Corresponding author: Alan F. Burke (
Academic editor: R. Gerstmeier
Six new
Se describen seis nuevas especies del género
The subfamily
The genus
This work is intended to be a small contribution towards a better understanding of the
The methods used for genitalia extraction and dissection are similar to those described by
Specimens were observed using a Leica MZ APO stereomicroscope. All measurements were made using a stereomicroscope ocular micrometer and the software Leica Application Suite V. 3.4.0. Optic images were taken using a Leica DFC 500 digital camera. Scanning electron photographs were taken using a Hitachi 3500N variable pressure scanning electron microscope.
The following abbreviations are used in the description of the holotypes:
Members of the genus
Size: TL= 12.5 mm, length of males: 8.8 - 13.5 mm, length of females 9.8 - 15.2 mm, n = 53 (
Color: head and pronotum fuscous-brown; rest of the body uniformly brown. Each elytron with a pair of median, slightly oblique, pale fascia that extends from elytral suture to epipleuron.
Head: HL= 2.1 mm, HW= 1.3 mm; length to width ratio: males average 1.68, females average 1.55; measured across eyes wider than pronotum; finely, rather punctate; somewhat clothed with short, recumbent setae intermixed with less numerous, erect setae; surface rugose, except frons shiny. Eyes rather big, somewhat rounded, inconspicuously longer than wide, emarginate in front, bulging laterally, separated by approximately 2.5 eye-widths (
Thorax: PL= 3.2 mm, PW= 1.9 mm; length to width ratio: males average 1.56, females average 1.62; pronotum widest at middle, middle slightly broader than anterior margin; sides constricted subapically, more strongly constricted behind middle; disc flat, indistinctly impressed in front of middle; clothed with short, recumbent setae intermixed with long, erect and suberect setae; surface somewhat rugose; slightly more densely punctate than head; subbasal tumescences indistinctly pronounced. Mesosternum scarcely, coarsely punctate. Metasternum convex, puncticulate; covered with fine, recumbent setae. Scutellum subquadrate, notched posteriorly, covered with short, erect setae.
Legs: vested with short, recumbent setae intermixed with long, suberect setae that become more densely arranged on proximal face of tibiae; femora rugulose, moderately, finely punctate; tibiae transversely rugose, moderately, coarsely punctate, vested with short, recumbent setae intermixed with occasional semierect setae.
Elytra: EL= 7.6 mm, EW= 3.4 mm; length to width ratio: males average 2.25, females average 1.78; anterior margin bisinuate; humeri rounded; sides subparallel; base wider than pronotum; widest behind middle; disc flattened apically; apices sinuate, feebly dehiscent; surface feebly rugose; vestiture composed of short, semirecumbent setae intermixed with less numerous, long, erect and semierect setae; sculpturing consisting of small, coarse punctations arranged in striae that gradually reduce in size behind middle; interstices smooth, 4.0 × the width of punctation.
Abdomen: ventrites 1-5 rugulose, vested with short, recumbent setae and some long, semierect setae. First visible ventrite indistinctly, finely punctate; ventrites 2-5 densely, finely punctate. Fifth visible ventrite convex; lateral margins oblique; posterior margin broadly, rather deeply, semicircularly emarginate (
Habitus of:
Antennae of:
Antennae and head in lateral view of:
Ungual structure of the protarsi of:
Terminalia in ventral view of:
Terminalia in dorsal view of:
Male genitalia of:
Female specimens differ from males in the following respects: antennal serration somewhat more evident (
Distinguishable from congeners based on its size, antennal shape, midelytral marking, terminal abdominal segments and male genitalia. This species appears most similar to the allopatric species
Known from the vicinity of the Chamela Biological Station, situated in the Chamela-Cuixmala region, on the western portion of Jalisco, Mexico.
I am very pleased to name this new species in honor of my mother, Rosalina Roco, a cornerstone in my life, and a person whose endless efforts have been of inspirational support during my professional career.
Size: TL= 11.3 mm, length of males 8.5 - 11.25 mm, length of female 10.5 - 11.5 mm, n= 14 (
Color: head and pronotum dark brown; rest of the body uniformly brown. Each elytron with a post median, irregular, narrow, obliquely directed, black fascia that extends from epipleuron to elytral suture, becoming somewhat paler near suture; this fascia is bordered anteriorly by an inconspicuous pale marking; ventrites 1-5 with a pair of irregular, testaceous maculae near sides.
