Corresponding author: Stjepan Krčmar (
Academic editor: D. Apanaskevich
The present paper is based on original and literature data. In Croatia the first studies on the occurrence of ixodid species were made about 80 years ago. The number of tick species recorded in Croatia considerably increased during the 1950s, 60s, 70s and 80s of the past century. A total of 21 species of hard tick belonging to 5 genera have been recorded in Croatia.
Hard ticks (Acari:
The following lists of species include all available literary records and new, still unpublished data. The samplings of ticks in Slavonia and Baranja were carried out during 2011 from the beginning of March to mid-September. Ticks were sampled by the flagging method, and were picked by hand from domestic animals and humans. The flagging method was used for tick sampling in two different vegetation types around the village of Mikleuš (YL 15). A 1 m2 white flag was dragged over the vegetation for about 50 m at each locality. The first locality was in the mixed forests of durmast oak and hornbeam (as.
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Sampling sites are identified by a running number referring to the map in
Sampling sites of hard ticks (
List of sampling sites
|
|
|
|
---|---|---|---|
1. | Dubrovnik<br/> Močiljska špilja<br/> Lokrum Island | BN 62 | |
2. | Kapelački Lug |
|
BR 87 |
3. | Đakovo |
|
BR 92 |
4. | Zmajevac |
|
CR 37 |
5. | Vrtlac reef |
|
VJ 88 |
6. | Baričevac Island<br/> Golac Island<br/> Mežanj Island<br/> Sparušnjak Island | VJ 98 | |
7. | Pula |
|
VK 16 |
8. | Unije Island |
|
VK 34 |
9. | Susak Island<br/> Vele Srakane Island | VK 42 | |
10. | Trstenik Island<br/> Školjić reef | VK 54 | |
11. | Cres Island: Belej<br/> Hrasta<br/> Zeča Island | VK 56 | |
12. | Cres Island: Cres<br/> Orlec<br/> Vrana | VK 57 | |
13. | Cres Island: Petrićevi- Špilja Čampari<br/> Vodice | VK 58 | |
14. | Kamenjak Island<br/> Lutrošnjak Island<br/> Premuda Island<br/> Južni Greben Island<br/> Zapadni Greben Island | VK 61 | |
15. | Grujica Island<br/> Ilovik Island<br/> Male Orjule Island<br/> Sveti Petar Island | VK 62 | |
16. | Lošinj Island: Veli Lošinj,<br/> Oruda Island,<br/> Palacol Island<br/> Ćunski-Like | VK 63 | |
17. | Morovnik Island<br/> Silba Island<br/> Kurjak reef | VK 71 | |
18. | Krk Island: Kozlja |
|
VK 79 |
19. | Dužac Island |
|
VK 80 |
20. | Olib Island<br/> Planičić reef<br/> Šip reef | VK 81 | |
21. | Mali Dolfin reef<br/> Mali Laganj Island | VK 93 | |
22. | Velebit mountain:<br/> Donja Klada<br/> Veliki Zavižan | VK 96 | |
23. | Lukovdol- Lipov Vrh |
|
VL 03 |
24. | Buzet-Rabakova Peć<br/> Ročko Polje | VL 22 | |
25. | Istra: Učka mountain |
|
VL 31 |
26. | Rijeka |
|
VL 51 |
27. | Gorski Kotar: Risnjak mountain |
|
VL 63 |
28. | Sungerski Lug |
|
VL 81 |
29. | Kamik od Tramuntane<br/> Palagruža Island | XG 09 | |
30. | Brač Island: Blato<br/> Nadsela<br/> Osridke<br/> Sveti Toma | XH 49 | |
31. | Brusnik Island<br/> Pod Mrčaru reef | XH 43 | |
32. | Brač Island:<br/> Zapadna Laščatna<br/> Sjeverna Laščatna | <br/> |
XH 59 |
33. | Mljet Island: Soline<br/> Veliko Jezero | XH 93 | |
34. | Baćinska jezera |
|
XH 97 |
35. | Kaštela, Kozjak mountain:<br/> Malačka | XJ 02 | |
36. | Split |
|
XJ 11 |
37. | Kaštela Bay |
|
XJ 12 |
38. | Brač Island: Supetar |
|
XJ 20 |
39. | Brač Island |
|
XJ 30 |
40. | Sinj |
|
XJ 34 |
41. | Srednja Posavina: Stružec |
|
XL 24 |
42. | Repušnica |
|
XL 33 |
43. | Križevci<br/> Stara Ves | XL 39 | |
44. | Nedelišće |
|
XM 12 |
45. | Čakovec |
|
XM 13 |
46. | Dekanovec |
|
XM 24 |
47. | Maslinovac Island |
|
WJ 07 |
48. | Mali Prišnjak<br/> Malo Šilo Island<br/> Mrtovac Island | WJ 16 | |
49. | Božikovac Island<br/> Mala Lavdara Island<br/> Mala Skala Island<br/> Rončić Island<br/> Trimulić Island<br/> Trstikovac Island<br/> Vela Skala Island | WJ 17 | |
