscutum: sparsely foveolate, with large smooth interspaces. Lateral margin of dorsal axillar area: with a semicircular expansion, broadest near midlength. Mesoscutellar sculpture: sparsely foveolate, with large smooth interspaces. Metascutellum in dorsal view: elongate-trapezoidal but with incised apex. Dorsal surface of metascutellum: apex protruding dorsally. Femoral depression: crossed by rounded carinae. Anterior corner of lateral propodeal carina: flat, without tooth. Posteromedial corner of lateral propodeal area: protruding posteriorly. Leg color: coxae and at least the last three tarsomeres dark, otherwise yellowish. T7: arched and posteriorly concave.
Males: Lateral margin of dorsal axillar area semicircularly expanded; mesosoma weakly rounded anteriorly; mesoscutellum without median carina; metascutellum elongate-trapezoidal with incised apex.
Holotype, male,
The male holotype of
Ocular setae: long and sparse. Frontal depression: smooth dorsally, torular triangle foveolate, areas lateral to it smooth. Smooth depression extending dorsolaterally from antennal foramen: present. Dorsal clypeal margin: arched, interrupted by broad median carina. Clypeal median carina: present. Ventral clypeal margin: with a small median point. Mandibular color: dark basally and at teeth, becoming lighter reddish brown between these areas. Mandibular teeth: three, but middle tooth tiny. Smooth area obliquely posterior to lateral ocellus: present. Genal sculpture: deeply reticulate-rugose with some septa much stronger than others, forming distinct rows differing in height.
Dorsal pronotal area: not set off by carina ventrally. Anterolateral corner of dorsal pronotal area: protruding anteriorly. Sculpture of posteromedian area of mesoscutum: foveolate with slightly stronger longitudinal septa. Lateral margin of dorsal axillar area: with a semicircular expansion, broadest near midlength. Mesoscutellar sculpture: densely foveolate with one or two median longitudinal channels. Metascutellum in dorsal view: elongate-trapezoidal but with incised apex. Dorsal surface of metascutellum: flat, or convex. Femoral depression: crossed by many sharply defined carinae. Leg color: yellowish-brown except for dorsal external part of coxae and sometimes tarsomeres 2-5. Anterior corner of lateral propodeal carina: flat, without tooth. Posteromedial corner of lateral propodeal area: protruding posteriorly.
Metasoma color: centrally reddish with some vague dark intrusions, peripherally becoming black. Median lobe of T1: with 7 or more longitudinal carinae. Metasoma at middle of T4: with metasomal bend and abrupt transition in sculpture. Posterolateral margins of metasomal terga: with tooth-like protrusions. T5 median carina: absent. Longitudinal sculptural septa on T5: strong, sharply raised. Transverse sculptural septa on T5: about as strong as the longitudinal septa. T5 setae: directed posteriorly, arising from anterior edge of sculptural mesh. T6: as broad or broader than long. T6 laterotergite: overlapped by rim from S6. S4 median carina: present. Transverse sculptural septa on S5: weak or absent, much weaker than the longitudinal septa. S5 setae: directed posteriorly, arising from anterior edge of sculptural mesh. Lateral carinae of S6: forming complete peripheral carina. Apex of S6: without notch.
Compound adjective using the Latin adjectives parvus and ruber.
Holotype, female: AUSTRALIA: WA, 20km N Denmark, 16.I.1987, J. S. Noyes, OSUC 238194 (deposited in WAMP). Paratypes: AUSTRALIA: 3 females, OSUC 238191, 238193, 238196 (CNCI).
Ocular setae: short and dense. Frontal depression: with many strong transverse carinae, sparsely foveolate at torular triangle. Smooth depression extending dorsolaterally from antennal foramen: present. Dorsal clypeal margin: absent between antennal foramina. Clypeal median carina: absent. Ventral clypeal margin: with a small median point. Mandibular color: dark basally and at teeth, becoming lighter reddish brown between these areas. Mandibular teeth: three of roughly equal size. Smooth area obliquely posterior to lateral ocellus: present. Genal sculpture: deeply reticulate-rugose with some septa much stronger than others, forming distinct rows differing in height.