Head: HL= 2.2 mm, HW 1.95 mm; length to width ratio: males average 1.14, females average 1.22; measured across eyes wider than pronotum; finely punctate; surface rugose; vested with short, recumbent setae intermixed with few long, semirecumbent setae that become more numerous toward epistoma. Eyes medium-sized, rather rounded, inconspicuously longer than wide, emarginate in front, somewhat bulging laterally, separated by approximately 3 eye-widths. Antennae extending to base of elytra; third antennomere 2.0 × the length of second
antennomere; fourth antennomere slightly shorter than third antennomere; antennomeres 4-10 subequal in length; antennomeres 2-4 subcylindrical; antennomere 5-10 gradually becoming serrate toward distal end; last antennomere irregularly elongate, sinuate internally, 1.5 × longer than tenth antennomere (
Thorax: PL= 2.7 mm, PW= 1.9 mm; length to width ratio: males average 1.36, females average 1.44; anterior and posterior margins of pronotum as wide as middle; sides feebly constricted subapically; slightly more constricted behind middle; disc flat, inconspicuously impressed in front of middle; moderately, coarsely punctate; less densely punctate than head; surface rugose; vested with short, recumbent setae, intermixed with long erect setae; less densely clothed than head; subbasal tumescences moderately projected. Mesosternum clothed with long, recumbent setae; coarsely punctate. Metasternum smooth, convex; rather puncticulate. Scutellum semicircular, covered with short, recumbent setae, posterior margin slightly notched.
Legs: somewhat covered with short and long semirecumbent setae that become more numerous on second half of tibiae; femora finely punctate, longitudinally rugose; tibiae coarsely punctate, transversely rugose.
Elytra: EL= 6.2 mm, EW= 3.45 mm; length to width ratio: males average 1.61, females average 1.76; anterior margin bisinuate, broader than pronotum; humeri rounded; sides subparallel; widest behind middle; disc subflattened above; surface rugose; apices rounded, feebly dehiscent; clothed with short, semirecumbent setae intermingled with less densely, longer, erect setae; sculpturing consisting on coarse punctations arranged in striae that gradually reduce in size behind middle; interstices 3.0 × the width of punctation.
Abdomen: ventrites 1-5 rugulose; inconspicuously vested with short, recumbent setae; sparsely, finely punctate. Fifth visible ventrite convex; lateral margins oblique; posterior margin truncate (
Fifth tergite rugulose; surface convex; lateral margins oblique; posterior margin broadly, shallowly, arcuately emarginate; posterior angles rounded (
Male specimens differ from females by having the posterior margin of the fifth visible ventrite broadly, shallowly, arcuately emarginate (
Separable from other
Known from two localities in the Guanacaste Province, Costa Rica: Playa Naranjo, adjacent to Santa Rosa, Guanacaste National Park; and Sector Las Pailas, Rincón de la Vieja National Park.
The specific epithet comes from the Latin word capax (= wide), a noun that makes allusion to the overall robust appearance of this new species.
Size: TL= 10.4 mm, length of males 10.8 - 12.2 mm, length of females 7.75 - 10.5 mm, n = 7 (
Color: head and pronotum fuscous; elytra, scutellum and legs brown, except posterior half of femora dark brown; antennae and mouthparts testaceous; abdomen, meso and metasternum light brown. Each elytron with three irregular, variably sinuate, pale fasciae; first on anterior fourth, slender, extending from elytral suture to epipleuron, surrounding scutellum; next fascia on second fourth, broader than preceding band, extending from elytral suture to epipleuron; third fascia on last fourth, slightly shorter and narrower than preceding fascia, extending from elytral suture to tenth stria, not reaching epipleuron.
Head: HL= 2.2 mm, HW= 1.95 mm; length to width ratio: males average 1.08, females average 1.17; measured across eyes wider than pronotum; densely, coarsely punctate; surface rugose; clothed with short, recumbent setae intermixed with long, erect setae; frons feebly bi-impressed. Eyes somewhat small, subsinuate, longer than wide, moderately emarginated in front, bulging laterally, separated by approximately 4 eye-widths (
Thorax: PL= 2.5 mm, PW= 1.75 mm; length to width ratio: males average 1.45, females average 1.39; pronotum widest at middle, middle slightly broader than anterior margin; sides constricted subapically, more strongly constricted behind middle; disc flat, feebly impressed in front of middle; subbasal tumescences pronounced; surface rugose, moderately, finely punctate;
less densely punctate than head; covered with short, recumbent setae interspaced with less numerous, long, erect and suberect setae. Mesosternum coarsely punctate. Metasternum convex; puncticulate; surface rugulose; median impression strongly indicated. Scutellum semicircular, covered with some short, recumbent setae.
