Research Article |
Corresponding author: Abraham S.H. Breure ( bbreure@xs4all.nl ) Academic editor: Eike Neubert
© 2015 Abraham S.H. Breure, Jonathan D. Ablett.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Breure ASH, Ablett JD (2015) Annotated type catalogue of the Megaspiridae, Orthalicidae, and Simpulopsidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London. ZooKeys 470: 17-143. https://doi.org/10.3897/zookeys.470.8548
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The type status is described for 65 taxa of the Orthalicoidea, classified within the families Megaspiridae (14), Orthalicidae (30), and Simpulopsidae (20); one taxon is considered a nomen inquirendum. Lectotypes are designated for the following taxa: Helix brephoides d’Orbigny, 1835; Simpulopsis cumingi Pfeiffer, 1861; Bulimulus (Protoglyptus) dejectus Fulton, 1907; Bulimus iris Pfeiffer, 1853. The type status of Bulimus salteri Sowerby III, 1890, and Strophocheilus (Eurytus) subirroratus da Costa, 1898 is now changed to lectotype according Art. 74.6 ICZN. The taxa Bulimus loxostomus Pfeiffer, 1853, Bulimus marmatensis Pfeiffer, 1855, Bulimus meobambensis Pfeiffer, 1855, and Orthalicus powissianus var. niveus
Megaspiridae , Orthalicidae , Simpulopsidae , types
This is the fourth paper on the types of Orthalicoidea in the Natural History Museum, London. Earlier papers (
References are given to the original publication, plus those of following papers where type material has been mentioned or is (re-)figured. Dates of publication are in accordance with
This systematic list follows
elata Gould, 1847.
achilles Pfeiffer, 1853; ascendens Pfeiffer, 1853; buckleyi Higgins, 1872; consimilis Reeve, 1848; foveolatus Reeve, 1849; hartwegi Pfeiffer in Philippi, 1846; inca d’Orbigny, 1835; insolitus Preston, 1909; integer Pfeiffer, 1855; loxostomus Pfeiffer 1853; magnificus Grateloup, 1839; plumbeus Pfeiffer, 1855; requieni Pfeiffer, 1853.
Molecular studies (
magnifica Pfeiffer, 1848; phoebus Pfeiffer, 1863; victor Pfeiffer, 1854.
gracilis E.A. Smith, 1902.
indentatus da Costa, 1901; thompsonii Pfeiffer, 1845; yanamensis Morelet, 1863.
murrea Reeve, 1849.
bensoni Reeve, 1849; bifulguratus Reeve, 1849; boucardi Pfeiffer, 1860; mars Pfeiffer, 1861; phlogera d’Orbigny, 1835.
subirroratus da Costa, 1898.
approximata Fulton, 1896; iris Pfeiffer, 1853; irroratus Reeve, 1849; saturanus Pfeiffer, 1860.
salteri Sowerby III, 1890.
Both this taxon described by Strebel and the next one (Scholvienia Strebel, 1910), previously treated as subgenera of Thaumastus (
alutaceus Reeve, 1849; brephoides d’Orbigny, 1835; porphyrius Pfeiffer, 1847.
deburghiae Reeve, 1859; fraseri Pfeiffer, 1858; gloriosus Pfeiffer, 1862; kelletti Reeve, 1850; niveus Preston, 1909; vicaria Fulton, 1896; yatesi Pfeiffer, 1855.
meobambensis Pfeiffer, 1855.
clouei Pfeiffer, 1857; demerarensis Pfeiffer, 1861; jeffreysi Pfeiffer, 1852; obliquus Reeve, 1849; opalinus Sowerby I, 1833; sarcochilus Pfeiffer, 1857; subtuszonata Pilsbry, 1899.
hyaloideus Pfeiffer, 1855; ovulum Reeve, 1849.
ephippium Ancey, 1904; marmatensis Pfeiffer, 1855.
aenea Pfeiffer, 1861; corrugatus Guppy, 1866; cumingi Pfeiffer 1861; decussata Pfeiffer, 1857; gomesae da Silva & Thomé, 2006; miersi Pfeiffer, 1857; salomonia Pfeiffer, 1953; simulus Morelet, 1851; vincentina E.A. Smith, 1895.
dejectus Fulton, 1907.
Bulimus achilles
Thaumastus (Thaumastus) taunaisii achilles;
[Brazil] “in ripis fluvii Amazonum”.
“Banks of Amazon”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.
“Long. 57, diam. 25 mill.”. Figured specimen H 58.0, D 25.5, W 6.4.
NHMUK 1975268, lectotype, 1975269, 2 paralectotypes (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based, but the species was described from the Cuming collection. The original label also mentions “between B. Largilliertii / + taunasii”; a label in a second (later?) hand has added “Prov. S. Paulo, Brazil / (Nehring)”.The current systematic position is according to
Megaspiridae, Thaumastus taunaisii (Férussac, 1822).
Simpulopsis aenea
“Parada, reipublicae mexicanae (Sallé)”.
“Parada, Mexico, Mr Sallé”, taxon label in Pfeiffer’s handwriting. M.C. label style III.
“Diam. maj. 9, min. 8, alt. 5 1/2 mill.”. Figured specimen H 8.98, D 10.6, W 2.5.
NHMUK 20140830, three syntypes, Sallé leg. (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based. The current systematic position follows
Simpulopsidae, Simpulopsis (Simpulopsis) aenea Pfeiffer, 1861.
Bulimus alutaceus
Strophocheilus alutaceus;
Thaumastus (Scholvienia) alutaceus;
“Cuzco, Bolivia; W. Lobb”.
“Cuzco”. M.C. label style III, V.
Not given. Figured specimen H 35.5, D 16.5, W 6.6.
NHMUK 1975148, lectotype, 1975149, one paralectotype. W. Lobb leg. (Cuming coll.).
Reeve did not state on how many specimens his description was based. As
?Orthalicidae, Scholvienia alutacea (Reeve, 1849).
Porphyrobaphe approximata
[Colombia] “Bogota”.
“Bogota”, in Fulton’s handwriting.
“Long. 67 millim., maj. diam. 31 millim.”. Figured specimen H 65.7, D 32.5, W 6.5.
NHMUK 1895.12.19.44, one syntype (ex Fulton).
Fulton did not state on how many specimens his description was based; the single specimen found corresponds to his figure and is herein considered as syntype. The current systematic position is according to
Orthalicidae, Porphyrobaphe (Porphyrobaphe) approximata Fulton, 1896.
Bulimus ascendens
Thaumastus (Thaumastus) ascendens;
Thaumastus ascendens;
“Brasilia”.
“Brazils”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.
“Long. 95 mill., diam. 34 mill.”. Figured specimen H 92.0, D 39.0, W –.
NHMUK 1975274, lectotype (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based, but the species was described from the Cuming collection. The top whorls of the specimen are missing. The current systematic position is according to
Megaspiridae, Thaumastus ascendens (Pfeiffer, 1853).
Bulimus bensoni
Oxystyla bensoni;
“Banks of the river Amazon”.
“Brazil”. M.C. label style I, V.
Not given. Figured specimen H 66.6, D 35.0, W 7.9.
NHMUK 1975582, three possible syntypes (Cuming coll.).
Reeve described this taxon from “Mus. Benson”, but did not state on how many specimens his description was based. According to
Orthalicidae, Orthalicus bensoni (Reeve, 1849).
Bulimus bifulguratus
Oxystyla bifulgurata;
Orthalicus bifulguratus;
[Colombia] “Andes of Columbia”.
“Andes of Colombia”. M.C. label style I, V.
Not given. Figured specimen H 56.9, D 32.8, W 5.8.
NHMUK 20140082, lectotype (Cuming coll.).
Reeve did not state on how many specimens his description was based. The lectotype is not full-grown as shown by the lip.
Orthalicidae, Orthalicus bifulguratus (Reeve, 1849).
Orthalicus boucardi
“Mexico (Mr. Boucard)”.
“Betaza Mexico / Mr. Boucard”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.
“Long. 43, diam. 25–26 mill.”. Figured specimen H 54.8, D 29.7, W 6.9.
NHMUK 20140081, three syntypes, Boucard leg. (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based. The specimens found are larger than the measurements given by Pfeiffer, but undoubtedly were collected by Mr. Boucard, bear Pfeiffer’s handwriting, and are considered syntypes herein. The type locality (Oaxaca, Sierra de Betaza) was specified by
Orthalicidae, Orthalicus boucardi (Pfeiffer, 1860).
Helix brephoides
Bulimus brephoides;
Strophocheilus brephoides;
“republica Peruviana”.
“Pérou”, in d’Orbigny’s handwriting.
“Longit. 52 millim., latit. 25 millim.”. Figured specimen H 51.9, D 25.1, W 5.6.
NHMUK 1854.12.4.117, lectotype (d’Orbigny coll.).
d’Orbigny did not state on how many specimens his description was based. The specimen found corresponds to the figures of d’Orbigny and is here designated lectotype (design. n.) to define the taxon, which has been compared to Bulimus taeniolus Nyst, 1845 (
?Orthalicidae, Scholvienia brephoides (d’Orbigny, 1835).
Orthalicus (Porphyrobaphe) buckleyi
Orthalicus (Methorthalicus) buckleyi;
Thaumastus (Thamastus) buckleyi;
[Ecuador, Prov. Loja] “San Lucas”.
“Ecuador”.
“Long. 93, lat. 36 mill.”. Figured specimen H 92.0, D 36.0, W 6.0.
NHMUK 1872.5.22.6, two syntypes (da Costa coll.).
Higgins did not state on how many specimens his description was based. Of the two syntypes mentioned by
Megaspiridae, Thaumastus buckleyi (Higgins, 1872).
Bulimus clouei
Drymaeus clouei;
Leiostracus (Leiostracus) clouei;
Leiostracus clouei;
“Brazil (Mr. Cloué)”.
“Brazils Mons Cloué”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.
“Long. 22, diam. 10 mill.”. Figured specimen H 22.2, D 11.1, W 7.2.
NHMUK 1975491, lectotype; 1975492, one paralectotype, Cloué leg. (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based. The current systematic position follows
Simpulopsidae, Leiostracus clouei (Pfeiffer, 1857).
Bulimus consimilis
“—?”.
“Brazil”. M.C. label style I, V.
Not given. Figured specimen H 52.9, D 22.8, W 6.5.
NHMUK 20030189, three syntypes (Cuming coll.).
Reeve did not state on how many specimens his description was based, but wrote “[t]his shell approaches nearest to the B. Taunaisii, but is certainly distinct”. Pfeiffer (1853: 406) considered this taxon a junior subjective synonym of Bulimus largillierti Philippi, 1842, which has been followed by later authors. The printed label also mentions this name, and this is consistent with the index (
Megaspiridae, Thaumastus largillierti (Philippi, 1842).
Simpulopsis corrugatus
“Trinidad”.
“Trinidad”, presumably in Guppy’s handwriting.
“Height 0.38 inch, greatest breadth 0.47 inch [H 9.65, D 11.9 mm]”. Figured specimen H 9.27 (damaged), D 10.8, W 3.5.
NHMUK 1866.1.3.7, one syntype (ex Guppy).
Guppy did not state on how many specimens his description was based; the single specimen found is damaged. Guppy emendated the name “qu’il me paraît préférable de féminiser, à l’exemple de Pfeiffer et de la plupart des auteurs” (
Simpulopsidae, Simpulopsis (Simpulopsis) corrugata Guppy, 1866.
Simpulopsis cumingi
“Mexico”.
“Mexico”, taxon label in Pfeiffer’s handwriting. M.C. label style III.
“Diam. maj. 20 1/2, alt. 12 mill.”. Figured specimen H 14.1, D 19.0, W 3.4.
NHMUK 1975486, lectotype and one paralectotype (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based. Two specimens have been found in the collection, of which one is designated lectotype (design. n.) to fixate this poorly understood species. This taxon was compared to Simpulopsis aenea by
Simpulopsidae, Simpulopsis (Simpulopsis) cumingi Pfeiffer, 1861.
Bulimus deburghiae
Sultana (Metorthalicus) deburghiae;
“Peruvian side of the Amazon”.
“Banks of Amazon River / (Reeve)”, in Dance’s handwriting; see below.
“Long. 2 3/4 in. Lat. 1 1/4 in. [H 69.9, D 31.8 mm]”. Figured specimen H 64.7, D 33.6, W 6+.
NHMUK 19601622, lectotype (ex DeBurgh).
Reeve wrote “[a] fine shell”, but otherwise it is not clear from the context that he had only one specimen at hand. The material is accompanied by a label written in 1961 by S.P. Dance “This specimen does not suit Reeve’s measurements but it is labelled by Mrs. de Burgh”; his selection as lectotype was interpreted as such by
Orthalicidae, Sultana (Metorthalicus) deburghiae (Reeve, 1859).
