Length: 15.0 mm; width: 7.0 mm. Habitus as in Fig. 2A.

Figure 2. 

Orthogonius sarawakensis Tian & Deuve, 2006 A habitus, male, holotype B median lobe, right lateral view C apical lamella, dorsal view.

Head dark brown, other parts of the body yellowish brown; head stout, as long as wide; labrum slightly emarginated at frontal margin; clypeus even near base, with two longitudinal furrows along both side, joining to frontal impressions respectively; mentum bisetose; pronotum transverse, PW/PL = 1.55, widest at about middle, strongly convex; lateral expanded margins almost flat, equal wide throughout; elytra elongate ovate, EL/LW = 1.54; base well-bordered; the 3^rd interval with three setiferous pores, all are located on the interval; tarsal claws pectinate; prosternal process unbordered at apex, abdominal ventrite VII shallowly emarginated at apical margin in male.

Male genitalia (Fig. 2B, C): Short and robust, expanded in median portion; in dorsal view, apical lobe elongated, gradually narrowed towards apex, with apical lamella evidently longer than wide.

Female: Unknown.

Easily separated from its congeners, O. perakicus Tian & Deuve, 2007 and O. morvanianus sp. n., by its unbordered prosternal process at apex, pointed elytral apex, and more elongated apical lobe of aedeagus.

1 male, the holotype, “Nord Borneo, Mont Kina Balu, 5-8, 1903, John Waterstradt”, in MNHN; 1 male, same data as in holotype, in SCAU.

Malaysia (Sarawak).

Length: 14.0 mm; width: 5.5 mm. Habitus as in Fig. 3A.

Figure 3. 

Orthogonius perakicus Tian & Deuve, 2007 A habitus, male, holotype B median lobe, right lateral view C apical lamella, dorsal view.

Elytra and sides of pronotum yellowish brown, head dark brown, pronotum light dark brown on disc; head as long as wide, clypeus smooth, front margin of labrum slightly emarginated; mentum bisetose; pronotum strongly transverse, PW/PL = 1.62; lateral expanded margins narrow and reflexed throughout, evenly widened; elytra well-bordered at base, elongate ovate, EL/EW = 1.58; apex broadly truncate, without outer angles, inner angles broadly obtuse; intervals flat; the 3^rd interval with three setiferous pores, all on median portion of the interval; legs moderately developed; the 4^th hind tarsomere emarginated as deep as 1/3 of the joint; all tarsal claws pectinate; prosternal process well-bordered at apex; abdominal ventrite VII in male slightly emarginated at apical margin.

Male genitalia (Fig. 3B, C): Median lobe of aedeagus stout, more or less bent near apex, apex pointed in profile, but obtuse in dorsal aspect, apical lamella longer than wide.

Allied to O. morvanianus sp. n. Differs from the latter by its labrum shallowly emarginated at front, the 4^th hind tarsomere having a deeper emargination, and apical lamella of aedeagus blunt at apex (see below for detail).

1 male, the holotype, “Perak”, in the Collection of Bates, MNHN.

Malaysia (Perak).

Male, “Thailand South, near Khao Sok NP, 3-6. II. 1997, A. Kudrna lgt”, in CPM.

Length: 14.0 mm; width: 5.5 mm. Habitus as in Fig. 4A.

Figure 4. 

Orthogonius morvanianus sp. n. A habitus, male, holotype B median lobe, right lateral view C apical lamella, dorsal view.

Yellowish brown, with head dark brown, the 2^nd–11^th antennomeres, disc of pronotum, tibiae and tarsi of legs brown; surface glabrous, impuctate, head and pronotum finely striate, the 3^rd, 5^th and 7^th elytral intervals each with a few tiny sparse punctures; surface moderately shiny; microsculptural engraved meshes densely isodiametric on the surface of head, pronotum and elytra; body rather flat.

Head stout, as long as wide; eyes very large and prominent; frons flat, vertex slightly convex, smooth; frontal impressions very small, foveate; clypeus bisetose, rather smooth near base; labrum 6-setose, evidently emarginate at apical margin; ligula very small and narrow, bisetose at apex; palps slender or elongate, subcylindrical, normally setose; palpiger asetose, mentum edentate, mentum and submentum each with one pair of setae, mental ones much shorter; antennae slender and long, reaching 1/4 of elytra from base, pubescent from apical 2/3 of the 4^th antennomere; the 3^rd antennomere as long as the 4^th, both shorter than the 1^st; the 1^st–3^rd antennomeres glabrous; evidently expanded laterally in the 4^th and 5^th antennomere; the 1^st antennomere 1.6 times as long as the 2^nd.

Pronotum strongly transverse, PW/PL = 1.78, sides evenly rounded, widest at about middle; both basal and fore margins beaded; lateral expanded margins well defined, subequal wide throughout, strongly reflexed, smooth though with punctate-like structures; both fore and hind angles rounded off; disc moderately convex, fore transversal impression evident, while hind one well-marked, joining basal foveae and the lateral expanded margins.

Elytra elongate ovate, much longer than wide, EL/EW = 1.63, moderately convex, apex broadly truncate, without outer apical angle, inner apical angle broadly obtuse; basal borders well-bordered; sides more or less paralleled in middle portion; striae quite deep; intervals nearly flat, subequal in width; the 3^rd interval with three setiferous pores, anterior one close to the 3^rd stria, while other two, median and posterior ones, on median portion of the interval; the 7^th interval not carinated and without setae throughout.

