♂ (SCAU), China, Guangdong, Qingyuan, Yingde Shi, Huanghua Xiang, Yanbei Cun, Cave Yangyan Dong, 24°18'32"N, 112°47'20"E, ca 450 m a.s.l., 2014-XII-30, leg. Mingyi Tian, Weixin Liu, Sunbin Huang & Xinhui Wang.

13 ♂, 7 ♀ (SCAU), 1 ♂, 1 ♀ (ZMUM), 1 ♂, 1 ♀ (ZFMK), same locality and collecting data as the holotype.

To emphasize the collum being the broadest segment; adjective.

Keys out to the superficially most similar D. eupterygota Golovatch, Li, Liu & Geoffroy, 2012 (Liu et al. 2014), especially so concerning metatergal ornamentation and a condensed solenophore, but differs by the collum being the broadest segment, as well as the femora, postfemora and tibiae conspicuously clavate in both sexes.

Length ca 26–27 mm (♂) or 28–29 mm (♀), width of midbody pro- and metazonae 2.0 and 2.5 (♂) or 2.2 and 2.5 mm (♀), respectively. Holotype 26 mm long, 2.0 and 2.5 mm wide on midbody pro- and metazonae, respectively, maximum width on collum 3.5 mm. In width, head < segment 8–16 < 5–7 < 4 < 3 < 2 < collum; starting with segment 17, body gradually tapering towards telson (Fig. 2). Live coloration rather uniformly yellowish to pallid (Fig. 1A–B). Head sparsely setose, epicranial suture distinct (Fig. 2A–B). Antennae long and slender, reaching back until posterior margin of segment 5 (♂) or middle of segment 4 (♀) when stretched dorsally; antennomeres 5 and 6 each with a compact apicodorsal group of bacilliform sensilla.

Figure 1. 

In vivo photographs. A–B a mating couple of Desmoxytes laticollis sp. n. C–D ♂ and ♀, Desmoxytes simplipoda sp. n. E–F2 ♀, Desmoxytes similis sp. n. G ♂, Desmoxytes phasmoides sp. n. H ♂, Desmoxytes variabilis sp. n.

Figure 2. 

Desmoxytes laticollis sp. n., ♂ paratype from Cave Yangyan Dong. A–B anterior part of body, dorsal and ventral views, respectively C–D midbody segments, dorsal and ventral views, respectively E–F posterior part of body, dorsal and ventral views, respectively.

Body with 20 segments. Tegument (Fig. 2) very strongly shining, prozonae faintly microalveolate; surface below paraterga of collum and those of segments 2–4 finely microgranulate. Collum (Figs 2A, 3A) with at least 2+2 setae at front margin, another 1+1 setae in the middle, hardly visible; paraterga on collum particularly strongly developed, wing-shaped, lying a little below a rather convex collum; paraterga on segments 2–4 clearly elevated above dorsum, thereafter rather poorly-developed, relatively small and crest-shaped, lying slightly below level of a faintly convex dorsum until segment 18, vestigial and lying far below level of a strongly convex dorsum on segment 19 (Fig. 2A, C, E); paraterga 1–4 with three especially strong denticles at lateral margin, two anterior of the denticles gradually disappearing towards segment 7, but caudalmost tooth persisting until segment 18 (Fig. 2A, C, E). Metaterga 2–4 each with 1+1 setae in anterior row; each of metaterga 5–19 additionally with 1+1 setae in posterior row, mostly poorly visible; paraterga a little more strongly developed in ♂ than in ♀, calluses very thin in poreless segments, slightly thicker in pore-bearing ones. Stricture between pro- and metazonae very narrow and deep. Ozopores entirely lateral, lying on top of caudal tooth on pore-bearing paraterga (Fig. 2A, C, E). Transverse sulcus evident and deep, smooth at bottom, reaching bases of paraterga on segments 7–16, incomplete (not reaching the bases of paraterga) in segments 5–6 and 17–18, vestigial in segment 19 (Fig. 2A, C, E). Epiproct subconical, clearly flattened dorsoventrally, mostly broad, subtruncate at a narrow apex, subapical lateral setae not borne on knobs (Fig. 2E–F). Hypoproct subtrapeziform, clearly emarginate at caudal margin, caudal setae distinctly separated (Fig. 2E–F). Pleurosternal carinae poorly-developed, only visible on segments 2 and 3 both in ♂ and ♀. Axial line missing.