Simpulopsis decussata
[Brazil] “Petropolis prope Rio Janeiro (Miers)”.
“Petropolis Rio / F. Miers E[sq.]”, taxon label in Pfeiffer’s handwriting. M.C. label style III.
“Diam. maj. 12 1/2, alt. 11 mill.”. Figured specimen H 14.3, D 12.5, W 4.7.
NHMUK 1975488, lectotype (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based, but described this taxon from “Mus. Cuming”. This is contrasting the statement in
Simpulopsidae, Simpulopsis (Simpulopsis) decussata Pfeiffer, 1857.
Bulimus demerarensis
Drymaeus demerarensis;
Leiostracus (Leiostracus) demerarensis;
Bostryx demerarensis;
[Guiana] “Demerara”.
“Demerara”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.
“Long. 20 1/2, diam. 10 mill.”. Figured specimen H 20.1, D 10.9, W 6.5.
NHMUK 1975501, lectotype (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based; only a single specimen was found.
Simpulopsidae, Leiostracus demerarensis (Pfeiffer, 1861).
Pupa (Megaspira) elata
Megaspira elata;
“Brazil”.
“Brazil (Gould)”.
“Long. 1 1/2, lat. 1/3 poll. [H 38.0, D 8.4 mm]”. Figured specimen H 37.2, D 8.4, W 18.1.
NHMUK 1987060, three paralectotypes (ex Gould).
Gould did not state on how many specimens his description was based. The lectotype is USNM 5503 (
Megaspiridae, Megaspira elata (Gould, 1847).
Bulimulus ephippium
Bulimulus ephippium;
“Bahia, Brazil (teste H. Fulton)”.
“Bahia”, in Fulton’s handwriting.
“Longit. 20, diam. 12 mill.”. Figured specimen H 20.5, D 13.3, W 5.3.
NHMUK 1905.12.30.12, one paralectotype.
Ancey did not state on how many specimens his description was based; the NHMUK specimens were considered syntypes by
Simpulopsidae, Simpulopsis (Eudioptus) ephippium Ancey, (1904).
Bulimus mahogani
Bulimus foveolatus
Strophocheilus foveolatus;
“Vitoe, near Sarma [sic, Tarma], Alto-Peru; W. Lobb”.
“Peru”. M.C. label style IV, V.
“Long. 3 poll., diam. 15 lin. [H 76.0, D 31.7 mm]”; see remarks. Figured specimen H 71.5, D 37.0, W 5.7.
NHMUK 1975275, lectotype; 1975276, one paralectotype (Cuming coll.).
Pfeiffer, in his original description, referred to Sowerby I 1838 in
Megaspiridae, Thaumastus foveolatus (Reeve, 1849).
Bulimus fraseri
Orthalicus fraseri;
Sultana (Metorthalicus) fraseri;
“in provincia Cuenca reipublicae Aequatoris (Fraser)”.
“Found on the road from Gualaquiza / to Mendez— and near to the latter / place”, “Province of Cuenca / Republic of Ecuador / Mr Fraser”, taxon label in Pfeiffer’s handwriting. M.C. label style III.
“Long. 89, diam. 37 mill.”. Figured specimen H 88.9, D 45.0, W 6.4.
NHMUK 20140083, lectotype, Fraser leg. (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based, but described this taxon from “Mus. Cuming”. The current systematic position follows
Orthalicidae, Sultana (Metorthalicus) fraseri (Pfeiffer, 1858).
Bulimus gloriosus
Sultana (Metorthalicus) deburghiae (Reeve);
“Republic of Ecuador”.
“Republic Ecuador / Mr Fraser”, taxon label in Pfeiffer’s handwriting. M.C. label style III.
“Long. 78, diam. 34 mill.”. Figured specimen H 75.2, D 39.3, W 5.7+.
NHMUK 1975243, lectotype (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based; the single specimen found has the top damaged. The lectotype designation by
Orthalicidae, Sultana (Metorthalicus) deburghiae (Reeve, 1859).
Simpulopsis gomesae
“Brasil, Rio Grande do Sul, São Francisco de Paula”.
No locality.
Not given (range H 1.60–10.96, D 1.55–8.63 mm). Figured specimen H 6.46, D 6.93, W 3.5.
NHMUK 20050238, one paratype in ethanol, J.W. Thomé leg.
This taxon was based on 17 specimens; the specimen present in NHMUK was mentioned in the original paper. Its systematic position may, however, need to be critically re-examined as many taxa have already been described from this region.
Simpulopsidae, Simpulopsis (Simpulopsis) gomesae da Silva & Thomé, 2006.
Corona pfeifferi gracilis
“Rio Caqueta, S.E. Colombia”.
“Rio Caqueta, / S.E. Colombia”, in Smith’s handwriting.
“[L]ength is 67 mm. and diameter 23”. Figured specimen H 67.3, D 24.8, W 8.8.
NHMUK 1902.5.27.4, holotype.
This taxon was described from a single specimen. The morphological variation within Corona pfeifferi needs further study.
Orthalicidae, Corona pfeifferi gracilis E.A. Smith, 1902.
Bulimus hartwegi Pfeiffer in
Strophocheilus hartwegi;
Thaumastus (Thaumastus) hartwegi;
“respublica [sic] Aequatoris, ubi ad ‘El Catamaija’ prope Loxa Hartweg”.
“El Catamaja near Loxa”. M.C. label style IV.
“Long 28, diam. 13´´´ [H 61.0, D 28.3 mm]”. Figured specimen H 57.0, D 30.0, W 4.8.
NHMUK 1975126, one syntype (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based, but said the material was in “Sammlung des Hrn. Hugh Cuming”. Given the context of the publication, it is here assumed that the dimensions were given in German lines (1 line = 2.18 mm);
Megaspiridae, Thaumastus hartwegi (Pfeiffer in Philippi, 1846).
Bulimus hyaloideus
Rhinus constrictus (Pfeiffer);
Rhinus hyaloideus;
“Mendez, Andes of New Granada”.
“Mendes Andes of Granada”, taxon label in Pfeiffer’s handwriting. M.C. label style I.
“Long. 12 1/2, diam. 7 1/2 mill.”. Figured specimen H 20.6, D 11.1, W 6.7.
NHMUK 1975412, lectotype; 1975413, one paralectotype (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based; he described this taxon from Cuming’s collection. The paralectotype specimen is juvenile.
Simpulopsidae, Rhinus constrictus (Pfeiffer, 1841).
Helix inca
Bulimus inca;
Strophocheilus inca;
Thaumastus (Atahualpa) inca;
“Tutulima (republica Boliviana)”.
“Tutulima, Bolivia”, in d’Orbigny’s handwriting.
“Longit 72 millim., latit. 30 millim.”. Figured specimen H 75.4, D 32.3, W 8.3.
NHMUK 1854.12.4.116, lectotype and three paralectotypes (d’Orbigny coll.).
d’Orbigny did not state on how many specimens his description was based. The lot found consists of four specimens, of which the one corresponding to d’Orbigny’s figure is now designated lectotype (design. n.) to fixate the taxon, which needs further study to clarify its status; the three paralectotypes are one subadult and two juveniles. Three other specimens are in the MNHN collection (
Megaspiridae, Thaumastus inca (d’Orbigny, 1835).
Strophocheilus (Dryptus) indentatus
Thaumastus (Thaumastus) indentatus;
“Ecuador”.
“Ecuador”, in da Costa’s handwriting.
“Long. 44, diam. 23 mm.”. Figured specimen H 44.0, D 24.0, W 4.8.
NHMUK 1907.11.21.115, lectotype; 1907.11.21.116, one paralectotype (da Costa coll.).
da Costa did not state on how many specimens his description was based. This species has been classified by Thaumastus s.str. by
Orthalicidae, Kara indentata (da Costa, 1901) (comb. n.).
Bulimus (Thaumastus) insolitus
“Chanchamayo, Peru”.
“Chanchamayo Peru”.
“Alt. 70, diam. maj. 29.5 mm.”. Figured specimen H 70.4, D 31.2, W 5.6.
NHMUK 1947.3.11.1, holotype (ex Preston).
Preston wrote “[a]n extraordinary shell”; the singular implies that he had only one specimen at hand, the specimen thus is the holotype. A label states “Purchased from / Preston many years ago / by Mayor Connolly with / others / A. M. N. H. viii p. 509”. The current systematic position follows
Megaspiridae, Thaumastus insolitus (Preston, 1909).
Bulimus integer
Porphyrobaphe integer;
Thaumastus (Thaumastus) integer;
“Quito, Ecuador”.
“Quito”. M.C. label style IV.
“Long. 82, diam. 39 mill.”. Figured specimen H 81.5, D 42.0, W 7.4.
NHMUK 1975244, lectotype; 1975245, one paralectotype (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based; he did, however, recognize a variety β for which he gave “Long. 65, diam. 31 mill.” as measurements. This was likely a shell from his own collection, as the paralectotype in the Cuming collection has a shell height of 71.6 mm. The protoconch of these type specimens is sculptured with axial wrinkles, becoming coarse granules on the lower part of the protoconch. The generic classification of
Megaspiridae, Thaumastus integer (Pfeiffer, 1855).
Bulimus iris
Porphyrobaphe iris;
“Le Ceja, Rio Negro Novae Granadae (Bland)”.
“La Ceja. Rio Negro / New Grenada”, taxon label in Pfeiffer’s handwriting. M.C. label style I.
“Long. 64, diam. 32 mill.”. Figured specimen H 72.6, D 41.1, W 5.8.
NHMUK 20100506, lectotype (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based, but described the species from “Mus. Cuming”. Although the specimen is larger than the measurements given by Pfeiffer, there is little doubt it is from the original series as the label confirms the original locality; it also bears the text “please to name this / ‘Blandi’ after the collector”. The specimen is now designated lectotype (design. n.) to fixate this taxon, which needs further study to clarify its status. The mentioning in
Orthalicidae, Porphyrobaphe (Porphyrobaphe) iris (Pfeiffer, 1853).
Bulimus irroratus
Porphyrobaphe irrorata;
“Brazil? New Granada?”.
“Quito Ecuador”, taxon label in Pfeiffer’s handwriting. M.C. label style III.
Not given. Figured specimen H 77.0, D 44.0, W 6+.
NHMUK 1975248, three syntypes, A.L. Gubba leg. (Cuming coll.).
Reeve did not state on how many specimens his description was based, but said it was the “Mus. Cuming (...) thanks to the liberality of A.L. Gubba, Esq., of [Le] Havre”. In Reeve (1850) the locality was mentioned as “—?”. The top of the specimen figured is damaged. The current systematic position is according to
Orthalicidae, Porphyrobaphe irrorata (Reeve, 1849).
Bulimus jeffreysi
Drymaeus obliquus (Reeve);
Leiostracus obliquus;
“Brasilia”.
“Brazils”, taxon label in Pfeiffer’s handwriting. M.C. label style I.
“Long. 19, diam. 11 mill.”. Figured specimen H 20.4, D 10.9, W 6+.
NHMUK 20110083, three syntypes (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based; he described the taxon “Ex Coll. Cl. Gruner” (
Simpulopsidae, Leiostracus obliquus (Reeve, 1849).
Bulimus kelletti
Orthalicus kellettii;
Sultana (Metorthalicus) kellettii;
“Ecuador?”.
“?Ecuador”. M.C. label style III, V.
Not given. Figured specimen H 61.2, D 33.2, W 5.7.
NHMUK 1975241, lectotype (Cuming coll.).
The material is accompanied by a label “the type specimen”. However, Reeve did not state on how many specimens his description was based, but only mentioned “this new and very beautiful species”. The specimen found should thus be regarded as lectotype, contradicting the statement by
Orthalicidae, Sultana (Metorthalicus) kelletti (Reeve, 1850).
Bulimus loxostomus
Strophocheilus loxostomus;
Thaumastus (Thaumastus) loxostomus;
“in Andibus Novae Granadae”.
“Andes N. Granada”. M.C. label style III.
“Long. 71, diam. 34 mill.”. Figured specimen H 71.3, D 37.3, W 5.8.
NHMUK 1975125, one syntype (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based, but said the material was in “Mus. Cuming”. The protoconch is sculptured with spaced, indistinct wrinkles, becoming closer towards the transition to the teleoconch. The lip is white, which is quite unusual for Thaumastuss.str. Further research should thus shed more light on the systematic position of this taxon, which is here figured for the first time.
Megaspiridae, Thaumastus loxostomus (Pfeiffer, 1853).
Achatina magnifica
Liguus (Hemibulimus) magnificus;
“Quito, Ecuador; in woods (De Lattre)”.
“Quito, Ecuador”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.
“Long. 47, diam. 21 mill.”. Figured specimen H 46.6, D 23.0, W 5.5.