Legs: vested with short, recumbent setae intermixed with occasional long, suberect and erect setae that become more densely arranged on tibiae; femora rugulose; tibiae transversely rugose.
Elytra: EL= 5.5 mm, EW= 2.5 mm; length to width ratio: males average 2.26, females average 2.19; anterior margin arcuately emarginate, as wide as pronotum; humeri feebly indicated; sides subovoid, widest at second third; disc moderately flattened apically; apex broadly, separately rounded, dehiscent, covering sixth tergite; surface smooth, clothed with short, recumbent setae combined with less numerous, long, erect setae; sculpturing consisting on rather numerous, coarse punctations arranged in striae that abruptly reduce in size and become less numerous on last fourth; interstices about 2.0 × the width of punctation.
Abdomen: ventrites 1-5 rugose; each segment with a pair of large, shallow impressions near sides; somewhat clothed with short, fine, pale, recumbent setae; moderately, coarsely punctate. Fifth visible ventrite convex; lateral margins oblique; posterior margin broadly, deeply, arcuately emarginate (
Males have the fifth visible ventrite convex, rugose, moderately, coarsely punctate, lateral margins oblique, posterior margin broadly, deeply, arcuately emarginate (
angles robust, strongly arcuate at apex (
This species recalls various
The species is known from two localities: La Muralla National Park, situated in the department of Olancho, Honduras and El Boquerón National Park, in the Department of San Salvador, El Salvador.
The specific name comes from the Latin noun
Size: TL= 7.8 mm, length of males 7.5 to 9.2 mm, length of females 6.5–7.8 mm, n = 9 (
Color: head fuscous-brown; pronotum, mouthparts, mesosternum, metasternum and abdomen testaceous; elytral ground predominantly testaceous except median region pale-testaceous; legs mostly testaceous, except posterior half of femora brown. Each elytron with two pairs of black, irregularly marked maculae; the first adjacent to anterior margin, extending from second stria to humeral angle; the second located on first half of second fourth, more faintly marked than preceding pair, extending from second to eighth stria; a long, irregular, transversally marked, black fascia located on third fourth, in the form of a vitta, extending from suture to epipleuron, covering about one fourth of elytral length.
Head: HL= 1.3 mm, HW= 1.25 mm; length to width ratio: males average 1.06, females average 1.12; measured across eyes wider than pronotum; densely, coarsely punctate; surface rugose; vested with short, semirecumbent setae interspaced with some long, erect setae. Eyes rather small, subsinuate, longer than wide, moderately emarginate in front, somewhat bulging laterally, separated by approximately 3.5 eye-widths. Antennae extending beyond basal sixth of elytra; antennomeres 2-3 subequal in length; fourth antennomere 1.5 × longer than preceding antennomere; antennomeres 4-5 subequal in length; sixth antennomeres slightly shorter than fifth antennomere; antennomeres 6-10 subequal in length; antennomeres 2-5 slender; antennomeres 6-10 weakly serrate; last antennomere 1.5 × longer than tenth antennomere, subsinuate, flattened apically (
Thorax: PL= 1.7 mm, PW= 1.2 mm; length to width ratio: males average 1.38, females average 1.46; pronotum widest at middle; sides constricted subapically, more strongly constricted behind middle; disc flat, rather impressed in front of middle; surface shiny; moderately, finely punctate; less densely punctate than head; somewhat covered with short, semirecumbent setae intermingled with long, erect setae; subbasal tumescences rather pronounced. Mesosternum moderately, coarsely punctate; vested with short, recumbent setae. Metasternum smooth; surface strongly convex, puncticulate laterally; median region with a sensory area consisting of rather dense, short, erect setae set on a rugose ground (
Legs: covered with short, semirecumbent setae intermixed with some long, erect and suberect setae, vestiture become more densely arranged on distal half of tibiae; femora rugulose, feebly punctate; tibiae rugose, somewhat punctate.