50. | Kali<br/> Vela Sestrica<br/> Zadar | WJ 18 | |
51. | Veseljuh Island |
|
WJ 25 |
52. | Gušteranski Island<br/> Koromašna Island<br/> Kosmerka Island<br/> Škrovada reef | WJ 26 | |
53. | Dugi otok Island<br/> Pašman Island | WJ 27 | |
54. | Bisaga Island<br/> Mali Babuljaš Island<br/> Veli Babuljaš Island | WJ 35 | |
55. | Biograd-Vrana |
|
WJ 36 |
56. | Babina Guzica Island<br/> Jančar Island<br/> Kameni Puh Island<br/> Mala Prduša Island<br/> Purara Island<br/> Veli Puh Island<br/> Vodeni Puh Island<br/> Vodenjak Island | WJ 43 | |
57. | Rovanjska |
|
WJ 49 |
58. | Čavlin Island<br/> Čerigul Island<br/> Mali Kamešnjak Island<br/> Mali Tetovišnjak Island<br/> Mikavica Island<br/> Samograd Island<br/> Veli Kamešnjak Island<br/> Veli Tetovišnjak Island<br/> Žirje Island | WJ 53 | |
59. | Velebit mountain: Crni Padež |
|
WK 04 |
60. | Velebit mountain: Baške Oštarije |
|
WK 12 |
61. | Velika Kapela mountain: Razvala |
|
WK 29 |
62. | Lika: Plitvice-Crna rijeka |
|
WK 46 |
63. | Lika: Plitvice-Mukinje |
|
WK 56 |
64. | Velebit mountain: Prezid, Pod ćelavcem |
|
WK 60 |
65. | Zagreb |
|
WL 77 |
66. | Zagreb- vicinity, Medvednica mountain: Sljeme |
|
WL 78 |
67. | Vukomeričke Gorice: Prkovec |
|
WL 84 |
68. | Petrinja |
|
WL 93 |
69. | Peščenica-Vratovo |
|
WL 95 |
70. | Dugo Selo |
|
WL 97 |
71. | Banjol Island<br/> Figarola Island<br/> Gustinja Island<br/> Pisulj Island<br/> Pulari Island<br/> Revera Island<br/> Rovinj<br/> Sturag Island<br/> Sveti Ivan Island<br/> Sveti Ivan np reef<br/> Velika Sestrica Island<br/> Veliki Piruzi reef<br/> Veštar Island | UK 99 | |
72. | Umag |
|
UL 83 |
73. | Koversada<br/> Lakal Island<br/> Lunga Island<br/> Tovarjež Island | UL 90 | |
74. | Mljet Island:<br/> Babino Polje<br/> Blato<br/> Zadublje | <br/> |
YH 03 |
75. | Metković |
|
YH 06 |
76. | Mljet Island |
|
YH 13 |
77. | Metković-Gabela |
|
YH 16 |
78. | Slano |
|
YH 33 |
79. | Mikleuš |
|
YL 15 |
The tick fauna of Croatia is now known to include 21 species. These species are classified into five genera:
For every species are mentioned, sex, developmental stage ( ♂ male, ♀ female N = nymph, L = larva), sampling place, month or date, host and data source.
♂♂,♀ ♀ Cres Island: Hrasta 10.XI.1948, from
♂♂,♀ ♀ ibidem: Vodice 6.–11.IV. 1948, from
♂♂,♀♀ ibidem: Hrasta 29.IX.–1.X.1948, from
♂♂,♀♀ ibidem: Belej 29.IX.–1.X.1948, 4.XI. 1948, from
♂♂, ♀♀ Pula, months: I. II. III. IV.V. IX. X. XI. XII. 1936–1963 from
♂♂, ♀♀ Krk Island, months: I. II. IX. X. XI. XII. 1936–1963 from
♂♂, ♀♀ Cres Island, months: I. II. III. IV. VIII. IX. X. XI. XII. 1936–1963 from
♂♂, ♀♀ Lošinj Island: Veli Lošinj, months: I. II. III. IV. VIII. IX. X. XII. 1936–1963 from
♂♂,♀♀ ibidem: Čunski-Like, month: VIII. 1948, from
♂♂,♀ ♀ Zadar, months: I. II. III. IV.V.VI. IX. X. XI. XII. 1936–1963 from
♂♂, ♀♀ Kaštela, month: VI. 1936–1963 from
♂♂, ♀♀ Split, months: I. II. III. IV.V.VI. XI. XII. 1936–1963 from
♂♂, ♀♀ Brač Island, months: I. II. III. IX. X. XI. XII. 1936–1963 from
♂♂, ♀♀ Dubrovnik, months: II. III. IV. IX. X. 1936–1963 from
♂♂, ♀♀ Srednja Posavina, months: III. IV. V. VI. XI. XII. from
♂♂,♀♀ Brač Island: Sveti Toma 4.–9.XI. 1965 from
♂♂,♀♀ ibidem: Osridke 4.–9.XI.1965 from
♂♂,♀♀ ibidem: Blato 4.–9.XI. 1965 from
♂♂,♀♀ ibidem: months: I. II. X. XI. XII. 1964/1965 collected on vegetation using the cloth dragging method (
♂♂,♀♀ ibidem: Sveti Toma, months: I. II. XI. XII. 1964/1965 collected on vegetation using the cloth dragging method (
♂♂, ♀♀ ibidem: Zapadna Laščatna, months: I. X. XI. XII.1964/1965 collected on vegetation and/or the ground by flagging/dragging method (
♂♂, ♀♀ ibidem: Sjeverna Laščatna, months: XI. XII. 1964/1965 collected on vegetation using the cloth dragging method (
♂♂, ♀♀ ibidem: months: I. II. III. IX. X. XI. XII. 1964/1965 from
NN ibidem: month:VII. 1964/1965 from