NHMUK 20100508, two syntypes (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based, but described this taxon from Cuming’s collection. Reeve’s figure (
Orthalicidae, Clathrorthalicus magnificus (Pfeiffer, 1848) (comb. n.).
Bulimus magnificus
Strophocheilus magnificus;
Thaumastus (Thaumastus) magnificus;
Thaumastus magnificus;
“Pérou”.
“Le Perou (Brésil)”; see remarks.
“Près de 3 pouces de longueur”; see remarks. Figured specimen H 78.0, D 36.0, W 6.9.
NHMUK 1907.11.22.24, lectotype (da Costa coll., ex Grateloup).
Grateloup did not state on how many specimens his description was based; in
Megaspiridae, Thaumastus magnificus (Grateloup, 1839).
Bulimus marmatensis
Bulimulus marmatensis;
[Colombia] “Marmato, New Granada”.
“Marmata / New Grenada”, taxon label in Pfeiffer’s handwriting. M.C. label style III.
“Long. 17, diam. 10 mill.”. Figured specimen H 15.0, D 11.0, W 5.0.
NHMUK 1975330, three syntypes (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based; he, however, described this taxon from the collection of Cuming. Three specimens were found, two damaged adults and one juvenile. The protoconch is sculptured with axial wrinkles and spiral lines; this taxon—classified by
Simpulopsidae, Simpulopsis (Eudioptus) citrinovitrea (Moricand, 1836).
Orthalicus mars
“republica Aequatoris (Mr. Fraser)”.
“Republic of Ecuador”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.
“Long. 77, diam. 35 mill.”. Figured specimen H 76.6, D 38.4, W 6+.
NHMUK 20100504, three syntypes (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based, but described “from the collection of H. Cuming”. Although on the label has been written in a later hand “none quite like fig.”, the type status is not disputed as the shell height matches the original data. The top of the largest specimen, herein figured, is damaged. Also the top of one of the other specimens is damaged. The protoconch of the third, undamaged, specimen is smooth. This taxon is tentatively classified with Orthalicus Beck, 1837; however, further anatomical and molecular research should provide evidence to assess if this classification is correct or needs to be adjusted.
Orthalicidae, Orthalicus mars Pfeiffer, 1861.
Bulimus meobambensis
Orthalicus meobambensis;
“Meobamba, Eastern Peru (Mr. Yates)”.
“Meobamba / East Peru”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.
“Long. 88, diam. 46 mill.”. Figured specimen H 84.9, D 52.8, W 6.4.
NHMUK 20100505, two syntypes (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based. There is no doubt, however, about the type status of the specimens found as he described this taxon from the Cuming collection and the taxon label is—although in pencil—in his handwriting. This is the first time this material is figured;
Orthalicidae, Sultana (S.) meobambensis (Pfeiffer, 1855).
Simpulopsis miersi
[Brazil] “Espirito Santo Brasiliae (Miers)”.
“Espirits Santo / F. Miers Esq.”, taxon label in Pfeiffer’s handwriting. M.C. label style III.
“Diam. maj. 24, alt. 17 1/2 mill.”. Figured specimen H 20.6, D 20.9, W 4.5.
NHMUK 1975489, lectotype; 1975490 one paralectotype (Cuming coll., ex Miers).
Pfeiffer did not state on how many specimens his description was based, but refers to “Mus. Cuming”, and Miers as source. The lectotype is slightly damaged at the body of the last whorl and the lip. The references of
Simpulopsidae, Simpulopsis (Simpulopsis) miersi Pfeiffer, 1857.
Achatina murrea
“—?”.
No locality label, taxon label in Pfeiffer’s handwriting. M.C. label style IV, V.
Not given. Figured specimen H 38.6, D 19.5, W 7.1.
NHMUK 1975482, 20230332, three + three syntypes (Cuming coll.).
Reeve did not state on how many specimens his description was based, but figured two different specimens from “Mus. Cuming”. Three specimens were found in lot NHMUK 1975482, one of which corresponds to Reeve's figure 22a; Pfeiffer has identified this lot as “A. fasciata / Müller juv.”. Lot 20120332 also contains three specimens, one of which was figured as fig. 22b. The current systematic position follows
Orthalicidae, Liguus murreus (Reeve, 1849).
Orthalicus powissianus var. niveus
“Jimenez, Rio Dagua, West Colombia”.
“Jimenez Rio Dagua / 1600 ft. Colombia”, in Preston’s handwriting.
Not given. Figured specimen H 65.2, D 31.3, W 7.2.
NHMUK 1909.8.18.85, holotype, M.G. Palmer leg., ex Preston.
Preston mentioned “taken with the animal alive”, from which may be inferred that he had only one specimen at hand. The specimen located is thus the holotype; the top is slightly damaged. The taxon is here figured for the first time. The current systematic position at species level follows
Orthalicidae, Sultana (Methorthalicus) powisiana (Petit de la Saussaye, 1843).
Bulimus obliquus
Drymaeus obliquus;
Leiostracus (Leiostracus) obliquus;
Leiostracus obliquus;
[Brazil] “Bahia”.
“Brazil”. M.C. label style I, V.
Not given. Figured specimen H 22.7, D 12.05, W 6+.
NHMUK 1975493, lectotype (Cuming coll.).
Reeve did not state on how many specimens his description was based, but mentioned “[a] pink shell”; this is herein not considered as sufficient evidence that he had only one shell for his description. The material was in “Mus. Cuming”. The top and the apertural lip of the specimen found are damaged. The current systematic position follows
Simpulopsidae, Leiostracus obliquus (Reeve, 1849).
Bulinus opalinus Sowerby I 1833 in
Bulimus opalinus;
Leiostracus perlucidus;
“Brazil”.
“Brazil”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.
Not given. Figured specimen H 27.8, D 14.3, W 7.4.
NHMUK 1975442, three probable syntypes (Cuming coll.).
Sowerby did not state on how many specimens his description was based; he wrote “Nob.”, thus “ours”, meaning the author claimed his right as describer of the new taxon, not necessarily proof of presence in his own collection. As
Simpulopsidae, Leiostracus perlucidus (Spix, 1827).
Bulimus ovulum
Rhinus ovulum;
“Philippine Islands; Cuming”.
“Pernambuco, Brazil”, taxon label in Pfeiffer’s handwriting. M.C. label style I, V.
Not given. Figured specimen H 20.1, D 12.3, W 6.4.
NHMUK 1975416, lectotype; 1975417, two paralectotypes (Cuming coll.).
Reeve did not state on how many specimens his description was based, but mentioned “A shell of rather solid growth...”; this is herein not considered as sufficient evidence that he had only one shell for his description. The material was in “Mus. Cuming”.
Simpulopsidae, Rhinus ovulum (Reeve, 1849).
Helix phlogera
Bulimus phlogerus;
“provincia Chiquitensi (republica Boliviana)”.
“Sn Xavier, Chquitos (Bolivia)”, in d’Orbigny’s handwriting.
“Longit. 55 millim.; latit. 24 millim.”. Figured specimen H 59.8, D 26.8, W 6+.
NHMUK 1854.12.4.86, six syntypes (d’Orbigny coll.).
Orthalicidae, Orthalicus phlogerus (d’Orbigny, 1835).
Bulimus phoebus
Plekocheilus phoebus;
Plekocheilus (Eurytus) phoebus;
“Ecuador”.
“Ecuador”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.
“Long. 31, diam. 15 mill.”. Figured specimen H 30.5, D 17.5, W 5.5.
NHMUK 1975143, lectotype (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based, but described his material “from the collection of H. Cuming”. This taxon has long been associated with Plekocheilus (Eurytus) Albers, 1850, but re-examination of the type—of which the protoconch proves to be smooth—plus recent collections in north-western Ecuador (Breure unpublished data) reveal that this taxon belongs to Clathrorthalicus Strebel, 1909. It may be closely allied to Bulimus corydon Crosse, 1869, B. magnificus Pfeiffer, 1848 and B. victor Pfeiffer, 1854; however, further anatomical and molecular studies should clarify the current systematic position.
Orthalicidae, Clathrorthalicus phoebus (Pfeiffer, 1863) (comb. n.).
Bulimus plumbeus
Strophocheilus plumbeus;
Thaumastus (Thaumastus) plumbeus;
“Venezuela”.
“Venezuela”. M.C. label style III.
“Long. 93, diam. 36 mill.”. Figured specimen H 93.0, D 40.5, W 5.9.
NHMUK 1975130, lectotype; 1975131, one paralectotype (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based, but described his material “from the collection of H. Cuming”. Although there is no label in Pfeiffer’s handwriting, the type status of these specimens is not disputed as the shell height matches the original data. This taxon has been considered a junior subjective synonym of Helix (Cochlogena) pardalis Férussac, 1821 (
Megaspiridae, Thaumastus plumbeus (Pfeiffer, 1855).
Bulimus porphyrius
Thaumastus (Scholvienia) porphyrius;
“Bolivia (T. Bridges)”.
“Bolivia”, “andes of Caxamarca / Peru”, taxon label in Pfeiffer’s handwriting. M.C. label style IV, V.
“Long. 51, diam. 20 mill.”. Figured specimen H 51.5, D 22.0, W 6.6.
NHMUK 1975277, lectotype; 1975278, two paralectotypes (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based; it was, however, one of the taxa from the Cuming collection.
?Orthalicidae, Scholvienia porphyria (Pfeiffer, 1847) (comb. n.).
Bulimus requieni
Strophocheilus requieni;
Thaumastus (Thaumastus) requieni;
Thaumastus requieni;
“Brasilia”.
“Brazils”, taxon label in Pfeiffer’s handwriting. M.C. label style IV.
“Long. 62, diam. 26 mill.”. Figured specimen H 62.0, D 29.0, W 5.3+.
NHMUK 1975301, lectotype; 1975302, one paralectotype (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based, but described his material from “Mus. Cuming”. The top of the lectotype is slightly damaged. The protoconch is sculptured with slightly waving axial riblets. Both specimens appear to be subadult; further studies are needed to ascertain the taxonomic position of this taxon. The current systematic position follows
Megaspiridae, Thaumastus requieni (Pfeiffer, 1853).
Vitrina salomonia
Simpulopsis salomonia; Pfeiffer 1854 in
Simpulopsis (?) salomonia;
Simpulopsis (S.) rufovirens (Moricand);
“in insulis Salomonis”.
“Solomons Isl”, taxon label in Pfeiffer’s handwriting. M.C. label style III.
“Long. 11, diam. 9 mill.”. Figured specimen H 11.1, D 10.7, W 4.5.
NHMUK 1975485, lectotype (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based, but noted “Mus. Cuming”. He also remarked “Diese Art ist mit der brasilianischen Gruppe Simpulopsis Beck sehr nahe verwandt”, which might have led
Simpulopsidae, Simpulopsis (S.) rufovirens (Moricand, 1846).
Bulimus salteri
“Catamarca, Andes Peruviae”.
“Andes of Peru”; printed label.
“Long. 70, maj. diam. 35 mill.”. Figured specimen H 69.9, D 35.2, W 6.0.
NHMUK 1907.11.21.118, lectotype (da Costa coll.).
Sowerby also described a (larger) “var. γ”, and remarked “[t]he two shells form part of the collection of Mr. S.J. Da Costa, and there is a specimen of each variety [typical one and var. γ] in the National Collection at South Kensington [= NHMUK]”. The original series thus seems to have comprised two specimens, and the reference in
?Orthalicidae, Quechua salteri (Sowerby III, 1890) (comb. n.).
Bulimus sarcochilus
Bulimulus sarcochilus;
Leiostracus (L.) sarcochilus;
Leiostracus sarcochilus;
“in Brasilia septentrionali (Miers)”.
“North of Brazils / F. Miers Esq”, taxon label in Pfeiffer’s handwriting. M.C. label style I.
“Long. 21–25, diam. 10–11 1/2 mill.”. Figured specimen H 24.7, D 13.1, W 7.7.
NHMUK 1975398, lectotype; 1975399, one paralectotype (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based, but from his dimensions it is clear that he had more than one specimen at hand. From
Simpulopsidae, Leiostracus sarcochilus (Pfeiffer, 1857).
Bulimus saturanus
Bulimus satuanus
Bulimus saturnus;
Porphyrobaphe saturnus;
“Pallatanga, Republic of Ecuador (Mr. Fraser)”.
“Pallatango Republic of Ecuador Mr Fraser”, taxon label in Pfeiffer’s handwriting. M.C. label style I.
“Long. 76, diam. 33 mill.”. Figured specimen H 75.8, D 38.4, W 6.7.
NHMUK 20140080, three syntypes, Fraser leg. (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based. In the original paper he made twice an error in the name, which was corrected in
Orthalicidae, Porphyrobaphe (Porphyrobaphe) saturnus (Pfeiffer, 1860).