Dorsal pronotal area: not set off by carina ventrally. Anterolateral corner of dorsal pronotal area: weakly rounded anteriorly. Sculpture of posteromedian area of mesoscutum: densely foveolate. Lateral margin of dorsal axillar area: with a semicircular expansion, broadest near midlength. Mesoscutellar sculpture: densely foveolate. Metascutellum in dorsal view: trapezoidal with broad apex. Dorsal surface of metascutellum: flat, or convex. Femoral depression: centrally smooth, peripherally foveolate. Leg color: coxae and femora dark, but tibiae, tarsi, and sometimes 2nd trochanters yellowish brown. Anterior corner of lateral propodeal carina: flat, without tooth. Posteromedial corner of lateral propodeal area: protruding posteriorly.
Metasoma color: black to dark reddish brown. Median lobe of T1: with a set of rugae that merge with one another. Metasoma at middle of T4: with metasomal bend and abrupt transition in sculpture. Posterolateral margins of metasomal terga: without protrusions. T5 median carina: absent. Longitudinal sculptural septa on T5: weak, blunt and hardly raised. Transverse sculptural septa on T5: about as strong as the longitudinal septa. T5 setae: directed posteriorly, arising from anterior edge of sculptural mesh. T6: as broad or broader than long. T6 laterotergite: overlapping S6. S4 median carina: present. Transverse sculptural septa on S5: about as strong as the longitudinal septa. S5 setae: directed posteriorly, arising from anterior edge of sculptural mesh. Lateral carinae of S6: absent. Apex of S6: without notch.
Latin noun, meaning “a small fin.” This is considered to be a noun in apposition to the generic name.
Holotype, female: AUSTRALIA: WA, WA122401, 7km N Stirling Range National Park,
Ocular setae: short and dense. Frontal depression: present as a vague smooth triangle. Smooth depression extending dorsolaterally from antennal foramen: present. Dorsal clypeal margin: absent between antennal foramina. Clypeal median carina: absent. Ventral clypeal margin: with a small median point. Mandibular color: dark basally and at teeth, becoming lighter reddish brown between these areas. Mandibular teeth: three, but middle tooth tiny. Smooth area obliquely posterior to lateral ocellus: present. Genal sculpture: deeply reticulate-rugose with some septa much stronger than others, forming distinct rows differing in height.
Dorsal pronotal area: not set off by carina ventrally. Anterolateral corner of dorsal pronotal area: weakly rounded anteriorly. Sculpture of posteromedian area of mesoscutum: foveolate with broad longitudinal septa. Lateral margin of dorsal axillar area: with a semicircular expansion, broadest near midlength. Mesoscutellar sculpture: sparsely foveolate, with large smooth interspaces. Metascutellum in dorsal view: very short, subrectangular. Dorsal surface of metascutellum: convex. Femoral depression: irregularly foveolate but not crossed by carinae. Leg color: coxae, femora (aside from their apices), and at least the last two tarsomeres dark, otherwise yellowish brown. Anterior corner of lateral propodeal carina: with longitudinal tooth-like crest extending dorsally above level of metascutellum. Posteromedial corner of lateral propodeal area: protruding posteriorly.
Metasoma color: black to dark reddish brown. Median lobe of T1: with 7 or more longitudinal carinae. Metasoma at middle of T4: with metasomal bend and abrupt transition in sculpture. Posterolateral margins of metasomal terga: without protrusions. T5 median carina: absent. Longitudinal sculptural septa on T5: weak, blunt and hardly raised. Transverse sculptural septa on T5: about as strong as the longitudinal septa. T5 setae: directed posteriorly, arising from anterior edge of sculptural mesh. T6: longer than broad. T6 laterotergite: overlapping S6. S4 median carina: present. Transverse sculptural septa on S5: about as strong as the longitudinal septa. S5 setae: not directed posteriorly, arising from center of sculptural mesh. Lateral carinae of S6: absent. Apex of S6: without notch.