♂♂, ♀♀, NN, LL Srednja Posavina: Stružec from 1970 to 1978 collected on vegetation using the cloth dragging method (
NN Peščenica-Vratovo 17.VII. 1974 from
NN Velebit mountain: Predzid (pod Ćelavcem) 1.VIII. 1975 from
♂♂,♀♀ Brač Island: Supetar 22.II. 1974 from
L, NN Repušnica 5.,6.VIII. 1982 from
N ibidem: 6.,7.VIII. 1982 from
♂♂,♀♀ Čakovec, months: III. IV. V. 2007 collected on vegetation using the cloth-dragging method (
♂♂, ♀♀ Kaštela Bay, month: X.2000 from
♂♂,♀♀ Sinj, month: X. 2000, from
♂♂,♀♀ Brač Island, Hvar Island (
♂♂, ♀♀ Đakovo 10.III.–30.III. 2011 from
♂♂, ♀♀ ibidem: 7.IV.–25.IV. 2011 from
♂♂, ♀♀ ibidem: 2.V.–25.V.2011 from
♂, ♀ Zmajevac 15.III.- 30.III. 2011 from
♀♀ ibidem: 10.IV.–24.IV. 2011 from
♀♀ ibidem: 25.IV.–30.IV. 2011 from
♀ ibidem: 7.V.–25.V.2011 from
♂♂, ♀♀ Gornja Posavina, months: III. IV. V. from
♀ Kapelački Lug, month: V. 1967 collected on vegetation using the cloth dragging method (
♂♂, ♀♀, NN, LL Srednja Posavina: Stružec -, from 1970 to 1978 collected on vegetation using the cloth dragging method (
♂♂, ♀♀ Čakovec, months: III. IV. V. 2007 collected on vegetation using the cloth dragging method (
♀♀ Mikleuš 27.IV.2011 collected on vegetation using the cloth dragging method
♀♀ ibidem: 14.V.2011 collected on vegetation using the cloth dragging method
♂♂, ♀♀ Cres Island: Hrasta 6.–11.IV.1948 from
♂♂, ♀♀ ibidem: 29.IX.–1.X. 1948 from
♂♂, ♀♀ ibidem: 10.XI. 1948 from
♂♂, ♀♀ ibidem: Vrana 6.–11.IV.1948 from
♂♂, ♀♀, N ibidem: Vodice 6.–11.IV.1948 from
♂♂, ♀♀ ibidem: Belej 29.IX.–1.X.1948 from
♂♂,♀♀ ibidem: 4.XI. 1948 from
♂♂, ♀♀ Lošinj Island: Veli Lošinj 29.IX.–1.X. 1948 from
♂♂, ♀♀ ibidem: Čunski-Like 29.IX.–1.X. 1948 from
♂♂, ♀♀, NN, LL Pula,months: I. II. , III. , IV. V. VI. , VII. , VIII. IX. , X. XI. XII. 1936–1963 from
♂♂, ♀♀ Krk Island,months: I. II. IV. IX. X. XI. XII. 1936–1963 from
♂♂, ♀♀ Cres Island,months: I. II. III. IV.V. IX. X. XI. XII. 1936–1963 from
♂♂, ♀♀ Lošinj Island: Veli Lošinj, months: I. II. III. IV. IX. X. XI. XII. 1936–1963 from
♂♂, ♀♀ Zadar,months: I. II. III. IV. V. VI. IX. X. XI. XII. 1936–1963 from
♂♂,♀♀, NN, LL Split,months: I. II. III. IV. X. XI. XII. 1936–1963 from
♂♂, ♀♀ Brač Island, months: II. III. VII. X. XI. 1936–1963 from
♂♂,♀♀ ibidem: months: I. II.III. IV. V. IX. X. XI. XII 1964/1965 from
♂♂, ♀♀, NN Dubrovnik,months: I. II. III. IV.V. IX. X. XI. XII.1936–1963 from
♂♂, ♀♀ Gornja Posavina,month: V. from
♂♂, ♀♀ Brač Island: Sveti Toma 4–9.XI.1965 from
♂♂, ♀♀ ibidem: Osridke 4.–9.XI. 1965 from
♂♂, ♀♀ ibidem: Blato 4.–9.XI. 1965 from
♂♂, ♀♀ ibidem: months: I. II. IV.V. XII. 1964/1965 collected on vegetation and/or the ground by flagging/dragging method (
♂♂, ♀♀ ibidem: Sveti Toma, months: I. II. IV.V. IX. X. XI. XII. 1964/1965 from
♂♂, ♀♀ ibidem: Blato 5.XI.1965 from
♂♂, ♀♀ ibidem: Zapadna Laščatna, months: I. IV. IX. X. XI. XII. 1964/1965 collected on vegetation and/or the ground by flagging/dragging method (
♂♂, ♀♀ ibidem: Sjeverna Laščatna, months: I. IV. XI. XII. 1964/1965 collected on vegetation and/or the ground by flagging/dragging method (
N Metković 23.IV. 1964 from
♂♂, ♀♀ Brač Island: Supetar 22.II. 1974 from
♂♂, ♀♀ Kaštela Bay, month: X.2000 from
♂♂, ♀♀ Sinj, month: X. 2000 from
♂♂, ♀♀ Brač Island, Hvar Island (
♂♂, ♀♀ Cres Island, months: I. II.1936–1963 from
♂♂, ♀♀ Lošinj Island: Veli Lošinj, month: IX. 1936–1963 from
♂♂, ♀♀ Zadar, month: V. 1936–1963 from
♂♂, ♀♀ Dubrovnik, month: V. 1936–1963 from
♂ Dekanovec 10.VI. 1977 from
NN Krk Island: 15.V.2002 from
♂♂ Mikleuš 14.V.2011 collected on vegetation using the cloth-dragging method
NN ibidem: 8.VI.2011 collected on vegetation using the cloth-dragging method
♀, NN ibidem: 9.VI.2011 collected on vegetation using the cloth-dragging method
♂ ibidem: 15.VI.2011 collected on vegetation using the cloth-dragging method
NN, LL ibidem: 3.VII.2011 collected on vegetation using the cloth-dragging method