Figure 3. 

Desmoxytes laticollis sp. n., ♂ paratype from Cave Yangyan Dong. A collum, dorsal view B sternal processes between coxae 4, ventral view C–D right gonopod, lateral and mesal views, respectively.

Sterna sparsely setose, nearly flat, cross-impressions shallow, axial groove being especially superficial (Fig. 2B, D, F). A pair of paramedian, densely setose, low cones between ♂ coxae 4 (Figs 2B, 3B). Legs 1 short, following ones growing slightly, but increasingly long and slender towards telson, midbody legs ca 2.5 (♂) or 2.0 (♀) times as long as body height; femora, postfemora and tibiae conspicuously clavate distad, neither tarsal brushes nor adenostyles in ♂ and ♀ (Fig. 2).

Gonopods (Fig. 3C–D) suberect. Coxite short, subcylindrical, sparsely setose distodorsally, nearly 1/3 as long as telopodite. Prefemur densely setose, almost half as long as acropodite. Femorite long, slightly curved ventrad. Solenophore (sph) clearly condensed and divided into a large rectangular lobe on lateral side (= lamella lateralis, ll) and a distinct coiled part on mesal side (= lamella medialis, lm). Seminal groove running entirely on mesal side of femorite before entering onto a short, but evident and flagelliform solenomere (sl), the latter lying between ll and lm.

Because of the pallid tegument and remarkably elongated antennae and legs, this species is most probably a troglobite.

♂ (SCAU), China, Guangdong, Qingyuan, Yangshan Xian, Chengjia Xiang, Dabei Cun, Cave Kuangzhanyan, 24°46'28"N, 112°48'16"E, ca 140 m a.s.l., 2014-XII-28, leg. Mingyi Tian, Weixin Liu, Sunbin Huang & Xinhui Wang.

1 ♂, 6 ♀ (SCAU), same locality and collecting data as the holotype.

To emphasize the legs being simple, devoid of modifications; adjective.

Using the latest key (Liu et al. 2014), this new species keys out to the superficially most similar D. longispina (Loksa, 1960), especially so due to spiniform paraterga and a condensed solenophore, but differs by the legs being devoid of modifications.

All characters as in D. laticollis sp. n., except as follows.

Length ca 28–29 mm (♂) or 31–33 mm (♀), width of midbody pro- and metazonae 1.8 and 4.0 (♂) or 2.5 and 4.5 mm (♀), respectively. Holotype 29 mm long, 1.8 and 4.0 mm wide on midbody pro- and metazonae, respectively. In width, head < segment 2–4 < collum < 5–16. Coloration (Fig. 1C–D) varying from dark brownish to nearly pallid, anterior part of body a little darker than posterior part. In holotype, head, as well as dorsal and both lateral sides of metaterga dark brownish; prozonae, paraterga, sterna, and legs pallid to yellowish (Fig. 4); apices of antennomeres 6 and 7 dark brownish (Fig. 1C–D). Antennae very long and slender, reaching back until posterior margin of segment 6 (♂) or segment 5 (♀) when stretched dorsally.

Figure 4. 

Desmoxytes simplipoda sp. n., ♂ holotype from Cave Kuangzhanyan. A–B anterior part of body, ventral and sublateral views, respectively C–D midbody segments, dorsal and lateral views, respectively E–F posterior part of body, dorsal and lateral views, respectively.

Prozonae very delicately microalveolate, metazonae rather finely shagreened and microgranulate (Fig. 4). Collum (Fig. 4A) with 3+3 evident setigerous spinules at fore margin, at least 1+1 much smaller spinules in the middle and 2+2 strongly enlarged spines (paramedian spines being larger than others) at posterior margin; following metaterga 2–18 showing a pattern of smaller 2+2 posterior spinules with invariably obliterated setae (paramedian two spinules a little larger, the other two located at base of each paratergum), while metaterga 19 with 1+1 posterior spinules (Fig. 4C, E). Paraterga (Fig. 4) very strongly developed, spiniform, on collum with four evident anteromarginal denticles; all following paraterga long, straight, also spiniform, about as high as metatergal height in ♂, a little shorter in ♀; paraterga 2–18 with 2–3 evident denticles frontally. Paraterga 2–9 directed more dorsad than laterad, nearly erect above dorsum; following paraterga directed a little caudad, but ending up clearly above dorsum. Ozopores conspicuous, located a little above first denticle from lateral side of pore-bearing paraterga (Fig. 4B–F). Transverse sulcus present on segments 2–19, but complete and reaching bases of paraterga only on segments 6–15 (Fig. 4C, E).

