COLOMBIA. Valle del Cauca. Buenaventura. Vereda El Salto. Pericos Natural Reserve, 350 m., 3°56'N, 76°47'W.

Holotype male, 9–11.VIII.2013. light trap, O. Saenz, N. Calderón. Deposited in the Entomology Museum, Universidad del Valle, Santiago de Cali, Colombia (MUSENUV slide No. 25775).

It is our pleasure to dedicate this species to Nadia Calderón, a graduate student at the Universidad del Valle, who, together with Oscar Saenz Manchola, collected the specimens of the two species of Edmockfordia here described.

Phallosome with three posterior projections, with posterior end broadly W shaped; external parameres short, little developed; aedeagal arch slightly projected posteriorly; epiproct broadly rounded posteriorly.

(in 80% ethanol). Body mostly pale brown, with creamy areas, as indicated below, pronotum, propleura and metapleura brown, upper half of mesopleura brown, lower half creamy; meso- and metanotal lobes creamy, bordered with brown. Abdomen creamy, clunium brown. Head (Fig. 2), with a brown transverse band between compound eyes enclosing the ocellar triangle; vertex creamy; genae brown; ocelli hyaline, with dark brown centripetal crescents; postclypeus brown, anteclypeus centrally brown, sides creamy; labrum pale brown anteriorly, fading towards the posterior margin. Antennae with scape and pedicel brown, flagella pale brown. Maxillary palpomeres 1–4 brown. Legs: front coxa, and trochanters of all legs creamy; coxae of mid and hind leg with a dark brown spot on outer border; femora mostly creamy, with a brown spot on proximal and distal ends; tibiae and tarsi pale brown. Forewings hyaline, pterostigma with a brown spot distally, and a brown band proximally; veins brown, each with a brown spot distally, at wing margin; a brown spot at nodulus. Hindwings hyaline, veins pale brown, with brown spots distally, at wing margin. Epiproct and paraprocts creamy. Hypandrium creamy, with postero-lateral corners brown.

Figures 1–5. 

Edmockfordia calderonae sp. n. Male. 1 Forewing and hindwing 2 Front view of head 3 Epiproct and paraprocts 4 Hypandrium 5 Phallosome. Scales in mm.

As in diagnosis, plus the following: outer cusp of lacinial tip broad, with eight denticles; compound eyes with interommatidial setae, mostly dorsally. Forewings (Fig. 1) symmetric, pterostigma elongate, rounded, widest in the middle; Rs with two branches, M dichotomously branched, resulting in four branches; areola postica low, apically rounded. Hindwings symmetric (Fig. 1), Rs of two branches, M simple. Coxae of hind leg without Pearman’s organ; trochanters of all legs dorsally with two long setae, tarsi of front legs without ctenidobothria, mid legs with 21 ctenidobothria on t1, t2 with two ctenidobothria. Hypandrium (Fig. 4): symmetric, setose, posterior border convex. Phallosome (Fig. 5): open anteriorly, side struts long, slender, sclerotized, external parameres short; aedeagal arch slightly projected posteriorly in the middle; endophallus membranous, without sclerites. Paraprocts (Fig. 3): elongate, ovoid, with short setae and a field of microsetae along inner margin, sensory fields with 22–23 trichobothria on basal rosettes. Epiproct (Fig. 3): trapeziform, broadly rounded posteriorly, setose, with a field of microsetae along posterior border; a field of papillae mesally, next to posterior border.

FW: 2675, HW: 2000, F: 625, T: 1125, t₁: 500, t₂: 120, ctt₁: 30, f₁: 550, f₂: 440, IO: 260, D: 240, d: 300, IO/d: 0.86, PO: 1.25.

COLOMBIA. Valle del Cauca. Buenaventura. Vereda El Salto. Pericos Natural Reserve, 350 m., 3°56'N, 76°47'W.

Holotype male, 28–29.III.2013. Light trap, O. Saenz, N. Calderón. Deposited in the Entomology Museum, Universidad del Valle, Santiago de Cali, Colombia (MUSENUV, slide code No. 25774).

2 male, COLOMBIA. Valle del Cauca. Buenaventura. Vereda El Danubio, 340 m., 3°36'58.8"N, 76°53'59.5"W. 28–29.VIII.2014. Light trap. R. González, O. Saenz, N. Calderón. Paratypes deposited in the Entomology Museum, Universidad del Valle, Santiago de Cali, Colombia (MUSENUV, slide code No. 26135–26136).

We take pleasure to dedicate this species to Oscar Saenz Manchola, one of its collectors, a graduate student at the Universidad del Valle, Santiago de Cali, Colombia.

Differing from E. calderonae and from E. chiquibulensis in having the phallosome aedeagal arch apically rhomboid, in having the external parameres extremely long, falcate, reaching the level of the aedeagal apex, and in having the postero-lateral corners of the epiproct rounded, protuberant.

(in 80% ethanol). Body mostly brown, with creamy areas, as indicated below, pronotum and propleura brown; meso- and metanotal lobes creamy, bordered with brown, upper halves of meso- and metapleura brown, lower halves creamy. Abdomen creamy. Head (Fig. 7), with a broad, transverse, dark brown band between compound eyes, enclosing the ocellar group; vertex creamy; genae proximally creamy, distally brown; ocelli hyaline, with ochre centripetal crescents; postclypeus brown, anteclypeus brown in the center, creamy on the sides; labrum creamy. Antennae with scape and pedicel brown, flagella pale brown. Maxillary palpomeres 1–4 brown. Legs: coxae and trochanters creamy; femora mostly creamy, with a brown spot on proximal and distal ends; tibiae and tarsi pale brown. Forewings hyaline, pterostigma with a brown spot at the apex, and a brown band proximally, veins pale brown, each with a brown spot distally, at wing margin, a brown spot at confluence of Cu₂ and A. Hindwings hyaline, veins pale brown, each with a brown spot distally, at wing margin. Clunium brown, epiproct pale brown, with sides dark brown; paraprocts pale brown, hypandrium almost unpigmented.

Figures 6–10. 

Edmockfordia saenzi sp. n. Male. 6 Forewing and hindwing 7 Front view of head 8 Epiproct and paraprocts 9 Hypandrium 10 Phallosome. Scales in mm.

As in diagnosis, plus the following: outer cusp of lacinial tip broad, with seven denticles; compound eyes with inter-ommatidial setae, mostly dorsally. Forewings (Fig. 6) asymmetric, pterostigma elongate, rounded, widest in the middle; Rs two branched; M dichotomously forked, resulting in four branches, right wing with M₄ incompletely forked; areola postica low, apically rounded; left wing with open areola postica, vein Cu1A arising from M. Hindwings (Fig. 6) symmetric: Rs of two branches, M simple. Coxae of hind leg without Pearman organ, trochanters of all legs dorsally with two long setae; tarsi of front legs without ctenidobothria, t1 of mid legs with 11 ctenidobothria, t2 without ctenidobothria. Hypandrium (Fig. 9), symmetric, setose, with posterior border obtusely concave. Phallosome (Fig. 10): open anteriorly, V-shaped, side struts broad, dilated proximally and curved outwards. Paraprocts (Fig. 8): elongate, ovoid, with short setae, and a field of microsetae along the inner border, sensory fields with 30–31 trichobothria on basal rosettes. Epiproct (Fig. 8): broad, trapeziform, with field of short setae and a field of microsetae along posterior border; a field of papillae mesally, next to posterior border.

Of the two paratypes for this species, one of them presents a forewing vein pattern with a four branched M, as described for the genus, however, a second specimen present the distal third of the forewing reticulated (Fig. 11) (e. g. R4+5 and M1 connected by a cross vein, R4+5 distally forked, a vein arising from the middle of R4+5 forked at wing margin, with the second branch fused at wing margin with M1, forming a closed cell, M1 absent, and M3 branched, with the branch near the areola postica incomplete.

Figure 11. 

Anomalous forewing venation in E. saenzi sp. n. Scale in mm.

FW: 2700, HW: 2025, F: 675, T: 1150, t₁: 500, t₂: 140, ctt₁: 26, f₁: 540, f₂: 470, IO: 280, D: 180, d: 280, IO/d: 1, PO: 1.5.

(19 years, three months in 80% ethanol). Essentially as in the male (see García Aldrete 2009). Scape and pedicel brown, flagella pale brown. Legs pale brown.

Wings (Fig. 12), distal labral sensilla and pretarsal claws as in the male (see García Aldrete 2009). Outer cusp of lacinial tip with seven denticles. Labral sclerites joined distally by a sclerotized band (Fig. 13). Subgenital plate (Fig. 15) broad, with field of short setae; pigmented area deeply concave anteriorly; posterior border rounded, a strongly sclerotized, concave band next apex, and a transverse crease anterior to it. Gonapophyses (Fig. 16) complete, v1 slender, about half the length of v2+3, joined to clunium by a membranous band. v2+3 wide based proximally, with a short, pointed “heel”; v3 a well defined, long slender lobe on side of v2, bearing eight setae; distal process almost straight, acuminate, with a field of microsetae proximally. Paraprocts (Fig. 14) broadly semicircular, with setae as illustrated, and a field of microsetae next distal border. Sensory fields elliptic, with 21 trichobothria on basal rosettes. Epiproct (Fig. 14) trapeziform, with field of setae on distal half, and three mesal macrosetae next anterior border.

Figures 12–16. 

Edmockfordia chiquibulensis Female. 12 Forewing and hindwing 13 Front view of head 14 Epiproct and paraprocts 15 Subgenital plate 16 Gonapophyses and IX sternum. Scales in mm.

FW: 2434, HW: 1784, F: 555, T: 998, t1: 449, t2: 113, ctt1: 27, f1: 608, f2: 493, IO: 343, D: 227, d: 159, Mx4: 184, IO/d: 2.15, PO: 0.46.

Belize. Cayo District. Chiquibul Forest Reserve. New María, 24.III.1995, 640 m. Malaise trap, paratype female. 17–20.I.1995, flight interception trap, female paratype. San Pastor, 560–580 m. 23–26.III.1995, paratype female. All specimens collected by T. King & A. Howe.