♂, NN ibidem: 4.VII.2011 collected on vegetation using the cloth-dragging method
NN, LL ibidem: 2.VIII.2011 collected on vegetation using the cloth-dragging method
NN, LL ibidem: 3.VIII.2011 collected on vegetation using the cloth-dragging method
N ibidem: 13.IX.2011 collected on vegetation using the cloth-dragging method
♂♂, ♀♀ Pula, months: X. XI. 1936–1963 from
♂♂, ♀♀ Krk Island, months: IV. XI. XII. 1936–1963 from
♂♂, ♀♀ Cres Island, months: I. II. III. IV. V. X. XI. XII. 1936–1963 from
♂♂, ♀♀ Lošinj Island: Veli Lošinj, months: I. IV. X. 1936–1963 from
♂♂, ♀♀ Zadar, months: I. II. III. IV. XI. XII. 1936–1963 from
♂♂, ♀♀ Split, months: I. II. III. IV. 1936–1963 from
♂♂, ♀♀ Brač Island, months: I. II. III. IV. V. X. XI. XII. 1964/1965 from
NN ibidem: months: VII. VIII. 1964/1965 from
♀ ibidem: Nadsela 1.X.1965 from
L Krk Island: 23.VII.1956 from
L ibidem: 23.VII.1956 from
N Cres Island: Vrana, month: VII.1949 from
NN, LL Trstenik Island: 20.VII.1965 from
NN Mali Dolfin reef: 20.IX.1961 from
NN, LL Lutrošnjak Island: 25.VII.1965 from
N, LL Kamenjak Island: 25.VII.1965 from
NN, LL Premuda Island: 1.VIII.1965 from
LL Zapadni Greben Island: 23.VII.1965 from
LL Južni Greben Island: 23.VII.1965 from
NN, LL Silba Island: 13.VIII.1963 from
NN, LL ibidem: 22.VII.1965 from
LL ibidem: 2.VIII.1965 from
NN, LL Morovnik Island: 24.VII.1965 from
NN, LL Olib Island: 3.VIII.1965 from
LL Planičić reef: 3.VIII.1965 from
LL Šip reef: 24.VII.1965 from
L Vela Sestrica Island: 11.VIII.1963 from
N, LL Dužac Island: 11.VIII.1963 from
NN, LL Golac Island: 9.VIII.1965 from
NN Mali Laganj Island: 12.VIII.1963 from
NN, LL Baričevac Island: 6.VIII.1965 from
NN, LL Mežanj Island: 8.VIII.1965 from
LL Kurjak reef: 24.VII. 1965 from
NN, LL Sparušnjak Island: 7.VIII.1965 from
NN, LL Maslinovac Island: 7.VIII.1963 from
NN, LL Vrtlac Island: 8.VIII.1963 from
LL Veseljuh Island: 28.VII.1964 from
NN, LL Bisaga Island: 7.VIII.1964 from
N, LL Mali Tetovišnjak Island: 22.VII.1964 from
LL Čerigul Island: 22.VII.1964 from
LL Mali Kamešnjak Island: 23.VII.1964 from
NN, LL Samograd Island: 6.VIII.1963 from
LL Koromašna Island: 20.VII.1964 from
NN, LL Gušteranski Island: 23.VII.1964 from
N, LL Palagruža Island: 8.VI.1960 from
LL Kamik od Tramuntane Island: 9.VI.1960 from
LL Brusnik Island: 7.VI.1960 from
NN Pod Mrčaru reef: 31.VII.1971 from
NN, LL Božikovac Island: 1.VIII.1964 from
N, LL Mala Lavdara Island: 1.VIII.1964 from
NN, LL Trimulići Island: 2.VIII.1964 from
NN, LL Trstikovac Island: 2.VIII.1964 from
LL Vela Skala Island: 2.VIII.1964 from
NN, LL Mala Skala Island: 2.VIII.1964 from
NN, LL Rončić Island: 1.VIII.1964 from
NN, LL Mali Babuljaš Island: 7.VIII.1963 from
NN, LL Malo Šilo Island: 6.VIII.1964 from
NN, LL Mrtovac Island: 6.VIII.1964 from
NN, LL Veli Babuljaš Island:7.VIII.1964 from
NN, LL Mali Prišnjak Island: 7.VIII.1964 from
NN, LL Vodenjak Island: 7.VIII.1964 from
NN, LL Jančar Island: 7.VIII.1964 from
NN, LL Purara Island: 7.VIII.1964 from
NN, LL Mala Prduša Island: 7.VIII.1964 from
NN, LL Babina Guzica Island: 8.VIII.1964 from
LL Veli Puh Island: 21.VII.1964 from
LL Vodeni Puh Island: 21.VII.1964 from
NN, LL Kameni Puh Island: 21.VII.1964 from
LL Čavlin Island: 22.VII.1964 from
LL Veli Tetovišnjak Island: 22.VII.1964 from
LL Veli Kamešnjak Island: 23.VII.1964 from
N, LL Žirje Island: 27.VII.1964 from
N, L Mikavica Island: 4.VIII.1964 from
LL Škrovada reef: 23.VII.1964 from
NN, LL Kosmerka Island: 4.VIII.1964 from
L Pod Mrčaru reef: 31.VII.1971 from
♂♂, ♀♀ Southern Croatia from
N Dubrovnik 10.VIII. 1964 from
♂♂, ♀♀ Brač Island: (
♂♂, ♀♀ Pula, months: III. IV. IX. 1936–1963 from
♂♂, ♀♀ Krk Island, months: II. III. IV. 1936–1963 from
♂♂, ♀♀ Lošinj Island: Veli Lošinj, month: IV. 1936–1963 from
♂♂, ♀♀ Zadar, months: II. III. IV. VI. VIII. 1936–1963 from