Sterna sparsely setose, cross-impressions evident. A large, median, sparsely setose process with two small pores at base between ♂ coxae 4 (Figs 4B, 5A). Legs devoid of modifications (Fig. 4A–B), ca 2.8–3.0 (♂) or 2.5 (♀) times as long as midbody height.

Figure 5. 

Desmoxytes simplipoda sp. n., ♂ holotype from Cave Kuangzhanyan. A sternal process between coxae 4, ventral view B–C entire right gonopod and its distal half, mesal and lateral views, respectively.

Gonopods (Fig. 5B–C) simple. Coxite stout, about 1/3 as long as telopodite. Prefemur short, less than half the length of acropodite. Femorite long, suberect. Solenophore strongly condensed and divided into a large subtriangular lamella lateralis (ll) and a terminally evidently bifid lamella medialis (lm).

Judging by the extremely elongated antennae and legs, this species seems to be a troglobite.

♂ (SCAU), China, Guangdong, Qingyuan, Yingde Shi, Qingkeng Zhen, Bangjiao Cun, Cave Bangjiao Dong, 24°25'09"N, 112°57'16"E, ca 230 m a.s.l., 2014-XII-29, leg. Mingyi Tian, Weixin Liu, Sunbin Huang & Xinhui Wang.

1 ♂, 4 ♀ (SCAU), 1 ♂, 1 ♀ (ZFMK), same locality and collecting data as the holotype.

To emphasize the particular similarities to D. simplipoda sp. n.; adjective.

This species seems to be especially similar to D. simplipoda sp. n., from Yangshan, Qingyuan, Guangdong, but differs by the setose process between ♂ coxae 4 showing a large pore and the lamella medialis of the gonopod solenophore a small lobule at about midlength.

All characters as in D. simplipoda sp. n., except as follows.

Length ca 25–26 mm (♂) or 27–29 mm (♀), width of midbody pro- and metazonae 1.5 and 4.0 (♂) or 2.5 and 4.5 mm (♀), respectively. Holotype 26 mm long, 1.5 and 4.0 mm wide on midbody pro- and metazonae, respectively. In width, head < collum < segment 2–8 < 9–16. Coloration rather uniformly light brownish to pallid (Figs 1E–F, 6).

Figure 6. 

Desmoxytes similis sp. n., ♂ holotype from Cave Bangjiao Dong. A–B anterior part of body, subventral and dorsal views, respectively C–D midbody segments, subventral and dorsal views, respectively E–F posterior part of body, ventral and dorsal views, respectively.

Paraterga of collum (Fig. 6A–B) directed laterad; following paraterga 2–18 directed dorsad and a little caudad; paraterga 19 directed caudad, but all paraterga ending up clearly much above dorsum (Fig. 6). Transverse sulcus very vague, only traceable in segments 3–18 (Fig. 6B, D, F).

A large, median, setose process with a large central pore at bottom between ♂ coxae 4 (Fig. 7A). Legs devoid of modifications, about 2.5 (♂) or 2.0 (♀) times as long as midbody height.

Figure 7. 

Desmoxytes similis sp. n., ♂ paratype from Cave Bangjiao Dong. A sternal process between coxae 4, ventral view B–C entire left gonopod and its distal half, mesal and lateral views, respectively.

Gonopods (Fig. 7B–C) short. Coxite less than 1/3 the length of telopodite. Prefemur about half as long as acropodite. Femorite very short, slightly enlarged distad. Solenophore strongly condensed and divided into a large subtriangular lamella lateralis (ll) and a terminally evidently bifid lamella medialis (lm), the latter with a very small lobule (l) at about midlength.

This species seems to be very similar to D. simplipoda sp. n., from Yangshan, Qingyuan, Guangdong. The only differences are outlined in the above diagnosis. Because of the nearly pallid tegument and extremely elongated antennae and legs, this species seems to be a troglobite.

♂ (SCAU), China, Guangxi, Baise, Lingyun Xian, Jiayou Zhen, Yangli Cun, Cave Fengliu Dong, 2015-VI-9, leg. Mingyi Tian, Weixin Liu, Xinhui Wang & Mingruo Tang.

1 ♂, 1 ♀ (SCAU), 1 ♂ (ZFMK), same locality and collecting data as the holotype.

