Corresponding author: Ziro Komiya (
Academic editor: S. Lingafelter
Two new species of the genus
Recently,
In the redescription of the genus in
The presence of
In this paper, we describe two new species of
The collection acronyms used in the text are as follows:
1 | Distance between anterior apex of upper eye lobe and posterior end of antennal insertion equal to the length of scape ( |
|
– | Distance between anterior apex of upper eye lobe and posterior end of antennal insertion much shorter than the length of scape ( |
2 |
2 (1) | Carina of ventral sensorial area of antennomeres III-XI not visible in lateral view ( |
|
– | Carina of ventral sensorial area of antennomeres III-XI distinctly visible in lateral view (primarily the apical antennomeres) ( |
3 |
3 (2) | Dorsal sensorial area of antennomere XI shorter than one-sixth of total length ( |
|
– | Dorsal sensorial area of antennomere XI longer than one-fourth of total length ( |
Dedicated to Dr. Akiko Saito of the Natural History Museum and Institute, Chiba, who collected the holotype.
Female: Integument dark-brown; parts of the mandibles, margins of the pronotum, elytral suture, pro- and mesosternal process, trochanters, extreme ventral apex of tibiae blackish; elytra laterally and apically darker than the remaining surface.
Dorsal area of head coarsely punctate; punctures sparser at central area, coarser, more abundant and confluent at side; punctures between upper eye lobes with short, apically spatulate setae; punctures between posterior eye margin and margin of pronotum with very short, thick setae. Area behind eyes coarsely, confluently punctate near apices of upper eye lobes, sparsely near lower eye lobes; nearly all punctures with single, very small, thick seta. Clypeus coarsely, densely punctate, except at smooth central area; punctures with short, spatulate setae. Labrum smooth, glabrous laterally; coarsely punctuate; with short, spatulate setae around central projection. Upper eye lobe not notably separated from base of antenna. Submentum coarse; densely punctate near anterior margin (confluently centrally), sparsely and coarsely near gena; each puncture with a small, spatulated seta; anterior edge slightly elevated throughout. Mentum with dense, spatulate setae, somewhat longer laterally. Mandibles punctate; external punctures coarse, deep, dense and accompanied with spatulate setae; internal punctures sparse. Carina of ventral sensorial area of antennomeres elevated and visible in lateral view (
Pronotum finely, sparsely punctate; punctures gradually becoming coarser and more abundant near lateral margin; lateral punctures with very small, spatulate seta. Prosternum coarsely, moderately densely punctate; nearly all punctures with small, spatulate seta. Prosternal process sparsely punctate (punctures with same type setae as prosternum). Mesosternum strongly, densely, deeply, confluently punctate, with a few very small, spatulated setae. Mesosternal process subglabrous and smooth. Mesepisterna coarsely, moderately densely punctate. Metasternum finely and sparsely punctate centrally; punctures gradually becoming coarser and denser towards lateral area; punctures close to metacoxae with small, spatulate setae. Elytra coarse; punctures dense on lateral parts and apical one-fourth; lateral punctures with very small thick setae, more conspicuously towards apices; punctures of apical one-third with small, spatulate setae; suture divergent at apex.
Urosternites densely punctate, mainly laterally; setae short, dense and spatulate, thick laterally; urosternite V with somewhat long, dense setae at margin, more conspicuous centrally (each seta apically spatulate). Coxae moderately sparsely punctate. Tibiae somewhat flat dorsally and sulcated laterally. Metatarsomere V (excluding claws) longer than I-III together.
Holotype ♀ from INDONESIA, Sulawesi,
The name refers to the continent where the species occurs.
Female: Integument dark-brown; parts of mandibles, margins of pronotum, elytral suture, pro- and mesosternal process, trochanters, extreme ventral apices of tibiae blackish.
Dorsal face of head punctate coarsely for the most part; punctures sparser at central area, and coarse, dense and confluent near each side; punctures on area between upper eye lobes with short setae (lateral ones apically spatulate), more abundant near clypeus; punctures on area between posterior eye lobe and pronotum without setae. Area behind eyes coarsely punctate near the apices of upper eye lobes, sparsely near lower eye lobes; most punctures at the area close to upper eye lobe with a very small seta, slightly thicker at apex. Clypeus coarsely and densely punctate; punctures with short, spatulate setae. Labrum smooth and subglabrous except area near central projection with coarse punctures and short setae, in part spatulate, some only thick or acute apically. Upper eye lobe not notably distant from the base of antenna. Submentum coarse, moderately, densely punctate near anterior margin (confluently so centrally), gradually becoming sparse and coarse towards gena; each puncture with single, small seta (nearly all thicker apically; some of them spatulate; some acute apically); anterior margin slightly elevated throughout. Mentum with moderately sparse spatulate setae which somewhat longer laterally. Mandibles coarse; external punctures deep and dense, with small setae (very few spatulate); internal ones sparse. Carina of ventral sensorial area of antennomeres elevated and visible in lateral view (
Pronotum finely, sparsely punctate centrally, gradually becoming more coarsely and densely punctate laterally; lateral punctures with very small, spatulate seta. Prosternum coarsely, moderately, densely punctate, and nearly all punctures with small, apically spatulate seta. Prosternal process with small and spatulate setae. Mesosternum, densely, deeply, confluently punctate, and with small, spatulate setae. Mesepisterna coarsely, moderately, densely punctate. Metasternum finely and sparsely punctate centrally; punctures gradually becoming coarse, moderately abundant towards lateral area; lateral punctures close to metacoxae with small setae some of which spatulate. Elytra coarse; punctures dense on lateral parts and apical third; lateral punctures with very small seta, thicker towards apex and conspicuous from base to apex; nearly all punctures with small seta, more distinct towards apex (many spatulate, at least apically); suture not divergent at apex.
Urosternites densely punctate, mainly laterally; setae short, dense and spatulate (at least apically), mainly laterally; urosternite V with somewhat long, dense setae at margin, more conspicuous centrally (very few of them apically spatulate). Coxae moderately, sparsely punctate. Tibiae somewhat flat dorsally, and sulcate laterally. Metatarsomere V (excluding claws) longer than I-III together.
Holotype ♀ from VIETNAM,
We are grateful to Drs. Shusei Saito of Japanese Wildlife Research Center, Akiko Saito of the Natural History Museum and Institute, Chiba, Gérard L. Tavakilian of Muséum National d’Histoire Naturelle, Paris, France, Mrs. Nobuhiko Katsura, Tokyo, Japan, Norbert Delahaye, Brussels, Belgium, Dr. Karl Adbauer, Graz, Austria, and Kiyoshi Matsuda, Takarazuka, Japan for their help in providing material and in many other matter in preparation of the present study.