NN ibidem: months: IX. XI. 1936–1963 from
♂♂, ♀♀ Split, months: I. II. III. IV. VI. 1936–1963 from
NN ibidem: months: XI. XII. 1936–1963 from
♂♂, ♀♀ Brač Island, months:V.VI. 1936–1963 from
♂♂, ♀♀ ibidem: 1964/1965 from
♂♂, ♀♀ ibidem: -, (
♂♂, ♀♀, NN Dubrovnik, month: I. 1936–1963 from
♂♂, ♀♀ ibidem: months: II. III. IV.V. IX. XI. XII.1936–1963 from
♂♂, ♀♀ Pula, months: I. II. III. IV. V. VI. VII. VIII. IX. X. XI. XII. 1936–1963 from
♂♂, ♀♀ Krk Island, months: II. III. IV. V. VI. VII. VIII. IX. X. 1936–1963 from
♂♂, ♀♀ Cres Island, months: IV. VI. VII. X. XII. 1936–1963 from
♂♂, ♀♀ Lošinj Island: Veli Lošinj, months: II. III. IV. V. VI. VII. 1936–1963 from
♂♂, ♀♀ Zadar, months: II. III. IV. V. VI. VII. VIII. X. 1936–1963 from
♂♂, ♀♀ Split, months: II. III. IV. V. VI. VII. VIII. IX. X. XII. 1936–1963 from
♂♂, ♀♀ Dubrovnik, months: III. IV. V. VI. VII. VIII. IX. XII. 1936–1963 from
♂, ♀ Petrinja, month: V. from
♂♂, ♀♀ Brač Island, months: I. II. III. IV. V. VI. VII. VIII. IX. X. 1936–1963 from
♂♂, ♀♀ ibidem: months: V. VI. VII. VIII. IX. X. 1964/1965 from
♂, ♀ ibidem: Blato, month: IV.1964/1965 from
♂♂, ♀♀ ibidem: Zapadna Laščatna, months: IV. VII.1964/1965 collected on vegetation and/or the ground by flagging/dragging method (
♂♂, ♀♀ ibidem: (
NN, LL ibidem: month: VII. 1967 from
♂♂, ♀♀ Kaštela Bay, month: X. 2000 from
♂♂, ♀♀ Sinj, month: X. 2000 from
♂♂, ♀♀ Lošinj Island: Veli Lošinj 29.IX.–1.X.1948 from
♂♂, ♀♀ ibidem: months: IV. V. X. 1936–1963 from
♂♂, ♀♀ Zadar, months: III. IV.V. VI. VII. VIII. IX. 1936–1963 from
NN ibidem: months: IV. VII. VIII. IX. 1936–1963 from
♂♂, ♀♀ Split, months: VI. VII. VIII. IX. X. XI. 1936–1963 from
NN ibidem: months: IX. XI. 1936–1963 from
♂♂, ♀♀ Brač Island: month: VII. 1936–1963 from
♂♂, ♀♀ ibidem: (
♂♂, ♀♀ Dubrovnik, months: III. IV. V. VII. 1936–1963 from
NN ibidem: months: IV.XI. 1936–1963 from
♂♂, ♀♀ Lošinj Island: Čunski-Like 6.–11.IV.1948 from
♂♂, ♀♀ Cres Island, months:, IV. V. 1948 from
♂♂, ♀♀ ibidem: month: VIII. 1948 from
LL ibidem: 29.IX.–1.X.1948 from
♂♂, ♀♀, NN, LL ibidem: Hrasta 6.–11.IV. 1948 from
♂♂, ♀♀, NN, LL ibidem: 29.IX.–1.X.1948 from
NN, L ibidem: 10.XI.1948 from
NN ibidem: Vrana 6.–11.IV. 1948 from
♂♂, ♀♀, NN ibidem: Vodice 6.–11.IV. 1948 from
♂♂, ♀♀ ibidem: Orlec 6.–11.IV. 1948 from
♂♂, ♀♀ ibidem: Belej 29.IX.–1.X.1948 from
NN ibidem: 4.XI.1948 from
♂♂, ♀♀ Pula, months: II. III. IV. V. VI. VIII. 1936–1963 from
NN ibidem: months: I. II. III. IV. X. XI. XII. 1936–1963 from
LL ibidem: months: III. IV. XI. 1936–1963 from
♂♂, ♀♀ Krk Island, months: II. III. IV. V. VI. VII.VIII. X. 1936–1963 from
NN ibidem: months: I. III. IV.VI. X. XI. XII.1936–1963 from
LL ibidem: months: III. IV. X. 1936–1963 from
♂♂, ♀♀ Cres Island: Cres, months: III. IV. V. VI. VII. IX. 1936–1963 from
NN ibidem: months: I. II. III. IV. V. XI. 1936–1963 from
LL ibidem: months: I. III. XI. 1936–1963 from
♂♂, ♀♀ Lošinj Island: Veli Lošinj, months: III. IV. V. VI. VII. VIII. IX. 1936–1963 from
NN ibidem: months: II. III. IV.X. XI. XII. 1936–1963 from
LL ibidem: months: III. IV. IX. XI. 1936–1963 from
♂♂, ♀♀ Zadar, months: III. IV. V. VI.VII. VIII. 1936–1963 from
NN ibidem: months: I. II. III. IV. V. XII. 1936–1963 from
LL ibidem: month: I. III. IV. 1936–1963 from
♂♂, ♀♀ Split, months: II. III. V. VI. VII. VIII. IX. 1936–1963 from
NN ibidem: months; II. III. X. XII. 1936–1963 from
LL ibidem: month: III. 1936–1963 from
♂♂, ♀♀ Brač Island, months: II. III. IV. V.VI. VII.VIII. 1936–1963 from
NN ibidem: months: I. II. III. IV. X. XI. XII. 1936–1963 from
♂♂, ♀♀ ibidem: 1964/1965 from
♂♂, ♀♀ ibidem: (
NN ibidem: Blato 4.–9.XI. 1965 from
♂, ♀ ibidem: Blato, month: IX.1964/1965 collected on vegetation and/or the ground by flagging/dragging method (
♂♂, ♀♀ ibidem: Sveti Toma, months: IX. X. 1964/1965 collected on vegetation and/or the ground by flagging/dragging method (