To emphasize that superficially this new species somewhat resembles a stick insect, Phasmatodea; noun in apposition.

Keys out to the superficially most similar D. minutubercula Zhang, 1986 (Liu et al. 2014), especially so due to long spiniform paraterga and a particularly condensed solenophore, but differs by a pair of rounded, setose processes present between ♂ coxae 4 and the gonopod lamella medialis showing a distinct spine.

All characters as in D. laticollis sp. n., except as follows.

Length of both sexes ca 27–29 mm, width of midbody pro- and metazonae 1.3–1.5 and 2.8–3.0 mm, respectively. Holotype 29 mm long, 1.5 and 3.0 mm wide on midbody pro- and metazonae, respectively. In width, segment 2–4 < collum < head < 5–7 < 8–18. Coloration rather uniformly light brownish, some metaterga and bases of paraterga pinkish (Fig. 1G). Antennae very long and slender, reaching back until posterior margin of segment 8 (7) (♂) or 6 (♀) when stretched dorsally.

Tegument shining and smooth, prozonae faintly microalveolate; metazonae finely microgranulate (Fig. 8). Collum (Fig. 8A) with at least 6+6 anterior, 4+4 (5) intermediate and 3+3 posterior setigerous spines; paraterga on collum spiniform, directed dorsolaterad, with a setigeous spine anteriorly at distal 1/3 (Figs 8A, 9A). Metaterga 2–4 each with 4+4 anterior, 3+3 intermediate and 5+5 posterior setigerous tubercles; metaterga 5–19 with a pattern of 5+5 setigerous tubercles anteriorly, these occasionally arranged in two transverse rows, as well as 4+4 between paraterga and at least 5+5 setigerous tubercles at posterior margin. Paraterga 2–18 (Fig. 8) extremely long, straight, spiniform, simple, usually with 2–3 very small setigerous denticles on lateral side; only paraterga 19 directed caudad. Ozopores inconspicuous, lying at base of pore-bearing paraterga on lateral side (Fig. 8D). Transverse sulcus incomplete, present on segments 6–18 (Fig. 8B, D, E).

Figure 8. 

Desmoxytes phasmoides sp. n., ♂ paratype from Cave Fengliu Dong. A–B anterior part of body, subventral and dorsal views, respectively C–D midbody segments, ventral and dorsal views, respectively E–F posterior part of body, ventral and dorsal views, respectively.

Sterna modestly setose, cross-impressions very shallow (Fig. 8C, E). A pair of paramedian, rounded, setose processes between ♂ coxae 4 (Fig. 9B). Legs long and slender, midbody legs ca 4.5 (♂) or 3.5 (♀) times as long as body height; ♂ femur 6 with a very evident apophysis at distal 1/4 (Fig. 9C).

Figure 9. 

Desmoxytes phasmoides sp. n., ♂ paratype from Cave Fengliu Dong. A collum, dorsal view B sternal processes between coxae 4, ventral view C femur 6, front view D–F entire right gonopod and its distal half, mesal, ventral and lateral views, respectively.

Gonopods (Fig. 9D–F) subfalcate. Coxite very short, less than 1/3 as long as telopodite. Prefemur short, less than half as long as acropodite. Femorite rather long, curved ventrad. Solenophore highly condensed, divided into a small, rectangular lamella lateralis and a simple lamella medialis, the latter with a distinct spine (s) at about midlength; solenomere very short and flagelliform.

Because of the pallid tegument and extremely elongated antennae and legs, this species may well be a troglobite.

♂ (SCAU), China, Guangxi, Hechi, Huanjiang Xian, near Cave Gui Dong II, Secondary forest, litter, Berlese extraction after sifting, 2007-V-18, leg. Louis Deharveng & Anne Bedos (CHIgx07-18-17).

1 ♂ (SCAU), same locality and collecting data as the holotype.

To emphasize the metaterga showing very evident, spiniform, setigerous paraterga; adjective.

Keys out to the superficially most similar D. draco Cook & Loomis, 1924 (Liu et al. 2014), judging from the ornamentation of metaterga, but differs by legs showing no modifications, in the metaterga supplied with more numerous setigerous spines, as well as the rather short femorite of the gonopod and the clearly coiled solenophore (cf. Kraus 2012).

All characters as in D. laticollis sp. n., except as follows.