An adjective, combining the anatomical term propodeum with the Latin adjectival suffix -alis, referring to its unique propodeum.
Holotype, female: AUSTRALIA: WA, Mount Cooke, 28.I-17.II.1991, malaise trap, M. S. Harvey & J. M. Waldock, OSUC 227590 (deposited in WAMP). Paratypes: AUSTRALIA: 2 females, OSUC 238190 (CNCI); OSUC 227591 (WINC).
The propodeum in this species is unique among the specimens studied. It is very difficult to determine in all examined specimens if the metasomal bend is present or not.
Ocular setae: short and dense. Frontal depression: with irregular rugae indicating large foveae. Smooth depression extending dorsolaterally from antennal foramen: present. Dorsal clypeal margin: angular, emarginate medially. Clypeal median carina: present. Ventral clypeal margin: with a small median point. Mandibular color: dark basally and at teeth, becoming lighter reddish brown between these areas. Mandibular teeth: three, but middle tooth tiny. Smooth area obliquely posterior to lateral ocellus: present. Genal sculpture: reticulate-rugose with strong dorsoventral carinae.
Dorsal pronotal area: not set off by carina ventrally. Anterolateral corner of dorsal pronotal area: truncate anteriorly. Sculpture of posteromedian area of mesoscutum: densely foveolate. Lateral margin of dorsal axillar area: with a semicircular expansion, broadest near midlength. Mesoscutellar sculpture: densely foveolate. Metascutellum in dorsal view: trapezoidal with broad apex. Dorsal surface of metascutellum: apex protruding dorsally. Femoral depression: centrally smooth, peripherally foveolate. Leg color: coxae and at least the last three tarsomeres dark, otherwise reddish. Anterior corner of lateral propodeal carina: flat, without tooth. Posteromedial corner of lateral propodeal area: protruding posteriorly.
Metasoma color: T2 to flat part of T4, troughs of S1 to variable part of S5 reddish, otherwise dark. Median lobe of T1: with 7 or more longitudinal carinae. Metasoma at middle of T4: with metasomal bend and abrupt transition in sculpture. Posterolateral margins of metasomal terga: without protrusions. T5 median carina: absent. Longitudinal sculptural septa on T5: strong, sharply raised. Transverse sculptural septa on T5: weak or absent, much weaker than the longitudinal septa. T5 setae: not directed posteriorly, arising from center of sculptural mesh. T6: longer than broad. T6 laterotergite: overlapping S6. S4 median carina: present. Transverse sculptural septa on S5: weak or absent, much weaker than the longitudinal septa. S5 setae: not directed posteriorly, arising from center of sculptural mesh. Lateral carinae of S6: not meeting apically. Apex of S6: without notch.
Used as a noun in apposition to the generic name, derived from a combination of the Latin participle sculptus and noun frons, referring to its strong network of frontal carinae.
Holotype, female: AUSTRALIA: WA, ~25km E Perth, John Forrest National Park, 23.XII-27.XII.1986, J. S. Noyes, OSUC 231799 (deposited in WAMP). Paratype: AUSTRALIA: 1 female, OSUC 148704 (CNCI).
Ocular setae: long and sparse. Frontal depression: with irregular rugae indicating large foveae. Smooth depression extending dorsolaterally from antennal foramen: present. Dorsal clypeal margin: forming a complete connection between antennal foramina medially. Clypeal median carina: absent. Ventral clypeal margin: with a small median point. Mandibular color: dark basally and at teeth, becoming lighter reddish brown between these areas. Mandibular teeth: two, separated by narrow incision. Smooth area obliquely posterior to lateral ocellus: present. Genal sculpture: reticulate-rugose without any strong carinae.