Bulimus simulus
[Guatemala] “sylvas Petenensis”.
“forêt de Dolores”, taxon label in Morelet’s handwriting.
“Longit. 11 – Diam. 9 [mm]”. Figured specimen H 8.26, D 6.79, W 4.3.
NHMUK 1893.2.4.1128–1129, two syntypes (Morelet coll.).
Morelet did not state on how many specimens his description was based. On the board on which the labels are glued has been written in a later hand “Type largest / Test. Noviss. No. 101”. The locality on the label probably refers to the village of Dolores, Petén, Guatemala, which is thus the exact type locality. The current systematic position follows
Simpulopsidae, Simpulopsis (Simpulopsis) simula (Morelet, 1851).
Strophocheilus (Eurytus) subirroratus
Porphyrobaphe subirroratus;
Porphyrobaphe (Oxyorthalicus) subirroratus;
“Paramba, Ecuador”.
“Paramba, Ecuador”, in da Costa’s handwriting.
“Long. 63, diam. 33 mm.”. Figured specimen H 62.6, D 36.6, W 5.9.
NHMUK 1907.11.21.114, lectotype (da Costa coll.).
da Costa did not state on how many specimens his description was based; the reference in
Orthalicidae, Porphyrobaphe (Oxyorthalicus) subirroratus (da Costa, 1898).
Bulimus onager
Drymaeus (Leiostracus) onager var. subtuszonata
Leiostracus subtuzonatus [sic];
Not given.
Not given [“Brazil” added in a later hand]. M.C. label style I, V.
“[L]ength of 28 mm”. Figured specimen H 29.0, D 14.8, W 7.9.
NHMUK 20130094, three probable paralectotypes (Cuming coll.).
Pilsbry did not state on how many specimens his description was based, and gave no type locality as he described what he regarded as a colour variation only. His figure was a black and white copy of Reeve’s figure.
Simpulopsidae, Leiostracus subtuszonatus (Pilsbry, 1899).
Bulimus thompsonii
Thaumastus (Kara) thompsoni [sic];
[Ecuador] “Quito. (Coll. Cuming)”.
“Quito”, taxon label in Pfeiffer’s handwriting. M.C. label style IV, V.
“Long. 70, diam. 31 mill.”. Figured specimen H 71.0, D 32.0, W 6.2.
NHMUK 1975464, lectotype; 1975465, two paralectotypes (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based. The specimen figured by Reeve has been selected lectotype by
Orthalicidae, Kara thompsonii (Pfeiffer, 1845).
Bulimus labeo
Porphyrobaphe vicaria
“Leimabamba, Peru, 8000 feet (O.T. Baron)”.
“Limabambo Peru”. M.C. label style IV.
Not given. Figured specimen H 82.2, D 46.7, W 6.3+.
NHMUK 20100507, holotype, Lobb leg. (Cuming coll.).
Fulton mentioned “[t]ype in British Museum (Cuming Collection)”, and said his taxon had been figured by Reeve. Reeve wrote: “It is with much gratification that I am enabled to give an original figure of the Bulimus labeo, figured at. Pl. XXXV, from a figure in the Zoological Journal. This shell, from the Cumingian collection, which I take to be identical with the lost specimen [see
Orthalicidae, Sultana (Metorthalicus) yatesi (Pfeiffer, 1855).
Bulimus victor
Plekocheilus victor;
“in provincia Antioquia, Columbiae (Schlim)”.
“Province of Antioquia / [...] Schlim [...]”, taxon label in Pfeiffer’s handwriting. M.C. label style III.
“Long. 65, diam. 29 mill.”. Figured specimen H 64.0, D 36.7, W 5+.
NHMUK 1975242, lectotype; 20100567, one paralectotype (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based; besides the specimen corresponding to Pfeiffer’s dimensions, and selected lectotype by
Orthalicidae, Clathrorthalicus victor (Pfeiffer, 1854) (comb. n.).
Simpulosis vincentina
[West Indies, St. Vincent] “Damp forest, Upper Richmond valley, 2000 ft, on leaves of Artanthe (Piperacea) (H.H. Smith)”.
“Damp forest, Upper Richmond valley, 2000 ft, on leaves of Artanthe (Piperacea), St. Vincent, B.W.I.”.
“Longit. 13, diam. maj. 10 mm”. Figured specimen H 11.4, D 10.2, W 2.8.
NHMUK 1895.6.17.458, holotype, H.H. Smith leg.
E.A. Smith wrote “[o]nly a single specimen was collected.” Both body of the last whorl and the lip are partly broken in the holotype. The current systematic position follows
Simpulopsidae, Simpulosis (S.) vincentina E.A. Smith, 1895.
Bulimus yanamensis
Strophocheilus yanamensis;
Thaumastus (Kara) yanamensis;
[Peru] “Yanama”.
“Yanama. Pérou”, taxon label in Morelet’s handwriting.
“Longit. 58; diam. 25 (...) mill.”. Figured specimen H 48.6, D 26.9, W 5.4.
NHMUK 1893.2.4.167–168, [two paralectotypes] (Morelet coll.).
Morelet did not state on how many specimens his description was based; the two specimens mentioned by
Orthalicidae, Kara yanamensis (Morelet, 1863).
Bulimus yatesi
Orthalicus yatesi;
Sultana (Metorthalicus) yatesi;
“Meobamba, Eastern Peru (Mr. Yates)”.
“Meobamba, East Peru”. M.C. label style III.
“Long. 82, diam. 32 mill.”. Figured specimen H 84.3, D 39.7, W 7.2.
NHMUK 1975239/1, lectotype; 1975239/2–3, two paralectotypes (Cuming coll.).
Pfeiffer did not state on how many specimens his description was based;
Orthalicidae, Sultana (Metorthalicus) yatesi (Pfeiffer, 1855).
The systematic position of the following taxon cannot be ascertained at present, and it is herein considered a nomen inquirendum.
Bulimulus (Protoglyptus) dejectus Fulton 1907: 153, pl. 10 fig. 1;
Protoglyptus dejectus;
“Santa Catarina (fide Linnaea Institute label)”.
“St. Catharina”.
“Maj. diam. 10, alt. 29 mm”. Figured specimen H 29.2, D 10.0, W 7.8.
NHMUK 1907.5.3.163, lectotype (ex Sowerby and Fulton).
Fulton did not state on how many specimens his description was based. The specimen found agrees with Fulton’s measurements and is now designated lectotype (design. n.). The sculpture of the protoconch is not with axial wrinkles as usual in Protoglyptus Pilsbry, 1897, but with axial wrinkles, partly broken into granules. It may be noted that all species currently classified with this genus occur in the West Indies (
?Bulimulidae, ?Naesiotus dejectus (Fulton, 1907). Nomen inquirendum.
Type material of the following taxa, previously known to be extant in the NHMUK, has not been found during our study.
Bulimus dennisoni
Hemibulimus (Myiorthalicus) dennisoni;
“—?”.
Not given.
The two syntypes mentioned by
Helix miliola
Pupa miliola
“imperio Brasiliano”.
“Latit. 2 millim., longit. 1 millim.”.
NHMUK 1854.12.4.239, [seven syntypes] (d’Orbigny coll.).
Seven specimens were known to be present (cf. the registration book, which has an undated note in pencil “6 missing”), but none could not be found. This taxon has been mentioned under two different species by
Helix progastor
Simpulopsis progastor;
Eudioptus progastor;
“Brasilianis oris”.
“Longit. 7 millim.”.
NHMUK 1854.12.4.72, [one syntype] (d’Orbigny coll.).
Bulimus vitrinoides
“—?” “Mus. Cuming”.
Not given.
The syntype material mentioned by
The following taxa were expected to be represented with type material; however, no material could be found matching the data in the original publication.
Bulinus adamsonii
Bulini guadaloupensis alba Sowerby I in
Achatina atramentaria
Bulimus aulacostylus
Bulinus bilabiatus
Bulinus bilineatus
Bulinus bivittatus
Bulimus blainvilleanus
Bulimus bolivianus
Bulinus cactivorus Broderip in
Bulimus cantatus
Bulimus cardinalis
Bulimus castelnaui
Bulimus castrensis
Bulimulus (Drymaeus) chacoensis
Bulimus coerulescens
Bulimus columellaris
Bulimus confinus
Bulimus coniformis
Bulimus constrictus
Bulimus contortuplicatus
Bulinus coquimbensis Broderip in
Bulinus corneus
Bulinus corrugatus King in
Bulimus curianianus
Bulinus decoloratus
Bulimus draparnaudi
Bulimus droueti
Bulimus eganus Pfeiffer 1853 in
Bulinus erythrostoma
Partula flavescens King in
Bulinus granulosus Broderip in
Bulinus gravesii King in
Plekocheilus glaber grenadensis
Bulimus guentheri
Bulimus guttula
Bulimus hegewischi
Bulimus hennahi
Bulimus hyematus
Otostomus (Drymaeus) lilacinus ictericus
Bulimus inaequalis
Bulinus inflatus Broderip, 1836: 45.—Described from a shell “brought home by Mr. Cuming”, but not encountered in the collection.
Bulimus iostoma
Bulinus janeirensis Sowerby I 1838 [
Bulimus jucundus
Bulinus labeo
Bulinus laurentii
Bulimus lindeni
Helix listeri
Orthalicus macandrewi
Bulinus mutabilis Broderip in
Bulimus navarrensis
Otostomus chiapensis nebulosus
Bulimulus (Drymaeus) nigroumbilicatus
Helix orobaena
Bulinus pallidior
Bulimus pardalis
Bulimus peelii
Bulimus pentlandi
Otostomus attenuatus pittieri
Bulimus primularis
Bulinus princeps Broderip in
Tomigerus principalis
Bulinus pulchellus Broderip in
Bulinus pulchellus Sowerby I 1838 [
Bulimus rhodacme
Bulinus rubellus
Bulimus rubescens
Bulimus rupicolus
Bulimus sarcodes
Bulimus sayi
Bulimus scytodes
Bulimus sporadicus
Bulinus striatulus
Bulinus striatus ‘King’
Clausilia? (Balea?) taylori
Bulinus tigris Broderip in
Otostomus (Drymaeus) lilacinus undulosus
Bulinus varians Broderip in
Tomigerus venezuelensis
Bulimus venosus
Bulimulus (Drymaeus) ventricosus
Bulinus vexillum Broderip in
Helix vexillum
Bulimulus (Drymaeus) vicinus
Bilinus vittatus Broderip in
Bulimus ziegleri
Bulimus senilis
Placostylus senilis;
[New Caledonia] “Baie du Sud (Nov. Cal.)” (see remarks).
“New Caledonia”.
“Long. 129 mill., diam. maj. 65”. Figured specimen H 126.1, D 56.3, W 7.0.
NHMUK 1883.11.10.1179, one paralectotype ex Sowerby ex Gassies.
Bothriembryontidae, Placostylus senilis (Gassies, 1869).
p. 3: salomonis (Pfeiffer, 1853) shoud be removed under Placostylus Beck, 1837, and inserted under Santacharis Iredale, 1927.
p. 25: Pupa spixii major d’Orbigny, 1837 under Type material: the lectotype has registration number 1854.12.4.230 instead of 1885.12.4.232. The latter lot is from “Corrientes, Argentina”, while lot 1854.12.4.230 is from “Guarayos, Bolivia”. The specimen figured in Figs
p. 30: Bulimus ouensis Gassies, 1870 under Type material: the holotype has registration number 1883.11.10.1176 where it should read 1883.11.10.1167.
Bulimus diaphanus
Bulimulus diaphanus;
Bulimulus (Bulimulus) diaphanus diaphanus; Breure 1974: 30, pl. III fis 11–12.
“S. Thomas, West Indies (Bland)”.
“Long. 15, diam. 7 mill.”.
The two specimens mentioned and figured by Breure (1974) have to be considered as lost, as—despite repetitive searches—they could not be re-found during our research.
Bulimulus (Drymaeus) interruptus var. pallidus
“Merida, Venezuela”.
“Merida, Venezuela”.
Not given. Figured specimen H 23.5, D 11.9, W 4.3+.
NHMUK 1914.4.3.41, lectotype (ex Preston).
This varietal name has been treated as unavailable under Art. 45.6 ICZN by
Our opinion is as follows:
Preston undoubtedly had a series of specimens at hand when describing Bulimulus (Drymaeus) interruptus; both the wording “to be greatly variable” and “its principle forms” are indicative of this.