Length ca 11 mm (holotype) or 12 mm (paratype), width of midbody pro- and metazonae 0.5 and 1.8 mm, respectively. In width, head < collum < segment 2–4 < 5–16. Coloration brownish to yellowish (Fig. 10). Antennae very long and slender, reaching back until posterior margin of segment 6.

Figure 10. 

Desmoxytes spiniterga sp. n., ♂ paratype from near Cave Gui Dong II. A–B body, dorsal and ventral views, respectively.

Prozonae faintly microalveolate; metazonae rather microgranulate and shagreened. Collum (Fig. 10A) with 4+4(5) anterior, 2+2 intermediate and 2+2 posterior setigerous spines; following metaterga 2–8 with 3+3 anterior and 3(4)+3(4) posterior setigerous spines; in metaterga 9–18 increasingly more numerous, with a pattern of 3(4)+3(4) anterior, 2–4+2–4 middle (behind transverse sulcus) and 5(4)+5 posterior setigerous spines, in posterior row lateral one or two spines being much larger than others; metatergum 19 with the same spination pattern, but setigerous tubercles smaller and similar in size. Paraterga (Figs 10A, 11A) very strongly developed, antler-shaped, usually three-branched, paraterga on collum with two branches; paraterga 2–8 directed more dorsad than laterad; paraterga 9–18 directed laterad, but clearly ending up above dorsum, each with an additional small denticle at last incision; paraterga 19 directed caudad. Ozopores normal, lying at base of last incision of paraterga (Fig. 11A). Transverse sulcus present on segments 3–18, incomplete (Figs 10A, 11A).

Figure 11. 

Desmoxytes spiniterga sp. n., ♂ paratype from near Cave Gui Dong II. A segments 9–10, dorsal view B sternal processes between coxae 4, ventral view C–D entire left gonopod and its distal half, mesal and lateral views, respectively.

Sterna moderately setose, cross-impressions shallow (Fig. 10B). A pair of paramedian processes between ♂ coxae 4 (Figs 10B, 11B). Legs without modifications, midbody legs ca 2.8 times (♂) as long as body height.

Gonopods (Figs 10B, 11C–D) with coxite about 1/3 as long as telopodite. Prefemur almost half as long as acropodite. Femorite short and slender, slightly enlarged distad. Solenophore clearly coiled and divided into a large spiniform lamella lateralis and a very distinct and coiled lamella medialis. Solenomere relatively long.

Compared to cave-dwelling congeners, this species is much smaller and darker.

♂ (SCAU), China, Guangxi, Hechi, Fengshan Xian, Fengcheng Zhen, Cave Huoji Dong, 24°28'32"N, 107°03'28"E, ca 500 m a.s.l., 2015-VIII-3, leg. Jujian Chen, Xinhui Wang & Mingruo Tang.

2 ♂, 3 ♀ (SCAU), same locality and collecting data as the holotype. 2 ♂, 1 ♀ (SCAU), 1 ♂, 1 ♀ (ZFMK), same County, Zhaiya Xiang, Laying Cun, Cave II Dong, 2015-VIII-4, leg. Jujian Chen, Xinhui Wang & Mingruo Tang. 3 ♂, 2 ♀ (SCAU), same County, Jinya Xiang, Hangdong Cun, Cave I Dong, 2014-VI-14, leg. Mingyi Tian, Weixin Liu, Haomin Yin & Xiaozhu Luo. 3 ♂, 2 ♀ (SCAU), Lingyun Xian, Sicheng Zhen, Wuzhishan, Cave Qianlongya, 2015-VI-10, leg. Mingyi Tian, Weixin Liu, Xinhui Wang & Mingruo Tang. 1 ♂, 1 ♀ (SCAU), Bama Xian, Yandong Xiang, Namen Cun, Cave Baiyan Dong, 2015-VIII-3; 3 ♂, 2 ♀ (SCAU), Tian’e Xian, Bala Xiang, Gandong Cun, Cave number VIII Dong, 2015-VIII-8. leg. Jujian Chen, Xinhui Wang & Mingruo Tang. 3 ♂, 3 ♀ (SCAU), China, Guangxi, Hechi, Huanjiang Xian, Mulun, way to Mashan Dong, 2007-V-21, leg. Louis Deharveng & Anne Bedos (CHIgx07-21-02). 2 ♂, 1 ♀ (SCAU), China, Guangxi, Hechi, Huanjiang Xian, near Midong Cun, 2007-V-23, leg. Louis Deharveng & Anne Bedos (CHIgx07-23-05).