♂♂, ♀♀ ibidem: Zapadna Laščatna, months: VII. VIII. 1964/1965 collected on vegetation and/or the ground by flagging/dragging method (
♂♂, ♀♀ Dubrovnik, months: III. IV.V. VI.VII. VIII. IX. 1936–1963 from
NN ibidem: months: II. IV. V. X. XI. XII. 1936–1963 from
♂♂, ♀♀ Kaštela Bay, month: X. 2000 from
♂♂, ♀♀ Sinj, month: X. 2000 from
♂♂, ♀♀ Mali Lošinj Island: Čunski-Like 6.–11.IV. 1948 -, V. 1948 from
♂♂, ♀♀ Cres Island: Hrasta 6.–11.IV. 1948 from
♂♂, ♀♀ ibidem: Belej 6.–11.IV. 1948 from
♂♂, ♀♀ ibidem: Vrana 6.–11.IV. 1948 from
♂♂, ♀♀ ibidem: Vodice 6.–11.IV. 1948 from
♂♂, ♀♀ ibidem: Orlec. months: IV. V. 1948 from
♂♂, ♀♀ ibidem: Cres, months: IV. V. VII. 1936–1963 from
♂♂, ♀♀ Pula, months: V.VI. 1936–1963 from
♂ ibidem: 18.V.1973 from
♂♂, ♀♀ Zadar, months: III. IV. V. VI. 1936–1963 from
♂♂, ♀♀ Split, months: III. IV. V. VI. 1936–1963 from
♂♂, ♀♀ Dubrovnik, months: III. IV. V. VI. VII. VIII. 1936–1963 from
♂♂, ♀♀ Brač Island, months: II. III. IV. 1936–1963 from
♂♂, ♀♀ ibidem: from 1964/1965 from
♂♂, ♀♀ ibidem: Blato, months; IV. V. VI. 1964/1965 collected on vegetation and/or the ground by dragging/flagging method (
♂♂, ♀♀ ibidem: Sveti Toma, months: IV. V. VI. VII. 1964/1965 collected on vegetation and/or the ground by dragging/flagging method (
♂♂, ♀♀ ibidem: Zapadna Laščatna, months: IV. VI. VII.1964/1965 collected on vegetation and/or the ground by dragging/flagging method (
♂♂, ♀♀ ibidem: Sjeverna Laščatna, months: IV. VI. 1964/1965 collected on vegetation and/or the ground by dragging/flagging method (
♂♂, ♀♀ ibidem: 1.–4.V. 1964 and 28.III.–1.IV.1965 collected on vegetation by dragging method (
♂♂, ♀♀ ibidem: 28.III. , 27.IV., 1.VI., 4.VI. 1965 from
♂♂, ♀♀ ibidem: (
♂♂, ♀♀ Umag 27.V.1978 from
♀ Slano (Dubrovnik) 10.V.1953 from
♂♂, ♀♀ Ugljan Island: Kali 9.–20.V.1955 from
♂♂, ♀♀ Rovanjska 23.IV.1958 from
♂♂, ♀♀ Zadar 25.IV.1958 from
♂♂ Pašman Island: 2.V.1985 from
♂,♀ Dugi otok Island: 29.IV.1986 from
♂♂, ♀♀ Mljet Island: Babino polje 8.–9.IV.1981, 16.–17.V. 1981 from
♂♂, ♀♀ ibidem: Zadublje 8.–9.IV.1981 from
♂♂, ♀♀ ibidem: Veliko jezero 8.–9.IV.1981 from
♂♂, ♀♀ ibidem: Soline 8.–9.IV.1981 from
♂♂, ♀♀ ibidem: Babino polje 16.–17.V. 1981 from
♂♂, ♀♀ ibidem: Blato 16.–17.V. 1981 from
♂♂, ♀♀ Ugljan Island: Kali 9.–20.V.1955 from
♂ Zadar 25.IV. 1958 from
♂ Lokrum Island: 9.IX.1963 from
♂ ibidem: 4.IX. 1963 from
♂♂, ♀ Brač Island: 1.–4.V.1964 and 28.III.–1.IV.1965 collected on vegetation by dragging method (
♂♂, ♀♀ ibidem: 28.III., 27.IV., 1.VI., 4.VI., 1965 from
♂♂, ♀♀ Pašman Island: 2.V.1985 from
♂♂, ♀ Dugi otok Island: 29.IV.1986 from
♂♂, ♀♀ Mljet Island: 8.–9.IV. 1981, 16–17.V. 1981 from
♂♂, ♀♀ ibidem: Babino polje 8.–9.IV.1981 from
♂♂, ♀♀ ibidem: Zadublje 8.–9.IV.1981 from
♂♂, ♀♀ ibidem: Veliko jezero 8.–9.IV.1981 from
♂♂, ♀♀ ibidem: Soline 8.–9.IV.1981 from
♂♂, ♀♀ ibidem: Babino polje 16.–17.V. 1981 from
♂♂, ♀♀ ibidem: Blato 16.–17.V. 1981 from
♀♀ Zagreb 29.IX. 1962 from
♀ Velebit mountain: Donja Klada, 1975 from
♀ Dubrovnik 22.II.1962 from
♀ ibidem: 1.XII. 1967 from
♀ Zagreb, vicinity, 20.IV. 1975 from
♂♂, ♀♀ Brač Island: Sveti Toma 4–9. XI. 1965 from
♂♂, ♀♀ ibidem: Osridke 4–9.XI.1965 from
♂♂, ♀♀ ibidem: Blato 4.–9.XI. 1965 from
♂♂, ♀♀, NN, LL ibidem: Zapadna Laščatna 24.–25.XI. 1964 collected on the vegetation by dragging method (
♂♂, ♀♀ ibidem: Blato, months: I. II. IV. V. XII. 1964/1965 collected on vegetation and/or the ground by dragging/flagging method (
♂♂, ♀♀ ibidem: Sveti Toma, months: I. II. IV. V. XI. XII. 1964/1965 collected on vegetation and/or the ground by dragging/flagging method (
♂♂, ♀♀ ibidem: Zapadna Laščatna, months: I. IV. XI. XII. 1964/1965 collected on vegetation and/or the ground by dragging/flagging method (
♂♂, ♀♀ ibidem: Sjeverna Laščatna, months: I. IV. XI. XII. 1964/1965 collected on vegetation and/or the ground by dragging/flagging method (