Köhler (1997) also concluded that this taxon was described from several specimens but said “A holotype has not been designated. Therefore, the present specimen [ZMB 59597] is a syntype. Consequently, the specimen in the BMNH referred to as holotype by
Reconsidering the (un)availability under Art. 45.6, it is important to note that var. pallidus was proposed before 1961 and has to be treated as subspecific (see the contributions of Steve Lingafelter and Doug Yanega on the Taxacom listing, http://to.ly/zFZO). It may be noted that the only reference to this taxon after Preston’s publication is in
While Bulimulus (Drymaeus) interruptus var. pallidus is an available name, we concur with
The specimen of var. pallidus in the NHMUK is now designated lectotype (design. n.) to fixate this synonymisation. The text in
Bulimulidae, Drymaeus (Mesembrinus) interruptus (Preston, 1909).
The ’Systematic list of taxa arranged in generic order’ on page 3–7 should be replaced by the following text:
swainsoni Pfeiffer, 1845.
acalles Pfeiffer, 1853; affinis Broderip in
barbadensis Pfeiffer, 1853; cacticolus Reeve, 1849; dysoni Pfeiffer, 1846; effeminatus Reeve, 1848; erectus Reeve 1849; haplochrous Pfeiffer, 1855; heloica d’Orbigny, 1835; ignavus Reeve, 1849; inutilis Reeve, 1850; istapensis Crosse and Fischer, 1873; juvenilis Pfeiffer, 1855; marcidus Pfeiffer, 1853; mollicellus Reeve, 1849; monachus Pfeiffer, 1857; montevidensis Pfeiffer, 1846; nubeculatus Pfeiffer, 1853; pervius Pfeiffer, 1853; pessulatus Reeve, 1848; petenensis Morelet, 1851; pliculatus Pfeiffer, 1857; rubrifasciatus Reeve, 1848; sporadica d’Orbigny, 1835; transparens Reeve, 1849; turritella d’Orbigny, 1835; vesicalis Pfeiffer, 1853.
abruptus Rolle, 1904; abscissus Pfeiffer, 1855; abyssorum d’Orbigny, 1835; aequatorianus E.A. Smith, 1877; acervatus Pfeiffer, 1857; acuminatus da Costa, 1906; alabastrinus da Costa, 1906; albolabiatus E.A. Smith, 1877; ambustus Reeve, 1849; angustus da Costa, 1906; antioquiensis Pfeiffer, 1855; arcuatostriatus Pfeiffer, 1855; auris Pfeiffer, 1866; baranguillanus Pfeiffer, 1853; bartletti H. Adams, 1867; bellus da Costa, 1906; bogotensis Pfeiffer, 1855; bolivarii d’Orbigny, 1835; bolivianus Pfeiffer, 1846; boucardi da Costa, 1907; bourcieri Pfeiffer, 1853; brachysoma d’Orbigny, 1835; buckleyi Sowerby III, 1895; canaliculatus Pfeiffer, 1845; castaneostrigatus da Costa, 1906; caucaensis da Costa, 1898; chamaeleon Pfeiffer, 1855; chimborasensis Reeve, 1848; chiriquensis da Costa, 1901; clathratus Pfeiffer, 1858; coarctatus Pfeiffer, 1845; confluens Pfeiffer, 1855; convexus Pfeiffer, 1855; cuticula Pfeiffer, 1855; cuzcoensis Reeve, 1849; dacostae Sowerby III, 1892; dombeyanus Pfeiffer, 1846; dunkeri Pfeiffer in Philippi, 1846; elsteri da Costa, 1901; exoticus da Costa, 1901; expatriatus Preston, 1909; fabrefactus Reeve, 1848; fallax Pfeiffer, 1853; farrisi Pfeiffer, 1858; felix Pfeiffer, 1862; fenestratus Pfeiffer, 1846; flexilabris Pfeiffer, 1853; flexuosus Pfeiffer, 1853; fucatus Reeve, 1849; fusoides d’Orbigny, 1835; gealei H. Adams, 1867; geometricus Pfeiffer, 1846; gueinzii Pfeiffer, 1857; hidalgoi da Costa, 1898; humboldtii Reeve, 1849; hygrohylea d’Orbigny, 1835; inclinatus Pfeiffer, 1862; incognitus da Costa, 1907; jansoni Martens, 1893; josephus Angas, 1878; knorri Pfeiffer in Philippi, 1846; lamas Higgins, 1868; lattrei Pfeiffer in Philippi, 1846; laxostylus Rolle, 1904; lilacinus Reeve, 1849; linostoma d’Orbigny, 1835; lophoica d’Orbigny, 1835; lucidus da Costa, 1898; malleatus da Costa, 1898; marmarina d’Orbigny, 1835; murrinus Reeve, 1848; musivus Pfeiffer, 1855; napo Angas, 1878; notabilis da Costa, 1906; notatus da Costa, 1906; nystianus Pfeiffer, 1853; ochrocheilus E.A. Smith, 1877; orthostoma E.A. Smith, 1877; patricius Reeve, 1849; perenensis da Costa, 1901; pergracilis Rolle, 1904; phryne Pfeiffer, 1863; plicatoliratus da Costa, 1898; poecila d’Orbigny, 1835; ponsonbyi da Costa, 1907; praetextus Reeve, 1849; protractus Pfeiffer, 1855; pseudofusoides da Costa, 1906; pulcherrimus H. Adams, 1867; punctatus da Costa, 1907; quadrifasciatus Angas, 1878; recedens Pfeiffer, 1864; regularis Fulton, 1905; rosenbergi da Costa, 1900; rubrovariegatus Higgins, 1868; saccatus Pfeiffer, 1855; schmidti Pfeiffer, 1854; scitulus Reeve, 1849; scitus H. Adams, 1867; selli Preston, 1909; serratus Pfeiffer, 1855; smithii da Costa, 1898; solidus Preston, 1907; spadiceus da Costa, 1906; spectatus Reeve, 1849; strigatus Sowerby I, 1833; subhybridus da Costa, 1906; subinterruptus Pfeiffer, 1853; subventricosus da Costa, 1901; sykesi da Costa, 1906; tigrinus da Costa, 1898; vespertinus Pfeiffer, 1858; volsus Fulton, 1907; xanthostoma d’Orbigny, 1835; yungasensis d’Orbigny, 1837; zhorquinensis Angas, 1879; ziczac da Costa, 1898; zoographica d’Orbigny, 1835.
aestivus Pfeiffer, 1857; amandus Pfeiffer, 1855; andicola Pfeiffer, 1847; apicepunctata Preston, 1914; apiculata J.E. Gray, 1834; attenuatus Pfeiffer, 1853; aureolus Guppy, 1866; aurifluus Pfeiffer, 1857; broadwayi E.A. Smith, 1896; bugabensis Martens, 1893; californicus Reeve, 1848; cancellata da Costa, 1906; castus Pfeiffer, 1847; championi Martens, 1893; citronellus Angas, 1879; columbiensis Pfeiffer, 1856; conicus da Costa, 1907; demotus Reeve, 1850; depictus Reeve, 1849; deshayesi Pfeiffer, 1845; discrepans Sowerby I, 1833; dubius Pfeiffer, 1853; dutaillyi Pfeiffer, 1857; electrum Reeve, 1848; erubescens Pfeiffer, 1847; feriatus Reeve, 1850; fidustus Reeve, 1849; flavidulus E.A. Smith, 1877; floridanus Pfeiffer, 1857; fuscobasis E.A. Smith, 1877; gabbi Angas, 1879; gruneri Pfeiffer, 1846; hachensis Reeve, 1850; hepatostomus Pfeiffer, 1861; hoffmanni Martens, 1893; hondurasanus Pfeiffer, 1846; hypozonus Martens, 1893; immaculatus C.B. Adams in Reeve, 1850; incarnatus Pfeiffer, 1855; inglorius Reeve, 1848; interruptus Preston, 1909; inusitatus Fulton, 1900; iodostylus Pfeiffer, 1861; jonasi Pferiffer in Philippi, 1846; keppelli Pfeiffer, 1853; koppeli Sowerby III, 1892; laetus Reeve, 1849; lascellianus E.A. Smith, 1895; lirinus Morelet, 1851; lividus Reeve, 1850; loxanus Higgins, 1872; loxensis Pfeiffer, 1846; lucidus Reeve, 1848; lusorius Pfeiffer, 1855; manupictus Reeve, 1848; meridanus Pfeiffer, 1846; monilifer Reeve, 1848; moricandi Pfeiffer, 1847; moritinctus Martens, 1893; mossi E.A. Smith, 1896; moussoni Pfeiffer, 1853; muliebris Reeve, 1849; nigrofasciatus Pfeiffer in Philippi, 1846; nitelinus Reeve, 1849; nitidus Broderip in
confusus Reeve, 1848; culminea d’Orbigny, 1835; edwardsi Morelt, 1863; gayi Pfeiffer, 1857; jussieui Pfeiffer, 1846; lithoica d’Orbigny, 1835; thamnoica d’Orbigny, 1835; tupacii d’Orbigny, 1835.
David Campbell (pers. commun.) kindly made us aware that the name Vermiculatus Breure, 1978 is preceded by Bocourtia Rochebrune, 1882.
aequatorius Pfeiffer, 1853; anthisanensis Pfeiffer, 1853; aquilus Reeve, 1848; badius Sowerby I, 1835; bicolor Sowerby I, 1835; caliginosus Reeve, 1849; coagulatus Reeve, 1849; cotopaxiensis Pfeiffer, 1853; filaris Pfeiffer, 1853; nucinus Reeve, 1850; ochraceus Morelet, 1863; peaki Breure, 1978; petiti Pfeiffer, 1846; polymorpha d’Orbigny, 1835; purpuratus Reeve, 1849; quechuarum Crawford, 1939; subfasciatus Pfeiffer, 1853.
achatellinus Forbes, 1850; albemarlensis Dall, 1917; apertus Pfeiffer, 1855; catlowiae Pfeiffer, 1853; chamayensis Weyrauch, 1967; chemnitzioides Forbes, 1850; cinereus Reeve, 1849; crepundia d’Orbigny, 1835; curtus Reibisch, 1892; darwini Pfeiffer, 1846; dentritis Morelet, 1863; durangoanus Martens 1893; eschariferus Sowerby I, 1838; exornatus Reeve, 1849; fernandezae Weyrauch, 1958; fontainii d’Orbigny, 1838; fourmiersi d’Orbigny, 1837; galapaganus Pfeiffer, 1855; irregularis Pfeiffer, 1848; jacobi Sowerby I, 1833; lycodus Dall, 1917; montivaga d’Orbigny, 1835; munsterii d’Orbigny, 1837; nucula Pfeiffer, 1853; nux Broderip, 1832; orbignyi Pfeiffer, 1846; pallidus Reibisch, 1892; paziana d’Orbigny, 1835; perspectivus Pfeiffer, 1846; phlegonis Dall and Ochsner, 1928; quitensis Pfeiffer, 1848; rimatus Pfeiffer, 1847; rivasii d’Orbigny, 1837; rocayana d’Orbigny, 1835; rugiferus Sowerby I, 1833; rugulosus Sowerby I, 1833; sculpturatus Pfeiffer, 1846; sugillatus Pfeiffer, 1857; terebra Reibisch, 1892; trichoda d’Orbigny, 1835; unifasciatus Sowerby I, 1833; ustulatus Sowerby I, 1833; ventrosus Reibisch, 1892; verrucosus Pfeiffer, 1855; wolfi Reibisch, 1892.
altoperuvianus Reeve, 1849; atahualpa Dohrn, 1863; binneyanus Pfeiffer, 1857; cora d’Orbigny, 1835; decussatus Reeve, 1849; excoriatus Pfeiffer, 1855; lobbii Reeve, 1849; myristicus Reeve, 1849; patasensis Pfeiffer, 1858; platystomus Pfeiffer, 1858; ptychostylus Pfeiffer, 1858.
crichtoni Broderip, 1836; illustris Rolle, 1905.
martinicensis Pfeiffer, 1846; pilosus Guppy, 1871; sanctaeluciae E.A. Smith, 1889.
juarezi Pfeiffer, 1866; liquabilis Reeve, 1848; ragsdalei Pilsbry, 1890; schiedeanus Pfeiffer, 1841.
baroni (Helix) Fulton, 1896; baroni (Bulimulus) Fulton, 1897; chiletensis Weyrauch, 1967; cretaceus Pfeiffer, 1855; grandiventris Weyrauch, 1960; latecolumellaris Preston, 1909; proteus Broderip in
meleagris Pfeiffer, 1853; nigrolimbatus Pfeiffer, 1853.
clarus Pfeiffer, 1857; dukinfieldi Melvill, 1900; gelidus Reeve, 1849; nivalis d’Orbigny, 1835; pallens Reeve, 1849; sowerbyi Pfeiffer, 1847.
p. 17: Bulimus amandus Pfeiffer, 1855: registration number should read 1975457.
p. 49: Drymaeus conicus da Costa, 1907: registration number for paralectotype should read 1907.11.21.32.
p. 69: Bulinus eschariferus Sowerby I, 1838: Type material should read NHMUK 1975173, five possible syntypes (Cuming coll.).
p. 101: Bulimus jussieui Pfeiffer, 1846: Type material should read NHMUK 1975170, lectotype and one paralectotype (Cuming coll.).
p. 180: Bulimus sisalensis Morelet, 1849: Remarks should read
p. 238: Bulimus gruneri Pfeiffer, 1846: figured specimen is not the lectotype but of paralectotype NHMUK 20100563/1.
p. 259, Fig. 45D–F: the figured specimen is not the lectotype but one of the pralectotypes.