Dorsal pronotal area: not set off by carina ventrally. Anterolateral corner of dorsal pronotal area: truncate anteriorly. Sculpture of posteromedian area of mesoscutum: densely foveolate. Lateral margin of dorsal axillar area: with a semicircular expansion, broadest near midlength. Mesoscutellar sculpture: densely foveolate. Metascutellum in dorsal view: trapezoidal but tapering to narrow apex. Dorsal surface of metascutellum: apex protruding dorsally. Femoral depression: crossed by rounded carinae. Leg color: entirely reddish. Anterior corner of lateral propodeal carina: flat, without tooth. Posteromedial corner of lateral propodeal area: protruding posteriorly.
Metasoma color: mostly reddish, variably dark at T4 bend, T5-T6, with variable smaller dark areas elsewhere. Median lobe of T1: with 7 or more longitudinal carinae. Metasoma at middle of T4: with metasomal bend and abrupt transition in sculpture. Posterolateral margins of metasomal terga: with tooth-like protrusions. T5 median carina: absent. Longitudinal sculptural septa on T5: strong, sharply raised. Transverse sculptural septa on T5: weak or absent, much weaker than the longitudinal septa. T5 setae: not directed posteriorly, arising from center of sculptural mesh. T6: as broad or broader than long. T6 laterotergite: overlapping S6. S4 median carina: present. Transverse sculptural septa on S5: weak or absent, much weaker than the longitudinal septa. S5 setae: not directed posteriorly, arising from center of sculptural mesh. Lateral carinae of S6: not meeting apically. Apex of S6: with notch.
Latin participle meaning “scattered,” recalling the sparsely distributed setae of this species.
Holotype, female: AUSTRALIA: WA, Toodyay, 31.X.1979, R. M. Bohart, OSUC 231798 (deposited in WAMP). Paratypes: AUSTRALIA: 4 females, 3 males, OSUC 230821 (ANIC); OSUC 148703, 238204-238206, 238208 (CNCI); OSUC 55870 (OSUC).
Ocular setae: short and sparse. Frontal depression: with many strong transverse carinae, sparsely foveolate at torular triangle. Smooth depression extending dorsolaterally from antennal foramen: present. Dorsal clypeal margin: bordering antennal foramina, absent between them. Clypeal median carina: absent. Ventral clypeal margin: with a small median point. Mandibular color: mostly yellowish brown, dark at teeth. Mandibular teeth: three, but middle tooth tiny. Smooth area obliquely posterior to lateral ocellus: present. Genal sculpture: deeply reticulate-rugose with some septa much stronger than others, forming distinct rows differing in height.
Dorsal pronotal area: not set off by carina ventrally. Anterolateral corner of dorsal pronotal area: weakly rounded anteriorly. Sculpture of posteromedian area of mesoscutum: foveolate with slightly stronger longitudinal septa. Lateral margin of dorsal axillar area: with a semicircular expansion, broadest near midlength. Mesoscutellar sculpture: sparsely foveolate, with large smooth interspaces. Metascutellum in dorsal view: trapezoidal with broad apex. Dorsal surface of metascutellum: convex. Femoral depression: crossed by rounded carinae. Leg color: coxae, femora (aside from their apices), and at least the last two tarsomeres dark, otherwise yellowish brown. Anterior corner of lateral propodeal carina: flat, without tooth. Posteromedial corner of lateral propodeal area: protruding posteriorly.