♂♂, ♀♀ ibidem: Supetar 22.II. 1974 from
♂♂, ♀♀ ibidem: (
♂♂, ♀♀ Gornja Posavina, month: III. from
NN Velika Kapela mountain: Razvala 16.VIII. 1969 from
♀ Dugo Selo 5.IV. 1959 from
♀♀ Đakovo 7.IV.–25.IV. 2011 from
♂♂ ibidem: 2.V.–25.V. 2011 from
♂♂, ♀♀, NN, LL Cres Island: Hrasta 6.–11.IV.1948 from
♂, ♀♀ ibidem: 10.XI.1948 from
♂♂, ♀♀, NN ibidem: Vrana 6.–11.IV.1948 from
♂♂, ♀♀, NN, LL ibidem: Vodice 6.–11.IV.1948 from
NN ibidem: Cres 6.–11.IV.1948 from
♂♂, ♀♀, NN ibidem: Belej 4.XI.1948 from
♂♂, ♀♀ Pula, months: I. II. III. IV. V. VI. VIII. IX. X. XI. XII. 1936–1963 from
NN ibidem, months: I. II. III. IV. X. XII. 1936–1963 from
LL ibidem, months: IV. VII. XII. 1936–1963 from
♂♂, ♀♀ Krk Island, months: I. II. III. IV. V. IX. XI. XII. 1936–1963 from
NN ibidem, months: I. II. III. IV. IX. X. 1936–1963 from
LL ibidem, months: I. II. III. IV. X. 1936–1963 from
♂♂, ♀♀ Cres Island, months: I. II. III. IV.V. IX. X. XI. XII. 1936–1963 from
NN ibidem, months: I. II. III. IV. IX. XI. 1936–1963 from
LL ibidem, months: II. III. 1936–1963 from
♂♂, ♀♀ Lošinj Island: Veli Lošinj, months: I. II. III. IV. V. IX. X. XI. XII. 1936–1963 from
NN ibidem: months: I. II. III. IV. V. IX. X. XI. 1936–1963 from
LL ibidem: months: II. III. IV. X. 1936–1963 from
♂♂, ♀♀ Zadar, months: I. II. III. IV. VI. IX. X. XI. XII. 1936–1963 from
NN ibidem: months: I. III. IV. IX. 1936–1963 from
LL ibidem: months: I. IV. IX. 1936–1963 from
♂♂, ♀♀ Split, months: I. II. III. IV. X. XI. XII. 1936–1963 from
NN ibidem: months: I. II. III.1936–1963 from
LL ibidem: month: II. 1936–1963 from
♂♂, ♀♀ Brač Island, months: I. III. IX. X. XI. XII. 1936–1963 from
NN ibidem: months: I. II. III. 1936–1963 from
♂♂, ♀♀ Dubrovnik, months: I. II. III. IV.V. IX. X. XI. XII. 1936–1963 from
NN ibidem: months: I. II. IV. XI. XII.1936–1963 from
N, L Rovinj 7.IV.1959 from
N, L ibidem: 7.IV.1959 from
L ibidem: 10.IV. 1959 from
NN, LL Tovarjež Island: 7.V.1959 from
N, L Lakal Island: 7.V.1959 from
NN, LL Lunga Island: 6.IV.1959 from
NN, LL Koversada Island: 4.IV.1959 from
N, L Figarola Island: 9.IV.1959 from
N, L Banjol Island: 11.IV.1959 from
N Veliki Piruzi reef: 4.IV.1959 from
LL Sturag Island: 12.VIII.1959 from
LL Sveti Ivan Island: 12.VIII.1959 from
L Sveti Ivan na Pučini reef: 4.IV.1959 from
NN, LL Pulari Island: 8.V.1959 from
LL Revera Island: 8.V.1959 from
N, LL Veštar Island: 8.IV.1959 from
NN, LL Velika Sestrica Island: 8.IV.1959 from
NN, LL Pisulj Island: 8.V.1959 from
NN, LL Gustinja Island: 8.V.1959 from
NN, LL Unije Island: 17.VII.1960 from
NN, LL ibidem: 17.VII. 1960 from
N, LL Zeča Island: 18.VIII.1963 from
N, LL Vele Srakane Island: 19.VII.1960 from
LL Susak Island: 19.VII.1960 from
NN, LL Oruda Island: 19.VII.1960 from
NN, LL Palacol Island: 19.VII.1960 from
N, LL Male Orjule Island: 19.VII.1960 from
N, LL Školjić Island: 19.VII.1960 from
NN, LL Sveti Petar Island: 29.VII.1965 from
LL Ilovik Island: 29.VII.1965 from
LL Grujica Island: 28.VII.1965 from
♂♂, ♀♀, NN, LL Srednja Posavina: Stružec, 1970–1978 collected on vegetation and/or the ground by dragging/flagging method (
♂♂, ♀♀ Gornja Posavina, months: III. IV. V. VI. VII. IX. X. XI. (
NN ibidem, months: III. IV. V. VI. (
LL ibidem, months: IV. V. (
♂♂, ♀♀ Stara Ves 22.III.1963 collected in the forest biotopes (
♀ Zagreb 5.IV. 1959 from
LL Velika Kapela mountain: Razvala 16.VIII.1968 from
N, LL Lika: Plitvice, Crna rijeka 12.VIII. 1968 from
N, LL ibidem: 12.VIII. 1968 from
N Vukomeričke gorice: Prkovec 19.VII. 1974 from
L Kozjak mountain: Malačka 29.IV. 1974 from
♀♀, N Dugo Selo 9.X.1964 from
♂, ♀♀, NN Dekanovec from
♂♂, ♀♀, NN Nedelišće 3.V.1978 from
N Dubrovnik 12.II. 1966 from
♀, NN, LL Stara Ves 21.IV. 1964 from
NN, LL ibidem: 10.V. 1964 from
NN, LL ibidem: 24.VI.1964 from
ibidem: 15.,16. VII. 1964 from
N Baćinska jezera 30.IV. 1976 from
L Vrana-Biograd 25.IV. 1974 from
NN ibidem: 25.IV.1974 from
NN Metković-Gabela 14.X. 1964 from
L Peščenica-Vratovo 26–27.IV.1984 from
N ibidem: month: V. 1984 from
N, LL ibidem: 3.VI. 1984 from
NN ibidem: 1.VII.1984 from
LL ibidem: 2.VII.1984 from
N Peščenica-Vratovo 22.–23.III.1984 from
N ibidem: 2.VII.1984 from
♂♂, ♀♀ Northern Croatia (
♂♂, ♀♀ Lukovdol, Lipov vrh months: V.- VIII. 2008 from
♂♂, ♀♀, NN Zmajevac 5.III.–30.III. 2011 from
♂♂, ♀♀, NN ibidem: 10. IV.–24.IV. 2011 from
♂♂, ♀♀, NN ibidem: 25.IV.- 30.IV. 2011 from
♂♂, ♀♀, N ibidem: 7.V.- 25.V. 2011 from
LL ibidem: 31.VII.2011 from
♀ ibidem: 26.X. 2011 from
♂, ♀♀, NN Đakovo 10.III.–30.III. 2011 from
♂♂, ♀♀, NN ibidem: 7.IV.- 25.IV. 2011 from
♀♀ ibidem: 2.V.–25.V. 2011 from
♀, NN Sungerski Lug 25.VI. 2011 from
♂♂, ♀♀, NN Mikleuš 27.IV. 2011 collected on vegetation using the cloth-dragging method
♂♂, ♀♀, NN, L ibidem: 14.V. 2011 collected on vegetation using the cloth-dragging method
♂♂, ♀♀, NN ibidem: 8.VI. 2011 collected on vegetation using the cloth-dragging method
♂, ♀, NN ibidem: 9.VI.2011 collected on vegetation using the cloth-dragging method
♂♂, ♀♀, NN ibidem: 15.VI.2011 collected on vegetation using the cloth-dragging method
♂♂, ♀♀, NN ibidem: 3.VII.2011 collected on vegetation using the cloth-dragging method
♂♂, ♀♀, NN ibidem: 4.VII.2011 collected on vegetation using the cloth-dragging method
♂♂, ♀♀, NN, LL ibidem: 2.VIII.2011 collected on vegetation using the cloth-dragging method
♂, ♀♀, NN, LL ibidem: 3.VIII.2011 collected on vegetation using the cloth-dragging method
♀♀, N ibidem: 13.IX.2011 from
♂♂, ♀♀ Rijeka (
♂♂, ♀♀ Split (
♂♂, ♀♀ Kozjak mountain: Malačka (
NN, LL ibidem: 29.IV. 1974 from
♂♂, ♀♀ Velebit mountain (
L ibidem: Veliki Zavižan 28.IX. 1975 from
♀ ibidem: Crni padež 1.VIII. 1975 from
L ibidem: Baške Oštarije 11.VIII. 1968 from
L ibidem: Predzid (pod Ćelavcem) 1.VIII.1975 from
♂♂, ♀♀ Istra: Učka mountain (
LL ibidem: 12.VII. 1969 from
♂♂, ♀♀ Gorski Kotar: Risnjak mountain (
L ibidem: 9.VIII. 1967 from
♀, L ibidem: 1.IX. 1978 from
♀ Velika Kapela mountain: Razvala 16.VIII. 1968 from
LL Lika: Plitvice, Crna rijeka 12.VIII. 1968 from
LL ibidem: 12.VIII. 1968 from
N ibidem: Mukinje 29.V. 1977 from
♀ ibidem: 29.V. 1975 from
♀, N, L Medvednica mountain: Sljeme 25.VI. 1975 from
N Vukomeričke gorice: Prkovec 19.VII.1974 from
N ibidem: 19.VII. 1974 from
♀♀ Peščenica-Vratovo 5.X. 1977 from
N, LL ibidem: 28.X. 1976 from
♀ Križevci, Stara Ves 20.IV.1963 from
♀ Peščenica-Vratovo 26.–27.IV. 1984 from
N ibidem: 2.VII.1984 from
NN ibidem: 3.VI. 1984 from
N ibidem: 1.VIII.1984 from
♀ ibidem: 20.–24.IX. 1983 from
L Peščenica-Vratovo 29.–30.XI.1983 from
♀, N Dubrovnik, Močiljska špilja 31.III.1953 from
♀, L ibidem: 6.V.1961 from
♂ Krbavsko polje: 17.IX.1963 cave habitats (
♀ Cres Island: Petrićevi-špilja Čampari 31.III.1975 from
♂ Buzet, Rabakova peć, Ročko polje, month: VII.1961 from
♂♂, ♀♀ Brač Island (
Creating a list of ticks present in the Croatian fauna is somewhat complicated because the territory of Croatia was formerly part of the Kingdom of Yugoslavia and later part of the Socialist Federal Republic of Yugoslavia. Before the disintegration of Yugoslavia, nearly all of the records were simply summarized as being from Yugoslavia and the faunas of the new European states were not differentiated. For instance,
I would like to thank Dr. Tomi Trilar (Slovenian Museum of Natural History, Ljubljana, Slovenia) for confirming the collected materials and for literature data. Furthermore, I would like to thank Dr. Đurđica Stubičan for assistance in collecting the literature data (Faculty of Veterinary Medicine, University of Zagreb, Croatia) and thank the students Marko Teni for help in field work. Also, I thank Dr. Lawrence J. Hribar of the Florida Keys Mosquito Control District, USA for all improvements in this manuscript.