This is additional to the taxa excluded in the previous papers (
Bulimus cucullus
Bulimulus glandiniformis
Thaumastus species. i–ii T. taunaisii (Férussac, 1822), lectotype of Bulimus achilles Pfeiffer, 1853 NHMUK 1975268 (H = 58.0) iii–iv T. largillierti (Philippi, 1842), holotype Bulimus consimilis Reeve, 1848 NHMUK 20030189 (H = 52.9) v–vi T. foveolatus (Reeve, 1849), lectotype NHMUK 1975275 (H = 71.5).
i–ii Leiostracus subtuszonatus (Pilsbry, 1899), probable paralectotype of Bulimus onager Reeve, 1848 NHMUK 20130094 (H = 29.0) iii–vi Rhinus species. iii–iv R. constrictus (Pfeiffer, 1841), lectotype of Bulimus hyaloideus Pfeiffer, 1855 NHMUK 1975412 (H = 20.6) v–vi Rhinus ovulum (Reeve, 1849), lectotype NHMUK 1975416 (H = 20.1).
This research received support from the SYNTHESYS Project (http://www.synthesys.info) which is financed by European Community Research Infrastructure Action under the FP7 Integrating Activities Programme. For all their help during ASHB’s stay at the National History Museum, he is very much indebted to J. Pickering, Andreia Salvador and Kathie Way of the Mollusca Section. A special word of thanks is due to staff of the NHMUK Photo Unit for handling the many requests for images (P. Crabb, P. Hurst, H. Taylor). Fred Naggs helped us with suggestions for non-Orthalicoid taxa. Paul Callomon and Gary Rosenberg (Philadelphia) commented on our previous paper, which has led us to add a reconsideration of one taxon. David Campbell (Boiling Springs) raised our awareness on a nomenclatural issue. Finally, the comments of Francisco Borrero and an anonymous reviewer has allowed us to improve this paper which concludes this series.
List of taxa for which Orthalicoidea types are extant, or discussed, in the NHMUK collection
Remarks. This list has been compiled from I: Amphibulimidae (
abruptus Rolle, 1904—III, 7
abscissus Pfeiffer, 1855—III, 8
abyssorum d’Orbigny, 1835—III, 9; ※III, 91
acalles Pfeiffer, 1853—III, 9
acervatus Pfeiffer, 1857—III, 10
achatellinus Forbes, 1850—III, 10
achilles Pfeiffer, 1853—IV, 5
acuminatus da Costa, 1906—III, 11
adamsonii J.E. Gray, 1834†—IV, 41
adoptus Reeve, 1849—I, 14
aenea Pfeiffer, 1861—IV, 5
aequatorianus E.A. Smith, 1877—III, 11; IV, 50
aequatorius Pfeiffer, 1853—III, 11
aestivus Pfeiffer, 1857—III, 12
affinis Broderip in
agueroi Weyrauch, 1960—III, 13
aileenae Breure, 1978—III, 13
alauda Hupé, 1857—※III, 193
alba Crosse, 1874—II, 5
alba Sowerby I in J.E.
alabastrinus da Costa, 1906—III, 14
albemarlensis Dall, 1917—III, 14
albicans Broderip in
albicolor Morelet, 1863—III, 15
albolabiatus E.A. Smith, 1877—III, 16
albus Sowerby I, 1833—III, 16
alexander Crosse, 1855—※II, 27
altoperuvianus Reeve, 1849—III, 17
alutaceus Reeve, 1849—IV, 6
amandus Pfeiffer, 1855—III, 17; IV, 51
ambagiosus Suter, 1906—※II, 32
ambustus Reeve, 1849—III, 18; ※III, 41
andicola Pfeiffer, 1847—III, 18
andoicus Morelet, 1863—III, 18
angasianus Pfeiffer, 1864—II, 5
angrandianus Pilsbry, 1897♦—※III, 164
angustus da Costa, 1906—III, 19
anthisanensis Pfeiffer, 1853—III, 19; ※III, 186; IV, 50
antioquensis Pfeiffer, 1855—III, 20
apertus Pfeiffer, 1855—III, 20
apicepunctata Preston, 1914—III, 21
apiculatus J.E. Gray, 1834—III, 21
apodemeta d’Orbigny, 1835—III, 22; ※III, 148
appendiculata Pfeiffer, 1847—I, 15
approximata Fulton, 1896—IV, 6
aquilus Reeve, 1848—III, 22; IV, 50
arcuatostriatus Pfeiffer, 1855—III, 23
ascendens Pfeiffer, 1853—IV, 7
asopeus Gassies, 1871—II, 6
atacamensis Pfeiffer, 1856—III, 23
atahualpa Dohrn, 1863—III, 24
attenuatus Pfeiffer, 1853—III, 24
atramentaria Pfeiffer, 1855†—IV, 41
aulacostylus Pfeiffer, 1853†—IV, 41
aureolus Guppy, 1866—III, 25
aurifluus Pfeiffer, 1857—III, 25
auriformis da Costa, 1904—I, 16
auris Pfeiffer, 1866—III, 26
aurissciuri Guppy, 1866—I, 16
backhuysi Delsaerdt, 2010—II, 6
badius Sowerby, 1835—III, 26; IV, 50
bairdii Reeve, 1848—II, 6
balsanus Morelet, 1863—III, 27
baranguillanus Pfeiffer, 1853—※III, 20; III, 27
barbadensis Pfeiffer, 1853—III, 28
baroni (Helix) Fulton, 1896—III, 28
baroni (Bulimulus) Fulton, 1897—III, 29
bartletti H. Adams, 1867—III, 29
bellus da Costa, 1906—III, 30
bensoni Reeve, 1849—IV, 7
bicolor Sowerby I, 1835—※III, 27; III, 30; ※III, 155; IV, 50
bifulguratus Reeve, 1849—IV, 8
bilabiatus Broderip and Sowerby I, 1829†—IV, 41
bilineatus Sowerby I, 1833†—IV, 41
binneyanus Pfeiffer, 1857—III, 31
binominis E.A. Smith, 1895—※III, 106
bivittatus Sowerby I, 1833†—※III, 77; IV, 41
blainvilleanus Pfeiffer, 1848†—IV, 41
blandi Pilsbry, 1897—※III, 30
bogotensis Pfeiffer, 1855—III, 31
bolivarii d’Orbigny, 1835—III, 32
bolivianus Pfeiffer, 1846—III, 32
bolivianus Reeve, 1848†—IV, 41
bonariensis Rafinesque, 1833—※III, 124
boucardi da Costa, 1907—III, 33
boucardi Pfeiffer, 1860—IV, 8
bourcieri Pfeiffer, 1853—III, 33
bowkeri Sowerby III, 1890—II, 7
brachysoma d’Orbigny, 1835—III, 34
brazieri Angas, 1871—II, 7
brephoides d’Orbigny, 1835—IV, 9
broadwayi E.A. Smith, 1896—III, 34
broderipii Sowerby I, 1832—II, 8; ※II, 16; ※II, 17; ※II, 35; ※II, 43
bruggeni Breure, 1978—I, 17
brunneum Verdcourt, 1991—II, 8
buckleyi Higgins, 1872—IV, 9
buckleyi Sowerby III, 1895—III, 35
bugabensis Martens, 1893—III, 35
bulbulus Gassies, 1871—II, 9
cacticolus Reeve, 1849—III, 36
cactivorus Broderip in
cactorum d’Orbigny, 1835—III, 36
caledonicus Petit de la Saussaye, 1845—※II, 16
californicus Reeve, 1848—III, 37
caliginosus Reeve, 1849—III, 37; IV, 50
calus E.A. Smith, 1891—II, 9
canaliculatus Pfeiffer, 1845—III, 37
cancellata da Costa, 1906—III, 38
cantatus Reeve, 1848†—IV, 41
cardinalis Pfeiffer, 1853†—IV, 41
carinatum Pfeiffer, 1853—II, 10
castaneostrigatus da Costa, 1906—III, 38
castaneus Pfeiffer, 1845—I, 17
castelnaui Pfeiffer, 1857†—IV, 41
castrensis Pfeiffer, 1847†—IV, 41
castus Pfeiffer, 1847—III, 39
catharinae Pfeiffer, 1857—II, 10
cathcartiae Reeve, 1848—I, 17
catlowiae Pfeiffer, 1853—III, 39
caucaensis da Costa, 1898—III, 40
ceratacme Pfeiffer, 1855—III, 40
cercicola Morelet, 1863—III, 40
chacoensis Preston, 1907†—IV, 41
chamaeleon Pfeiffer, 1855—III, 41
chamayensis Weyrauch, 1967—III, 41
championi Martens, 1893—III, 42
charpentieri Pfeiffer, 1850—II, 10
chemnitzioides Forbes, 1850—III, 42
chiletensis Weyrauch, 1967—III, 43
chimborasensis Reeve, 1848—III, 43
chiriquensis da Costa, 1901—III, 43
chrysostoma Moricand, 1836—※III, 190
cinereus Reeve, 1849—III, 44
citrinovitrea Moricand, 1836—※IV, 24
citronellus Angas, 1879—III, 44
clarus Pfeiffer, 1857—III, 45
clathratus Pfeiffer, 1858—III, 45
clouei Pfeiffer, 1857—IV, 10
coagulatus Reeve, 1849—III, 46; IV, 50
coarctatus Pfeiffer, 1845—III, 46
coerulescens Pfeiffer, 1858†—IV, 41
coloratus Nyst, 1845※—I, 23
columbianus Lea, 1838—※III, 84; ※III, 86
columbiensis Pfeiffer, 1855—III, 47
columellaris Reeve, 1849†—IV, 41
compactus Fulton, 1902—III, 47
confinus Reeve, 1850†—IV, 41
confluens Pfeiffer, 1855—III, 48
confusus Reeve, 1848—III, 48
conicus da Costa, 1907—III, 49; IV, 51
coniformis Pfeiffer, 1847†—IV, 41
consimilis Reeve, 1848—IV, 10
conspersus Sowerby I, 1833—III, 49
constrictus Pfeiffer, 1841—※IV, 18
constrictus Reeve, 1848†—IV, #42
contortuplicatus Reeve, 1850†—IV, 42
convexus Pfeiffer, 1855—III, 50; ※III, 153
coquimbensis Broderip in
cora d’Orbigny, 1835—III, 50
corderoi Klappenbach, 1958—II, 11
coriaceus Pfeiffer, 1857—III, 51
corneus Sowerby I, 1833†—※III, 135; IV, 42
corpulentus Gassies, 1871—II, 11
corrugatus Guppy, 1866—IV, 10
corrugatus King in
corticosus Sowerby III, 1895—I, 18
costatus Pfeiffer, 1848—II, 12
costatus Weyrauch, 1960—III, 51
costifer Weyrauch, 1960—III, 51
cotopaxiensis Pfeiffer, 1853—III, 52; IV, 50
coturnix Sowerby I, 1832—II, 12
crassilabrum Garrett, 1872—II, 12
crepundia d’Orbigny, 1835—III, 52
cretaceus Pfeiffer, 1855—※III, 29; III, 53
crichtoni Broderip, 1836—III, 54; ※III, 93
cucullus Morelet, 1849*—IV, 51
culminea d’Orbigny, 1835—※III, 49; III, 54; ※III, 101; ※III, 111
cumingi Pfeiffer, 1861—IV, 11
cumingii ‘Newcomb’ Pfeiffer, 1849—II, 13
cuninculinsulae Cox, 1872—II, 13
curianianus Reeve, 1849†—IV, 42
curtus Reibisch, 1892—III, 55
cuticula Pfeiffer, 1855—III, 55
cuzcoensis Reeve, 1849—III, 56
cylindricus Fulton, 1907—II, 14
dacostae Sowerby III, 1892—III, 56
darwini Pfeiffer, 1846—III, 56
dealbatus Say, 1821—※III, 110; ※III, 165
deburghiae Reeve, 1859—IV, 12; ※IV, 16
decoloratus Sowerby I, 1833†—IV, 42
decussata Pfeiffer, 1856—IV, 12
decussatus Reeve, 1849—III, 57; ※III, 130