Metasoma color: black to dark reddish brown. Median lobe of T1: with a set of rugae that merge with one another. Metasoma at middle of T4: with very weak bend. Posterolateral margins of metasomal terga: without protrusions. T5 median carina: absent. Longitudinal sculptural septa on T5: weak, blunt and hardly raised. Transverse sculptural septa on T5: about as strong as the longitudinal septa. T5 setae: directed posteriorly, arising from anterior edge of sculptural mesh. T6: as broad or broader than long. T6 laterotergite: overlapped by rim from S6. S4 median carina: absent. Transverse sculptural septa on S5: weak or absent, much weaker than the longitudinal septa. S5 setae: directed posteriorly, arising from anterior edge of sculptural mesh. Lateral carinae of S6: forming complete peripheral carina. Apex of S6: without notch.
Latin participle meaning “tricky,” named for its strong similarity to some other species and its very subtle metasomal bend.
Holotype, female: AUSTRALIA: WA, 20km N Denmark, 16.I.1987, J. S. Noyes, OSUC 238541 (deposited in WAMP). Paratypes: AUSTRALIA: 3 females, 3 males, OSUC 148699, 238192, 238195, 238197, 238199-238200 (CNCI).
Ocular setae: long and sparse. Frontal depression: with many irregularly transverse rugae. Smooth depression extending dorsolaterally from antennal foramen: present. Dorsal clypeal margin: arched, interrupted by broad median carina. Clypeal median carina: present. Ventral clypeal margin: with a small median point. Mandibular color: dark basally and at teeth, becoming lighter reddish brown between these areas. Mandibular teeth: three, but middle tooth tiny. Smooth area obliquely posterior to lateral ocellus: present. Genal sculpture: deeply reticulate-rugose with some septa much stronger than others, forming distinct rows differing in height.
Dorsal pronotal area: not set off by carina ventrally. Anterolateral corner of dorsal pronotal area: truncate anteriorly. Sculpture of posteromedian area of mesoscutum: densely foveolate. Lateral margin of dorsal axillar area: with a semicircular expansion, broadest near midlength. Mesoscutellar sculpture: densely foveolate. Metascutellum in dorsal view: trapezoidal with broad apex, or elongate-trapezoidal but with incised apex. Dorsal surface of metascutellum: apex protruding dorsally. Femoral depression: crossed by 4-6 dorsal carinae, 7-8 ventral carinae interrupted by central smooth area. Leg color: yellowish-brown except for dorsal external part of coxae and sometimes tarsomeres 2-5. Anterior corner of lateral propodeal carina: flat, without tooth. Posteromedial corner of lateral propodeal area: protruding posteriorly.
Metasoma color: black to dark reddish brown. Median lobe of T1: with 7 or more longitudinal carinae. Metasoma at middle of T4: with metasomal bend and abrupt transition in sculpture. Posterolateral margins of metasomal terga: without protrusions. T5 median carina: absent. Longitudinal sculptural septa on T5: strong, sharply raised. Transverse sculptural septa on T5: about as strong as the longitudinal septa. T5 setae: not directed posteriorly, arising from center of sculptural mesh. T6: as broad or broader than long. T6 laterotergite: overlapping S6. S4 median carina: absent. Transverse sculptural septa on S5: weak or absent, much weaker than the longitudinal septa. S5 setae: not directed posteriorly, arising from center of sculptural mesh. Lateral carinae of S6: not meeting apically. Apex of S6: with notch.
Latin adjective, named for the usually 3-pronged pattern formed apically by dark cuticle on T6.
Holotype, female: AUSTRALIA: SA, 32km N Renmark, Amalia Dam, xeric mallee scrub, MT 4, ROM 2000041, Bookmark Biosphere Reserve,
We thank L. Musetti, J. Jennings, J. Cora, A. Valerio, S. Mantel, D. Stringer, and other members of the PBI team, for assistance with specimens, photographs, and databasing. This work was funded by the National Science Foundation under grant No. DEB–0614764 to N.F. Johnson and A.D. Austin, and by the Australian Biological Resources Study grant 208-058 to A.D. Austin.