To emphasize the metaterga showing a variable pattern of setigerous spines; adjective.

This species seems to be especially similar to D. nodulosa Liu, Golovatch & Tian, 2014, from Cave II, Xiao’ao Xiang, Du’an Xian, Hechi, Guangxi, because both share very close patterns of metatergal ornamentation and particularly stout gonopods, but differs by the metaterga showing a variable pattern of setigerous spines, and the particularly complex gonopod solenophore.

All characters as in D. laticollis sp. n., except as follows.

Length ca 17–21 (♂) or 20–24 mm (♀), width of midbody pro- and metazonae 1.0–1.2 and 2.8–3.8 (♂) or 1.5–1.8 and 3.0–4.0 mm (♀), respectively. Holotype 19 mm long, 1.0 and 2.8 mm wide on midbody pro- and metazonae, respectively. In width, head < collum < segment 2–4 < segment 5–16. Coloration (Figs 1H, 12, 14–15) varying from dark to light brownish; paraterga and posterior parts of metaterga finely yellow-brownish. Antennae long and slender, reaching back until posterior margin of segment 5 (♂) or segment 4 (♀) when stretched dorsally (Fig. 12A–B).

Figure 12. 

Desmoxytes variabilis sp. n., ♂ holotype from Cave Huoji Dong. A–B anterior part of body, lateral and ventral views, respectively C legs 5–7, ventral view D–F midbody segments, lateral, ventral and dorsal views, respectively G–I posterior part of body, lateral, ventral and dorsal views, respectively.

Prozonae faintly microalveolate; metazonae rather microgranulate and shagreened. Collum (Figs 12B, 13B) with 4+4 (or 5+5) anterior and 2+2 (or 3+3) posterior setigerous spines. Metaterga 2–18 (Figs 12–13) each with 2+2 (or 3+3) anterior and 2+2 (or 3+3) posterior setigerous spines, lateral spine of posterior row being much larger than others; metatergum 19 with 3+3 anterior and 2+2 posterior, similar, small, setigerous tubercles. Paraterga very strongly developed, antler-shaped, usually three-branched, each tip with an evident lateral seta (Figs 12–13). Paraterga of collum stout, two-branched; paraterga 2–5 directed more dorsad than laterad; paraterga 19 directed caudad, horizontal, lying level to dorsum; paraterga 6–18 directed obliquely upwards at ca 45°, in ♂ ending up clearly above dorsum (Figs 12–13), but paraterga slightly lower, shorter, subhorizontal and lying level to dorsum in ♀. Ozopores conspicuous, located at last incision of poriferous paraterga (Fig. 14A). Transverse sulcus obscure on collum and metaterga 2–4; more evident, but incomplete on metaterga 5–17 (Figs 12–13).

Figure 13. 

Desmoxytes variabilis sp. n., ♂ paratype from near Midong Cun. A–B anterior part of body, dorsal and ventral views, respectively C–D midbody segments, dorsal and ventral views, respectively E–F posterior part of body, dorsal and ventral views, respectively.

Sterna sparsely setose, cross-impressions very shallow (Figs 12–13). A paramedian pair of separated, short, rounded, poorly setose processes between ♂ coxae 4 (Fig. 14B). Legs long and slender, ca 2.5–2.8 (♂) or 2.0–2.2 (♀) times as long as body height; ♂ femora 5–7 each with a conspicuously densely pilose apophysis ventrally at about midlength (Figs 12C, 13A–B, 14C–E).

Figure 14. 

Desmoxytes variabilis sp. n., ♂ paratype from Cave Huoji Dong. A segment 10, dorsal view B sternal processes between coxae 4, ventral view C–E femora 5–7, front view F–G entire right gonopod and its distal half, mesal and lateral views, respectively.

Gonopods (Fig. 14F–G) very short. Coxite less than 1/3 as long as telopodite. Prefemur less than half the length of acropodite. Femorite stout, apically with a distinct sulcus. Solenophore quite complex and compact, divided into two well differentiated lobes, a higher, bipartite and apically acuminate lamina medialis, plus a lower and curved lamina lateralis; solenomere short and flagelliform.

This obviously troglophilic species is rather eurytopic, occurring both outside and inside caves. It shows a remarkably variable pattern of spination on collum and metaterga (Figs 12–13), but the gonopod structure remains stable.