Elytra: EL= 4.7 mm, EW= 1.85 mm; length to width ratio: males average 2.49, females average 2.38; anterior margin bisinuate, slightly broader than pronotum; sides subparallel; widest behind middle; base slightly wider than pronotum; humeri moderately indicated; disc subflattened above; apices rounded, dehiscent, covering sixth tergite; surface smooth, somewhat covered with short, erect and semierect setae interspaced with long, erect setae; sculpturing consisting of coarse, deep punctations arranged in striae that gradually reduce in size behind middle; interstices 2.0 × the width of punctation.
Abdomen: ventrites 1-5 rugulose; moderately, finely punctate; somewhat covered with short, recumbent setae combined with some long, erect setae. Fifth visible ventrite convex; lateral margins oblique; posterior margin broadly, deeply, arcuately emarginate (
Metasterna of:
Females differ from male specimens as follows: antennomeres 5-10 weakly serrate, sixth visible ventrite subtriangular, lateral margins oblique, posterior margin broadly, very shallowly emarginate (
Its distinct elytral ground color, slender form, shape of terminal abdominal segments and male genitalia will readily separate this species from congeners.
Known from three localities in Panamá: El Llano, Panamá Province; 4 km south of Gamboa, Colón Province; and 4 km south of El Valle, Coclé Province.
The specific epithet comes from the Latin noun
Size: TL= 7.7 mm, length of males 7 - 9.2 mm, length of females 6.8 - 9.8 mm, n = 13 (
Color: head, pronotum, prosternum, mesosternum, metasternum, and abdomen ferrugineous; elytral ground dark-testaceous; legs, antennae and mouthparts testaceous; sixth visible ventrite light brown, slightly lighter than remaining visible ventrites (
Head: HL= 1.2 mm, HW= 1.4 mm; length to width ratio: males average 0.91, females average 1.02; measured across eyes wider than pronotum; densely, moderately coarsely punctate; surface rugose; covered with short, recumbent setae intermixed with less abundant, long, erect setae that become more densely arranged behind eyes; frons moderately bi-impressed. Eyes rather small, subsinuate, longer than wide, feebly emarginate in front, somewhat bulging laterally, separated by approximately 3.5 eye-widths. Antennae extending to basal sixth of elytra; antennomeres 2-3 subequal in length; fourth antennomere inconspicuously longer than third antennomere; antennomeres 4-10 subequal in length; antennomeres 2-5 slender; antennomeres 6-10 weakly serrate; last antennomere subsinuate, flattened apically, about 1.5 × longer than tenth antennomere (
Thorax: PL= 1.7 mm, PW= 1.2 mm; length to width ratio: males average 1.39, females average 1.33; pronotum widest at middle; sides constricted subapically, more constricted behind middle; surface somewhat rugose; disc flat, somewhat impressed in front of middle; moderately, finely punctate; less densely, deeply punctate than head; clothed with short, semirecumbent setae intermingled with long, erect setae; subbasal tumescences pronounced. Mesosternum rugulose, coarsely punctate. Metasternum rugose, moderately, coarsely punctate; somewhat clothed with short, recumbent setae; absence of sensory area (
Legs: vested with short, recumbent setae, intermixed with longer semierect setae, vestiture becomes more abundant on internal face of tibiae; femora rugulose, finely punctate; tibiae rugose, moderately, coarsely punctate.
Elytra: EL= 4.5 mm, EW= 2.0 mm; length to width ratio: males average 2.18, females average 2.12; anterior margin bisinuate, slightly broader than pronotum; sides subparallel, inconspicuously wider behind middle; humeri moderately indicated; apices rounded, dehiscent; covering sixth tergite; surface shiny, vested with short, recumbent setae interspaced with some long, erect setae; sculpturing consisting of coarse, deep punctations arranged in striae that gradually reduce in size behind middle; interstices smooth, 2.0 × the width of punctuation.
Abdomen: ventrites 1-4 moderately, finely punctate; smooth; somewhat vested with fine, short, recumbent setae interspaced with few long, erect setae; hind margins truncate. Fifth visible ventrite smooth; surface convex; lateral margins oblique; posterior margin broadly, deeply, arcuately emarginate (
Female specimens have the sixth visible ventrite subquadrate, rugulose, surface strongly convex, lateral margins oblique, posterior margin truncate (
Known from five localities in Panamá: the Canal Zone, 5 km south of Gamboa; Fuerte Kobbe; the mountainous region of Cerro Azul; the surroundings of Palenque; and El Llano-Carti road, 12 km north of El Llano.
The specific epithet comes from the Latin noun
TL= 9.8 mm, length of males 10.2–11 mm, length of female 9.4 mm, n = 4 (
Color: head predominantly black (
Head: HL= 1.3 mm, HW= 1.5 mm; length to width ratio: males average 0.92, female 0.86; measured across eyes wider than pronotum; surface rugose; frons feebly bi-impressed; moderately, rather coarsely punctate; clothed with short, recumbent setae intermixed with some long, semierect and erect setae. Eyes medium-sized, rather rounded, inconspicuously longer than wide, feebly emarginate in front, somewhat bulging laterally, separated by approximately 3 eye-widths. Antennae reaching humeral angles; antennomeres 2-3 subcylindrical, slender; antennomeres 4-10 strongly serrate, longer than broad; third antennomere somewhat longer than second antennomere; fifth antennomere slightly shorter than fourth antennomere; antennomeres 5-10 subequal in length; last antennomere 2.5 × longer than tenth antennomere (
Thorax: PL= 2.1 mm, PW= 1.15 mm; length to width ratio: males average 1.85, female 1.75; pronotum rugose; widest at middle; middle slightly wider than front margin; sides constricted subapically, more strongly constricted behind middle; disc flat, feebly impressed in front of middle; subbasal tumescence pronounced; vested with short and long semirecumbent setae interspaced with some erect setae; surface moderately, finely punctate. Mesosternum rugose; moderately, coarsely punctate; somewhat clothed with long, semirecumbent setae.
Metasternum shiny; surface convex, puncticulate; covered with long, semirecumbent setae. Scutellum subquadrate; wider than long; notched medially.
Legs: clothed with long, erect setae and some short, recumbent setae that become more abundant on posterior half of tibiae; femora shiny, finely, indistinctly punctate, transversely rugose; tibiae coarsely, densely punctate, longitudinally rugose.
Elytra: EL= 6.1 mm, EW= 2.8 mm; length to width ratio: males average 2.15, female 2.18; anterior margin bisinuate, wider than pronotum; disc smooth, flattened above; humeri indicated; sides subparallel, widest on third fourth; apices weakly dehiscent, rounded, covering sixth tergite; elytral declivity somewhat procurved, clothed with short, semirecumbent setae intermingled with long, erect, less densely arranged setae; sculpturing consisting of coarse punctations arranged in striae that gradually reduce in size behind middle; interstices smooth, about 1.5 × the width of punctation.
Abdomen: ventrites 1-5 rugose; moderately, coarsely punctate; somewhat clothed with short, recumbent setae interspaced with some long, erect setae. First visible ventrite strongly convex; posterior margin elevated, with a transverse carina that initiates next to hind angles and produces a broad, deep, arcuate emargination (
The only female in the type series differs from the male by having the first visible ventrite moderately longer and the posterior margin of ventrites 1-2 truncate (
The distinctive coloration and elytral markings of this species will serve to separate it from congeners. No other species in the
This species is known from three localities in the state of Chiapas, Mexico: El Aguacero, in the periphery of Tuxtla Gutierrez; El Chorreadero, 8 km northeast of Chiapa de Corzo; and on highway 190, 17 km west of Tuxtla Gutierrez.
The specific epithet comes from the Latin noun
(5 males, 2 females). 2 males: Texas, Hidalgo Co., Bentsen Rio Grande State Park, 16-VI-1974, G. H. Nelsen (FSCA); 3 males and 1 females: USA, Texas, Hidalgo Co., LRGVNWR, MacManus unit, (
I am thankful to the following persons and institutions for all the material allocated to me: Jacques Rifkind, Weston Opitz, James E. Wappes, Charles R. Turnbow Jr., the Texas A&M Insect Collection, the Florida State Collection of Arthropods, the Universidad Nacional Autónoma de México National Insect Collection, and the Instituto Nacional de Biodiversidad de Costa Rica. My special thanks go to Gregory Zolnerowich for allowing me the use of his microscopy equipment and Frank Merickel for his helpful comments and suggestions. I am especially indebted to Roland Gerstmeier, Jaime Solervicens and Weston Opitz, for all the observations and suggestions that improved this document. Finally, I am most grateful to Jacques Rifkind for his invaluable observations and recommendations prior to and during the preparation of this manuscript.