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1. entirely dark
2. entirely reddish
3. mostly dark, extreme base of scape becoming reddish
4. reddish-brown, darker at scape apex, pedicel, and A3
5. mostly dark, ventral parts of pedicel apex, A4-A12 variably lighter
6. yellowish-brown
7. dark except for pedicel apex
8. claval segments brown, basal segments yellowish
1. same as scape
2. base lighter than scape
3. lighter than scape
1. 6
2. 7
1. 1
2. 2
3. 0
1. short and dense
2. long and sparse
3. short and sparse
4. long and dense
5. absent
1. with transverse rugae
2. with large areoles
3. smooth or foveolate
4. with median longitudinal carina
5. inapplicable (frontal depression absent)
1. with transverse rugae
2. with large areoles
3. smooth or foveolate
4. with median longitudinal carina
5. inapplicable (frontal depression absent)
1. absent
2. present
1. absent between antennal foramina
2. interrupted by a median areole below interantennal process
3. interrupted by median carina
4. complete only between antennal foramina medially
5. wrinkle-like with median peak
6. angular, emarginate medially
7. uniformly arched without interruption
1. present
2. absent
1. straight
2. with a small median point
3. convex
4. concave
1. dark basally and at teeth, becoming lighter reddish brown between these areas
2. entirely dark
3. mostly yellowish brown, dark at teeth
4. mostly reddish brown, dark at teeth
1. three of roughly equal size
2. two, separated by narrow incision
3. three, but middle tooth tiny
4. two, broadly separated by an angled flat area
1. present
2. absent
1. reticulate-rugose with strong dorsoventral carinae
2. reticulate-rugose without any strong carinae
3. deeply reticulate-rugose with some septa much stronger than others, forming distinct rows differing in height
1. not set off by carina
2. set off by carina
1. protruding anteriorly
2. truncate anteriorly
3. weakly rounded anteriorly
1. sparsely foveolate, with large smooth interspaces
2. with many longitudinal carinae
3. densely foveolate
4. foveolate with broad longitudinal septa
5. foveolate with slightly stronger longitudinal septa
6. foveolate, with some interspersed additional tiny foveae
1. not expanded
2. broadly expanded
3. triangularly expanded or with posterior tooth, broadest posteriorly
4. with a semicircular expansion, broadest near midlength
1. densely foveolate
2. sparsely foveolate, with large smooth interspaces
3. densely foveolate with one or two median longitudinal channels
4. densely foveolate but with smooth central area
5. densely foveolate with slightly stronger longitudinal septa
6. foveolate, with some interspersed additional tiny foveae
7. densely foveolate with a longitudinal carina
8. medially smooth except for a few irregular carinae
1. strongly transverse, subrectangular
2. trapezoidal with broad apex
3. trapezoidal but tapering to narrow apex
4. elongate-trapezoidal but with incised apex
5. very short, subrectangular
6. semicircular
7. elongate-rectangular
1. flat
2. apex protruding dorsally
3. convex
1. crossed by rounded carinae
2. irregularly foveolate but not crossed by carinae
3. smooth, not crossed by carinae
4. crossed by 4-6 dorsal carinae, 7-8 ventral carinae interrupted by central smooth area
5. centrally smooth, peripherally foveolate
6. shallowly sculptured but not crossed by carinae
7. crossed by many sharply defined carinae
1. entirely dark
2. entirely reddish
3. coxae dark, all else reddish
4. coxae, femora (aside from their apices), and at least the last two tarsomeres dark, otherwise yellowish brown
5. coxae and at least the last three tarsomeres dark, otherwise reddish
6. reddish but with at least last two tarsomeres dark
7. dark except for 2nd trochanter of fore leg and mid leg, tip of mid and hind metatibia
8. reddish except for variable antero-dorsal portion of coxae
9. dark except for trochanters, tips of femora and tibiae, and tarsomeres 1-4