dejectus Fulton, 1907♦—IV, 38
delumbis Reeve, 1849—III, 57
demerarensis Pfeiffer, 1861—IV, 13
demotus Reeve, 1850—III, 58
denickei J.E. Gray, 1852—III, 58
dennisoni Reeve, 1848¶—IV, 39
dentata Wood, 1828—II, 14
dentritis Morelet, 1863—III, 59
depictus Reeve, 1849—III, 59
depstus Reeve, 1849—III, 60
derelictus Broderip, 1832—III, 60
deshayesi Pfeiffer, 1845—III, 61
devians Dohrn, 1863—III, 61
diaphanus Pfeiffer, 1855¶—IV, 45
dillwynianus Pfeiffer, 1853—I, 18
discrepans Sowerby I, 1833—III, 62
dissimulans Preston, 1909—I, 19
doliarius da Costa, 1898—I, 19
dombeyanus ‘Férussac’ Pfeiffer, 1846—III, 62
draparnaudi Pfeiffer, 1847†—IV, 42
droueti Pfeiffer, 1857†—IV, 42
dubius Pfeiffer, 1853—III, 62
dukinfieldi Melvill, 1900♦—III, 63
dunkeri Pfeiffer, 1846—III, 64
durangoanus Martens, 1893—III, 64
dutaillyi Pfeiffer, 1857—III, 65
dux Pfeiffer, 1861—II, 15
dysoni Pfeiffer, 1846—III, 65
eddystonensis Pfeiffer, 1855—II, 15
edwardsi Morelet, 1863—III, 65
edwardsianus Gassies, 1863—II, 16
effeminatus Reeve, 1848—III, 66
eganus Pfeiffer, 1853†—IV, 42
elaeodes Pfeiffer, 1853—I, 46
elata Gould, 1847—IV, 13
electrum Reeve, 1848—III, 66
elegans Pfeiffer, 1842—II, 16
elongata d’Orbigny, 1837—II, 17
elongatus Röding, 1789—※III, 22
elsteri da Costa, 1901—III, 67
emaciatus Morelet, 1863—III, 67
emeus Say, 1830—※III, 92
ephippium Ancey, 1904—IV, 14
episcopalis Pfeiffer, 1855—I, 20
erectus Reeve, 1849—III, 68
eros Angas, 1878—I, 20
erosus Broderip in
erubescens Pfeiffer, 1847—III, 69
erythrostoma Sowerby I, 1833†—IV, 42
eschariferus Sowerby I, 1838—III, 69; IV, 51
euryomphalus Jonas, 1844※—I, 27
excoriatus Pfeiffer, 1855—III, 70
exornatus Reeve, 1849—III, 70
exoticus da Costa, 1901—III, 70
expansus Pfeiffer, 1848—※III, 177
expatriatus Preston, 1909—III, 71
fabrefactus Reeve, 1848—III, 71
falcicula Gassies, 1871—II, 17
fallax Pfeiffer, 1853—III, 72
farrisi Pfeiffer, 1858—III, 72
felix Pfeiffer, 1862—III, 73; IV, 51
fenestratus Pfeiffer, 1846—III, 73
fenestrellus Martens, 1864—※III, 82
feriatus Reeve, 1848—III, 74
fernandezae Weyrauch, 1958—III, 74
ferrugineus Reeve, 1849—III, 75
fibratus Martyn, 1784—※II, 6; ※II, 7; ※II, 8; ※II, 17; ※II, 30; ※II,41
fidustus Reeve, 1849—III, 75
filaris Pfeiffer, 1853—III, 76; IV, 50
flavescens King in
flavidulus E.A. Smith, 1877—III, 76
flexilabris Pfeiffer, 1853—III, 76
flexuosus Pfeiffer, 1853—III, 77
floridanus Pfeiffer, 1857—III, 77
fontainii d’Orbigny, 1838—III, 78
fourmiersi d’Orbigny, 1837—III, 78
foveolatus Pfeiffer, 1848—IV, 14
foxi Clench, 1950—II, 18
fraseri Pfeiffer, 1858—IV, 15
fucatus Reeve, 1849—III, 79
fuligineus Pfeiffer, 1853—II, 18
funckii Nyst, 1843—※I, 14
fuscagula d’Orbigny, 1837—II, 19
fuscobasis E.A. Smith, 1877—III, 79
fuscus Guilding, 1828—※III, 28
fusiformis Menke, 1828—※II, 10
fusoides d’Orbigny, 1835—III, 80
gabbi Angas, 1879—III, 80
galapaganus Pfeiffer, 1855—III, 81
gargantua Férussac, 1821—※II, 30
gatopensis Crosse, 1870—II, 19
gayi Pfeiffer, 1857—III, 81; ※III, 193
gealei H. Adams, 1867—III, 82
gelidus Reeve, 1849♦—III, 82
geometricus Pfeiffer, 1846—III, 83
glandiniformis Sowerby III, 1892*—IV, 51
glomeratus Weyrauch, 1960—III, 83
gloriosus Pfeiffer, 1862—IV, 15
gomesae da Silva & Thomé, 2006—IV, 16
goroensis Souverbie, 1870—※II, 20
gracilis E.A. Smith, 1902—IV, 16
grammica Crosse, 1870—II, 20
granadensis Pfeiffer, 1848—※III, 58; ※III, 93; ※III, 112
grandiventris Weyrauch, 1960—III, 84
granulosus Broderip in
gravesii King in
grayanus Pfeiffer, 1845—II, 20
grenadensis Guppy, 1868†—IV, 42
gruneri Pfeiffer, 1846—III, 84
guadalupensis Bruguière, 1789—※III, 171
guarani d’Orbigny, 1835—II, 20
gueinzii Pfeiffer, 1857—III, 85
guentheri Sowerby III, 1892†—IV, 42
guerini Pfeiffer, 1846—I, 21
guestieri Gassies, 1869—※II, 19; ※II, 22; ※II, 32
guppyi E.A. Smith, 1891—II, 21
guttatus Broderip, 1832—III, 85
guttula Pfeiffer, 1854†—IV, 42
haasi Weyrauch, 1960—※III, 121
habeli Dall, 1892—※III, 191
hachensis Reeve, 1850—III, 86
hamiltoni Reeve, 1849—III, 86
haplochrous Pfeiffer, 1855—III, 87
hargravesi Cox, 1871—II, 21
hartwegi Pfeiffer, 1846—IV, 17
hegewischi Pfeiffer, 1842†—IV, 42
heloica d’Orbigny, 1835—III, 87
hennahi J.E. Gray, 1828†—※III, 36; IV, 42
hepatostomus Pfeiffer, 1861—III, 88
hidalgoi da Costa, 1898—III, 88
hoffmanni Martens, 1893—※III, 38
holostoma Pfeiffer, 1846—III, 89
hondurasanus Pfeiffer, 1846—III, 89
huascensis Reeve, 1848—III, 90
humboldtii Reeve, 1849—III, 90
hyaloideus Pfeiffer, 1855—IV, 17
hyematus Reeve, 1848†—IV, 42
hygrohylaea d’Orbigny, 1835—III, 91
hypozonus Martens, 1893—III, 91
ictericus Martens, 1893†—IV, 42
ignavus Reeve, 1849—III, 92
illustris Rolle, 1904—III, 92
imeldae Weyrauch, 1958—※III, 54
immaculatus C.B. Adams in Reeve, 1850—III, 93
imperfectus Guppy, 1866—※III, 197
inca d’Orbigny, 1835—IV, 18
inaequalis Pfeiffer, 1857†—IV, 42
incarnatus Pfeiffer, 1855—III, 93
inclinatus Pfeiffer, 1862—III, 94
incognita da Costa, 1907—III, 94
incrassatus Pfeiffer, 1853—III, 94
indentatus da Costa, 1901—IV, 18
inermis Morelet, 1851*—III, 214
inflatus Broderip, 1836†—IV, 42
infundibulum Gassies, 1871—II, 22
infundibulum Pfeiffer, 1853—III, 95
inglorius Reeve, 1848—III, 95
insolitus Preston, 1909—IV, 19
integer Pfeiffer, 1855—IV, 19
interruptus Preston, 1909—III, 96; ※IV, 42
inusitatus Fulton, 1900—III, 96
inutilis Reeve, 1850—III, 97
iodostylus Pfeiffer, 1861—III, 97
iostoma Sowerby I, 1824†—IV, 43
iris Pfeiffer, 1853—IV, 20
irregularis Pfeiffer, 1848—III, 98
irroratus Reeve, 1849—IV, 20
ischnus Pilsbry, 1902—※III, 192
istapensis Crosse & Fischer, 1873—III, 98
jacobi Sowerby I, 1833—III, 99; ※III, 142
janeirensis Sowerby I, 1833†—IV, 43
jansoni Martens, 1893—III, 99
jeffreysi Pfeiffer, 1852—IV, 21
jonasi Pfeiffer, 1846—III, 100
josephus Angas, 1878—III, 100
juarezi Pfeiffer, 1866—III, 101
jubeus Fulton, 1908—I, 21
jucundus Pfeiffer, 1855†—IV, 43
jussieui Pfeiffer, 1846—III, 101; IV, 51
juvenilis Pfeiffer, 1855—III, 102
kathiae Breure, 1978—III, 102
kelletti Reeve, 1850—IV, 21
keppelli Pfeiffer, 1853—III, 102
kingii J.E. Gray, 1825—II, 22
knorri Pfeiffer in Philippi, 1846—III, 103
koppeli Sowerby III, 1892—III, 103
koroensis Garrett, 1872—II, 23
kreftii Cox, 1872—II, 23
labeo Broderip, 1828†—IV, 43
labeo Reeve, 1848—※IV, 35
lacerta Pfeiffer, 1855—I, 22
lactifluus Pfeiffer, 1857—III, 104
laetus Reeve, 1849—III, 104
lamarckianus Pfeiffer, 1848—I, 23
lamas Higgins, 1868—III, 105
largillierti Philippi, 1842—※IV, 10
lascellianus E.A. Smith, 1895—III, 105
latecolumellaris Preston, 1909—III, 106
latilabris Pfeiffer, 1855—I, 23
lattrei Pfeiffer in Philippi, 1846—III, 106
laurentii Sowerby I, 1833†—IV, 43
laxostylus Rolle, 1904—III, 107
leeuwinensis E.A. Smith, 1894—II, 24
lesueureanus Morelet, 1860—III, 107
lichnorum d’Orbigny, 1835—III, 108
lilacinus Reeve, 1849—III, 108; ※III, 144
limensis Reeve, 1849—III, 109
limonoica d’Orbigny, 1835—III, 109
lindeni Reeve, 1848†—IV, 43
linostoma d’Orbigny, 1835—III, 110
linterae Sowerby III, 1890—I, 24
liquabilis Reeve, 1848—III, 110
lirinus Morelet, 1851—III, 111
listeri Wood, 1828†—IV, 43
lithoica d’Orbigny, 1835—III, 111
lividus Reeve, 1850—III, 112
lobbii Reeve, 1849—III, 112; ※III, 159
longinquus Morelet, 1863—III, 112
longulus ‘Behn’ Pfeiffer, 1859—II, 24
lophoica d’Orbigny, 1835—III, 113
loveni Pfeiffer, 1848—I, 24
loxanus Higgins, 1872—¶III, 113
loxensis Pfeiffer, 1846—III, 114
loxostomus Pfeiffer, 1853—IV, 22
lucidus da Costa, 1898—III, 114
lucidus Reeve, 1848—III, 115
luridus Pfeiffer, 1863—III, 115
lusorius Pfeiffer, 1855—III, 116
lycodus Dall, 1917—III, 116
macandrewi Sowerby III, 1889†—IV, 43
macgillivrayi Pfeiffer, 1855—II, 25
magnifica Pfeiffer, 1848—IV, 22
magnificus Grateloup, 1839—IV, 23
mahogany Pfeiffer, 1841—※IV, 14
major d’Orbigny, 1837—II, 25; IV, 45
malleatus da Costa, 1898—III, 117
manupictus Reeve, 1848—III, 117
marcidus Pfeiffer, 1853—III, 117
marmarina d’Orbigny, 1835—III, 118
marmatensis Pfeiffer, 1855—IV, 24
marmoratus Dunker in Philippi, 1844—I, 25
martinicensis Pfeiffer, 1846—III, 118
mars Pfeiffer, 1861—IV, 24
mejillonensis Pfeiffer, 1857—III, 119
meleagris Pfeiffer, 1853—III, 119
melo Quoy and Gaimard, 1832—※II, 31
meobambensis Pfeiffer, 1855—IV, 25
meridanus Pfeiffer, 1846—III, 120
metagyra Pilsbry & Olsson, 1949—III, 120
mexicanus Lamarck, 1822—※III, 90
miersi Pfeiffer, 1856—IV, 25
miliola d’Orbigny, 1835¶—IV, 39
miltocheilus Reeve, 1848—II, 26
minor d’Orbigny, 1837—II, 26