Legs moderately developed; hind femur long and elongate, with 2 setae posteriorly; middle and hind coxae smooth and glabrous; fore tibia deeply sinuate at apical margin, outer angle sharp, outer margin distinctly serrate; middle tibia slender, not dilated medially in male; hind tibia slender, apex slightly dilated, apical spurs very long and sharp, the 1^st tarsomere slightly longer than the 2^nd, the 3^rd tarsomere about 1.2 times longer than the 4^th which is bilobed at apex, with outer lobe slightly shorter than half as long as the joint, outer lobe evidently longer than the inner; all tarsal claws moderately pectinate.

Male genitalia (Fig. 4B, C): Robust, expanded in median portion, then deeply concave on ventral margin, apex suddenly contracted, sharp, dorsal opening long and large, apex more or less pointed, with a fine tooth at subapex; in dorsal view the apical lamella quite long, slightly longer than wide, roundly obtuse at apex.

Prosternal process bordered at apex; apical margin of abdominal ventrite VII feebly emarginated in male.

Female: Unknown.

Close to O. perakicus Tian & Deuve, 2007, but labrum strongly emarginated at frontal margin (slightly emarginated in O. perakicus), the 4^th hind tarsomere deeply emarginated, but with outer lobe about 1/3 as long as the joint (half as long as the joint in O. perakicus).

To be dedicated to Mr. Pierre Morvan (Carentoir, France), a good expert of Carabidae.

Southern Thailand.

male, “Malaisie, Sabah, Crocker Range, Avril 94, Chew”, in MNHN; paratypes: 1 male, “Malaysia, N. Borneo, Sabah, Keningau distr. Trus Madi Mt. 1150 m, 7.VII.2011, A. Klimenko legit”, in CIB; 1 female, IBID, in SCAU.

Length: 16.0–17.0 mm; width: 6.5–6.6 mm. Habitus as in Fig. 5A.

Figure 5. 

Orthogonius sabahicus sp. n. A habitus, male, holotype B median lobe, right lateral view C apical lamella, dorsal view.

Head dark brown, other parts reddish brown; smooth and glabrous, impunctate, elytral intervals covered with fine and sparse punctures; head and pronotum faintly wrinkled; microsculptural engraved meshes isodiametric on head, pronotum and elytra.

Head stout, slightly longer than wide, HL/HW = 1.05–1.10; vertex convex; labrum quite long, nearly as long as wide, widely and feebly emarginated at frontal margin, 6-setose; clypeus bisetose, base raised and sides longitudinal furrowed, joining to the frontal impressions; frontal impressions pit-like; eyes moderately developed and prominent; palps slender, subcylindrical, the 3^rd maxillary palpomere as long as the 4^th, both glabrous, with two short setae at apex of the 3^rd; the 2^nd labial palpomere slightly longer than the 3^rd, bisetose on inner margin, the 3^rd faintly and sparsely setose; ligula narrow, bisetose at apex; mentum and palpigers unsetose, submentum bisetose; antennae extending beyond basal 1/5 of elytra, pubescent from the 4^th antennomere, starting from basal 1/4; the 1^st–3^rd antennomeres and basal 1/4 of the 4^th antennomere glabrous; the 1^st antennomere stouter than other, with a long seta at subapex, several shorter setae present at apexes of each joint from the 2^nd antennomere; the 1^st antennomere less than twice as long as the 2^nd, and slightly longer than the 3^rd.

Pronotum transverse, notably wider than long, PW/PL = 1.66–1.68, widest at about middle; lateral expanded margins flat, even, feebly reflexed throughout; front and base well bordered; fore transversal impression faint, basal one evident; basal foveae small; disc markedly convex; front margin nearly straight at middle, basal margin feebly bisinuate; hind and front angles completely rounded off; base slightly shorter than front.

Elytra elongate-ovate, distinctly longer than wide, EL/EW = 1.61–1.63; nearly parallel-sided, widest at middle; basal border interrupted against the 1^st–3^rd intervals; humeri rather square; apex roundly truncate, without outer angles, inner angle broadly blunt; disc convex; striae finely and deeply striate; intervals convex, subequal in width; the 3^rd interval with median and posterior setiferous pores (anterior one wanted), both are located on middle of the interval.

Femora moderately expanded, hind femur bisetose posteriorly; for tibia dilated at apex, apical margin obtusely truncate or sinuate towards outer angle, outer margin notably serrate; middle tibia slightly curved; hind legs slender, apical spurs long and sharp, the 1^st and 3^rd tarsomeres much longer than the 2^nd and 4^th respectively, the 4^th tarsomere deeply emarginated at apex, with lobes about half as long as the joint; fore and middle tarsal claws weakly pectinate, hind claws simple.

Prosternal process bordered at apex; abdominal ventrite VII finely and shallowly emarginate at apical margin in male.

Male genitalia (Fig. 5B, C): Stout and robust, apex suddenly narrowed and pointed in lateral view; the apical lamella short, wider than long.

It is a very peculiar species, differing from other congeners by its mentum asetose, elytral base incompletely bordered, anterior dorsal pores missed on the 3^rd interval, and hind tarsal claws simple.

To refer to the type locality.

Malaysia (Sabah).