10. coxae and femora dark, but tibiae, tarsi, and sometimes 2nd trochanters yellowish brown
11. dark except for tips of tibiae and at least 1st tarsomere
12. yellowish-brown except for dorsal part of coxae and sometimes tarsomeres 2-5
13. dark except for tibiae, 2nd trochanter, and tarsomeres 2-5
14. dark, except trochanters, femoral apices, variable parts of tibiae, and tarsi pale
15. coxae, femora, and apical two tarsomeres dark, trochanters, tibiae, and other tarsomeres yellowish brown
16. coxae dark, leg becoming gradually lighter apically
17. coxae and at least the last three tarsomeres dark, otherwise yellowish
18. entirely yellowish
1. flat, without tooth
2. with longitudinal tooth-like crest extending dorsally above level of metascutellum
1. not protruding posteriorly
2. protruding posteriorly
1. black to dark reddish brown
2. entirely reddish
3. reddish, with last segment variably dark
4. reddish, with last two segments dark
5. reddish, with last 2.5 segments dark
6. T2 to flat part of T4, troughs of S1 to variable part of S5 reddish; otherwise dark
7. mostly reddish, variably dark at T4 bend, T5-T6, with variable smaller dark areas elsewhere
8. centrally reddish with some vague dark intrusions, peripherally becoming black
1. with 7 or more longitudinal carinae
2. with a set of rugae that merge with one another
3. with median smooth area interrupting carinae
4. with 6 carinae
5. with 5 carinae
1. with metasomal bend and abrupt transition in sculpture
2. without bend
3. with very weak bend
1. with tooth-like protrusions
2. without protrusions
1. absent
2. present
1. strong, sharply raised
2. weak, blunt and hardly raised
1. about as strong as the longitudinal septa
2. weak or absent, much weaker than the longitudinal septa
1. directed posteriorly, arising from anterior edge of sculptural mesh
2. not directed posteriorly, arising from center of sculptural mesh
1. longer than broad
2. as broad or broader than long
1. overlapped by rim from S6
2. overlapping S6
1. absent
2. present
1. weak or absent, much weaker than the longitudinal septa
2. about as strong as the longitudinal septa
1. directed posteriorly, arising from anterior edge of sculptural mesh
2. not directed posteriorly, arising from center of sculptural mesh
1. forming complete peripheral carina
2. not meeting apically
3. absent
4. weak and ruga-like, becoming indistinct posteriorly
1. without notch
2. with notch
3. roundly concave
1. A3 over 1.5× as long as broad, most others as long or longer than broad
2. A3 1–1.2× as long as broad, most others transverse
1. arched and posteriorly concave
2. flat and posteriorly truncate
Chromoteleia sp. 7002444010221101223052200H02111000100020011
Bracalba_cuneata 50110[23][23]16111000[01]132[04]11001B0[02]101001100020011
Bracalba_globosa 02110[23][23]161110100242111001C01101001100020011
Bracalba_plana 4211000101110200222150401F00101001001020111
Bracalba_clavata 41020[03][02]13112020023230[02]400300200001110020001
Bracalba_hesperia 00102021401001002232124018000[01]1000000120?1?
Bracalba_intermedia 0202022061220100242400401301101000110020011
Bracalba_laminata ??11302101120200143631001???????????????0?0
Bracalba_magnirubra 221012211012010002303[12]001520010110110111000
Bracalba_nigrescens ????300101120230203131001G??????????????0?0
Bracalba_parvirubra 201012012012020004323[02]601B70000001010000?0?
Bracalba_pinnula 021[01]00010110020022301[02]401901001001111020011
Bracalba_sculptifrons 00110[13][13]150120000123011401450000110110110?1?
Bracalba_propodealis 0110022101120200233142111300001000111120?1?
Bracalba_tricorata 0010200101120220243112001301201001000000010
Bracalba_sparsa 3210122131110100123021001160000111110111000
Bracalba_tridentata 00101001201202001230[13]1301B00000011100111?1?
Taxonomic records for all records used in the present paper. (doi:
Locality records for all records used in the present paper. (doi: