Corresponding author: Miguel A. Alonso-Zarazaga (
Academic editor: Terry Erwin
A new genus
The family
Their systematics is extremely difficult and based mainly on structures of the male genitalia (
A study of the type specimens of Oriental
Specimens of the new genus
Descriptions were made using a binocular LeicaWild MZ8, provided with a photographic tube. Photographs were taken with an Olympus C7070WZ. Extended focus images were generated with CombineZP 7.0 by Alan Hadley and edited with Adobe Photoshop CS 5.0 if required. Microscope slides were studied under a light microscope and drawings were made using a drawing tube.
With the characters of the tribe
This genus is named in honour of our good colleague and friend Dr Christopher H.C. Lyal (Natural History Museum, London), one of the best world experts on weevils (
This genus includes for the moment three nominal species in the Oriental Realm, two of them having been wrongly placed in the Afrotropical genus
(holotype).
(paratype).
As male, but
The male paratype has a length (without rostrum) of 4030 µm, a length (standard) of 3770 µm and a ratio Lrostrum/Lpronotum of 1.44. Golden coloration of scales is a bit more extended than in the holotype, and all femora show 1+3 teeth. In general in
Holotype: 1 male, labelled as follows: yellowish: Jardin Botanique; yellowish: SAIGON / 17-VII-50 / J. BARBIER; yellowish: MUSEUM PARIS / Coll. / J. Barbier; white: AZ-0001; red: HOLOTYPE ♂/ LYALIA CURVATA / n. sp./ Alonso-Zarazaga / et Perrin 2011 (Perrin’s handwriting). Specimen coming from Coll. Barbier, now included in the General Collection.
Paratypes: 1 female, labelled: yellowish: Jardin Botanique; yellowish: SAIGON / 15-VI-50 / J. BARBIER; white: COTYPE (in red); yellowish: MUSEUM PARIS / Coll. / J. Barbier; white: Nanophyes maximus nsp (Pic’s handwriting); white: AZ-0002; red: PARATYPE ♀/ LYALIA CURVATA / n. sp./ Alonso-Zarazaga / et Perrin 2011 (Perrin’s handwriting). To our knowledge this nominal taxon by Pic has never been described. These two specimens are close to a bottom label stating: “Nanophyes major / maximus, barbieri / et testaceicollis in litt. / selon Zherikhin 1997”. 1 male: Saigon; Nanophyes / maximus / Pic (locality and identification: Hoffmann’s handwriting); MUSEUM PARIS / 1968 / Col. A. HOFFMANN; AZ-0313; red: PARATYPE ♂/ LYALIA CURVATA / n. sp./ Alonso-Zarazaga / et Perrin 2011 (Perrin’s handwriting); white: Exchange /M.N.H.N. It lacks the left antenna beyond the scape, left midleg beyond base of tibia (broken), onychium of right midleg and tarsi of both hindlegs. This specimen was probably part of the Barbier’s series and transferred to Hoffmann’s collection “à la Hoffmann” (Perrin, 1998). It has been agreed that this specimen will be transferred to the Coll. Alonso-Zarazaga (MNCN, Madrid).
This species is named
This species is known only from its type locality, Saigon (now Thành phố Hồ Chí Minh or Ho Chi Minh City) in Southern Vietnam. Nothing is known about its host plant, but it will probably be a species of
Types of
Types of
(lectotype of
Lectotype of
As in male, but 1st elytral interstria immaculate.
Lectotype of
Syntypes of
This species is known from the island of Java (Indonesia), without any further detail. It has been recently recorded from the island of Bali (Pura Taman musi, Buleleng prov.) (
This species seems to be trophically linked to
It is curious that this synonymy was already known to Van Emden (1938: 27), who treated the Javan Kalshoven’s specimens as
This species was incorrectly placed in the Afrotropical genus
This species is perhaps a synonym of
It is only known from the Indian state of Assam: Kaziranga and Tejpur, Chardaur, without any definite host plant (“ex a forest tree”).
The single originally included and type species was thoroughly described by
The generic description, thus, coincides with the specific one given by
It is a great pleasure to name this new genus after Mr. Junnosuke Kantoh, young Japanese entomologist, with our wish for a long and fruitful career.
The penis and the spiculum gastrale of this species are distinctive. It is known only from Taiwan and was captured on flower buds of
Abdominal ventrites:
The new genus
On the other hand,
The genera of
1 | Stria 10 complete | 2 |
– | Stria 10 erased or fused to 9 between metacoxal and suture II level | 4 |
2 | Interstria 8 with a complete crenulate keel running up to the basal third of elytron. Flagellum of penis conspicuously longer than penis |
|
– | Interestria 8 not visibly crenulate in basal third. Flagellum of penis conspicuously shorter than penis. | 3 |
3 | Procoxae acutely projecting in both sexes, trochanters inserted laterally on procoxa. Rostrum almost straight. Eyes visibly separated on frons |
|
– | Procoxae rounded at apex, trochanters apical or subapical. Rostrum conspicuously curved. Eyes closely approximated on frons, leaving one row of hairs on midline |
|
4 | Interstria 8 not visibly crenulate in basal third. Humeral callus obsolete or absent. Intermesocoxal distance 0.3–0.4 × the intermetacoxal one |
|
– | Interstria 8 crenulate, at least partially. Humeral callus developed. Intermesocoxal distance more than 0.75 × the intermetacoxal one | 5 |
5 | Interstria 8 not crenulate on callus, keel reaching basal third. Arms of spiculum gastrale unwinged. Tegmen with parameroid lobes not or hardly developed, at most separated by a small notch. Flagellum conspicuously shorter than penis. Body integument monochrome black |
|
– | Interstria 8 crenulate on callus, keel reaching basal or apical third. Arms of spiculum gastrale winged. Tegmen with parameroid lobes deeply cleft. Flagellum at least as long as penis. Body integument maculate | 6 |
6 | Apex of tarsomere 1 deeply emarginate in V. Crenulate keel of interstria 8 reaching apical third of elytron. Parameroid lobes very long, widely separated |
|
– | Apex of tarsomere 1 subtruncate to roundly notched. Crenulate keel of interstria 8 reaching basal 3rd of elytron (mid-metasternum level). Parameroid lobes short, narrowly separated |
|
This artificial key, however, does not show the real affinities of the genera.
The known species of the genus
1 | Second and third elytral striae strongly shifted towards suture in basal third, 2nd interstria usually less than 1.4 × as wide as interstria 1 at same level ( |
|
– | Second and third elytral striae not or weakly shifted towards suture in basal third, 2nd interstria at least 1.5 × as wide as interstria 1 at same level ( |
2 |
2 | Male rostrum longer, 1.45–1.47 × as long as pronotum. Scape |
|
– | Male rostrum shorter, ca. 1.35 × as long as pronotum. Scape 1.5 × as long as funicle in male. Male pronotum 1.67 × as wide as long, base |
|
Studies in the MNHN (Paris) by the senior author were facilitated by the attribution of a grant of Invited Professor during January-February 2010, for which he thanks the authorities of the Museum. We also thank Mr Maxwell V. L. Barclay and Dr Christopher H. C. Lyal (NHM, London) for the loan of specimens, Dr Wang Zhiliang (IZCAS, Beijing) for his help with the graphic material and Dr Christian Salcedo (IZCAS, Beijing) for the linguistic review. This research has been supported by project grant CGL2010-15786 (Ministerio de Ciencia e Innovación, Spain).