minor Weyrauch, 1960—III, 121
modestus Broderip in
mollicellus Reeve, 1849—III, 121
monachus Pfeiffer, 1857—III, 122
moniezi Dautzenberg, 1896—III, 122
monilifer Reeve, 1848—III, 123
montagnei d’Orbigny, 1837—III, 123
montevidensis Pfeiffer, 1846—III, 124
montivaga d’Orbigny, 1835—III, 124
mordani Breure, 1978—III, 125
moricandi Pfeiffer, 1847—III, 125
moritinctus Martens, 1893—III, 126
mossi E.A. Smith, 1896—III, 126
moussoni Pfeiffer, 1853—III, 127
muliebris Reeve, 1849—III, 127
multilineatus Say, 1825—※III, 180
multispira da Costa, 1904—III, 128
munsterii d’Orbigny, 1837—III, 128
murrea Reeve, 1849—IV, 26
murrinus Reeve, 1848—III, 129; ※III, 151
musivus Pfeiffer, 1855—III, 129
mutabilis Broderip in
myristicus Reeve, 1849—III, 130
nanus Reeve, 1849—III, 130
napo Angas, 1878—III, 130
navarrensis Angas, 1878†—IV, 43
nebulosus Martens, 1893†—IV, 43
necouensis Gassies, 1871—II, 27
neglectus Pfeiffer, 1847—II, 27; ※II, 28
nigrofasciatus Pfeiffer in Philippi, 1846—III, 131
nigrolimbatus Pfeiffer, 1853—III, 132
nigropileatus Reeve, 1849—※III, 27; III, 132; ※III, 166
nigroumbilicatus Preston, 1907†—IV, 43
nitelinus Reeve, 1849—III, 132
nitidus Broderip in
nivalis d’Orbigny, 1835♦—III, 134
niveus Preston, 1909—IV, 26
notabilis da Costa, 1906—III, 134
notatus da Costa, 1906—III, 134
nubeculatus Pfeiffer, 1853—III, 135
nubilus Preston, 1903—III, 135
nucinus Reeve, 1850—III, 136; IV, 50
nucula Pfeiffer, 1853—III, 136
nuptialis Melvill & Ponsonby, 1894—II, 28
nux Broderip, 1832—※III, 95; III, 137; ※III, 204
nystianus Pfeiffer, 1853—III, 137
obliquistriatus da Costa, 1901—III, 138
obliquus Reeve, 1849—※IV, 21; IV, 26
oblitus Reeve, 1848—II, 28
occultus Reeve, 1849—II, 28; ※II, 31
ochraceus Morelet, 1863—III, 138; IV, 50
ochrocheilus E.A. Smith, 1877—III, 139
ochrostoma Garrett, 1872—II, 29
odontostoma Sowerby I, 1824—II, 29
onager Reeve, 1848—※IV, 34
onca d’Orbigny, 1835—I, 25
opalinus Sowerby I, 1833—IV, 27
orbignyi Pfeiffer, 1846—※III, 21; III, 139
oreades d’Orbigny, 1835—III, 140
orobaena d’Orbigny, 1835¶—IV, 43
orophilus Morelet, 1860—※III, 15; ※III, 41; ※III, 108; III, 140
orthostoma E.A. Smith, 1877—III, 141
otostomus Pfeiffer, 1855—I, 27
ouensis Gassies, 1870—II, 30, IV, 45
ouveanus Dotzauer in Mousson, 1869—※II, 5; ※II, 39
ovulum Reeve, 1849—IV, 27
pallens Reeve, 1849♦—III, 141
pallidior Sowerby I, 1833†—IV, 43
pallidus Preston, 1909—IV, 46
pallidus Reibisch, 1892—III, 142
panamensis Broderip, 1832—III, 142
papillatus Morelet, 1860—III, 143
paposensis Pfeiffer, 1856—III, 143
parallelus Pfeiffer, 1857—II, 30
pardalina Guppy, 1868—I, 27
pardalis Reeve, 1848†—IV, 43
patagonica d’Orbigny, 1835—II, 31
patasensis Pfeiffer, 1858—III, 144
patricius Reeve, 1849—III, 144
paucicostatus Breure, 1978—III, 144
paziana d’Orbigny, 1835—III, 145
peaki Breure, 1978—III, 145; IV, 50
peelii Reeve, 1859†—IV, 43
pentadinus d’Orbigny, 1835—※I, 26
pentlandi Reeve 1849†—IV, 43
perdix Pfeiffer, 1848—I, 27
perenensis da Costa, 1901—III, 146
pergracilis Rolle, 1904—III, 146
perlucidus Spix, 1827—※IV, 27
perspectivus Pfeiffer, 1846—III, 147
pertristis Pfeiffer, 1855—※III, 197
pervariabilis Pfeiffer, 1853—III, 147
pervius Pfeiffer, 1853—III, 148
pessulatus Reeve, 1848—III, 148
petenensis Morelet, 1851—III, 149
petiti Pfeiffer, 1846—III, 149
philippii Pfeiffer, 1842—III, 150
phlegonis Dall & Ochsner, 1928—III, 150
phlogera d’Orbigny, 1835—IV, 28
phoebus Pfeiffer, 1863—IV, 28
phryne Pfeiffer, 1863—III, 151
physoides Reeve, 1849—II, 31
pictus Pfeiffer, 1855—III, 151
pilosus Guppy, 1871—III, 151
pinicola Gassies, 1870—II, 32
piperitus Sowerby I, 1837—I, 28
pittieri Martens, 1893†—IV, 43
platystomus Pfeiffer, 1858—III, 152
plectostylus Pfeiffer, 1848—I, 30
plicatoliratus da Costa, 1898—III, 152
pliculatus Pfeiffer, 1857—III, 153
plumbeus Pfeiffer, 1855—IV, 29
poecila d’Orbigny, 1835—III, 153
polymorpha d’Orbigny, 1835—III, 154; IV, 50
ponsonbyi da Costa, 1907—III, 155
porphyrius Pfeiffer, 1847—IV, 30
porphyrostomus Pfeiffer, 1851—※II, 39
powisiana Petit de la Saussaye, 1843—※IV, 43
praetextus Reeve, 1849—III, 155
prestoni da Costa, 1906—III, 156
primula Reeve, 1848—III, 156
primularis Reeve, 1849†—IV, 43
princeps Breoderip in
principalis Sowerby II, 1849†—IV, 43
priscus Powell, 1938—II, 32
progastor d’Orbigny, 1835¶—IV, 40
proteus Broderip, 1832—III, 157
protractus Pfeiffer, 1855—III, 157
pruinosus Sowerby I, 1833—III, 158
pseudofusoides da Costa, 1906—III, 158
ptychostylus Pfeiffer, 1858—III, 159
puellaris Reeve, 1850—III, 159
pulchellus Broderip in
pulchellus Sowerby I, 1833†—IV, 43
pulcherrimus H. Adams, 1867—III, 160
pulicarius Reeve, 1848—I, 31
pumilio Rehder, 1945—※III, 130
punctatus da Costa, 1907—III, 160
punctulifer Sowerby I, 1833—II, 33
pupiformis Broderip, 1832—III, 161
purpuratus Reeve, 1849—III, 161; IV, 50
pustulosus Broderip, 1832—III, 162
pygmeus Weyrauch, 1960—※III, 51
pyrostomus Pfeiffer, 1860—II, 33
quadingensis Connolly, 1929—II, 34
quadrifasciatus Angas, 1878—※III, 131; III, 162
quadricolor Pfeiffer, 1848—I, 31
quechuarum Crawford, 1939—III, 163; IV, 50
quitensis Pfeiffer, 1848—※III, 39; ※III, 98; III, 163
radiatus Morelet, 1863—III, 164
ragsdalei Pilsbry, 1890—III, 164
ramagei E.A. Smith, 1890—II, 34
rawsonis H. Adams, 1873—III, 165
recedens Pfeiffer, 1864—III, 165
recluzianus Pfeiffer, 1847—※III, 135
reconditus Reeve, 1849—III, 166
rectilinearis Pfeiffer, 1855—III, 166
reflexa Pfeiffer, 1842—II, 34
regularis Fulton, 1905—III, 167
rehderi Weyrauch, 1960—III, 167
requieni Pfeiffer, 1853—IV, 30
revinctus Hupé, 1857—※III, 193
rhodacme Pfeiffer, 1842†—IV, 43
rhodinostoma d’Orbigny, 1835—II, 35
rhodocheilus Reeve, 1848—I, 34
rhodolarynx Reeve, 1849—III, 167
rhodostomus J.E. Gray, 1834—II, 35
ridleyi E.A. Smith, 1890—II, 36
rimatus Pfeiffer, 1847—III, 168
rivasii d’Orbigny, 1837—III, 168; ※III, 188
rocayana d’Orbigny, 1835—III, 169
rodriguezae Weyrauch, 1967—III, 169
roseatus Reeve, 1848—III, 170
rosenbergi da Costa, 1906—III, 170
roseolabrum E.A. Smith, 1877—I, 36
rubellus Broderip, 1832†—IV, 44
rubescens Reeve, 1848†—IV, 44
rubrifasciatus Reeve, 1848—III, 171
rubrovariegatus Higgins, 1868—III, 171
rufescens J.E. Gray, 1825—※III, 69
rufovirens Moricand, 1846—※IV, 31
rugiferus Sowerby I, 1833—III, 171
rugulosus Sowerby I, 1833—III, 172
rupicolus Reeve, 1848†—IV, 44
rusticellus Morelet, 1860—III, 172
saccatus Pfeiffer, 1855—III, 173
salomonia Pfeiffer, 1853—IV, 31
salomonis Pfeiffer, 1853—※II, 33; II, 36; IV, 45
salteri Sowerby III, 1890—IV, 31
sanchristovalensis Cox, 1870—II, 37
sanctaeluciae E.A. Smith, 1889—III, 173
sarcochilus Pfeiffer, 1857—IV, 32
sarcodes Pfeiffer, 1846†—IV, 44
saturnus Pfeiffer, 1860—IV, 33
savesi Crosse, 1886—II, 37
sayi Pfeiffer, 1847†—IV, 44
scabiosus Sowerby I, 1833—III, 174
scalaricosta Morelet, 1860—III, 174
scalariformis Broderip, 1832—III, 175
schiedeanus Pfeiffer, 1841—III, 175
schmidti Pfeiffer, 1854—III, 176
scytodes Pfeiffer, 1853†—IV, 44
scitulus Reeve, 1849—III, 176
scitus H. Adams, 1867—III, 176
sculpturatus Pfeiffer, 1846—III, 177
scytodes Pfeiffer, 1853—I, 37
sellersi Cox, 1872—II, 38
selli Preston, 1909—III, 177
senilis Gassies, 1869—IV, 44
serotinus Morelet, 1860—III, 178
serparastrus Say, 1830—※III, 133
serratus Pfeiffer, 1855—III, 178
signifer Pfeiffer, 1855—III, 179
simpliculus Pfeiffer, 1855—III, 179
simulus Morelet, 1851—IV, 33
singularis Morelet, 1857—II, 38
sinistrorsa Crosse, 1884—II, 39
sisalensis Morelet, 1849—III, 180; IV, 51
smithii da Costa, 1898—III, 180
solidus Preston, 1907—III, 181
souvillei Morelet, 1857—II, 39
sowerbyi Pfeiffer, 1847♦—III, 181
spadiceus da Costa, 1906—III, 182
speciosus Pfeiffer, 1854—I, 38
spectatus Reeve, 1849—III, 182
spenceri Tate, 1894—II, 40
spiculatus Morelet, 1860—III, 183
sporadica d’Orbigny, 1835—III, 183
sporadicus Reeve, 1848†—IV, 44
stenacme Pfeiffer, 1857—III, 184
stramineus Guilding, 1824—※III, 115
strangei Pfeiffer, 1855—II, 40
striata Spix, 1827—※II, 26
striatulus Sowerby I, 1833†—IV, 44
striatus ‘King’ Sowerby I, 1833†—IV, 44
strigatus Sowerby I, 1833—※III, 129; ※III, 173; III, 185
studeri Pfeiffer, 1847—※III, 156; III, 185
subfasciatus Pfeiffer, 1853—III, 186; IV, 50
subhybridus da Costa, 1906—III, 186
subinterruptus Pfeiffer, 1853—III, 187
subirroratus da Costa, 1898—IV, 34
subpellucidus E.A. Smith, 1877—III, 187
subroseus Fulton, 1915—II, 41
subtuszonatus Pilsbry, 1899—IV, 34
subventricosus da Costa, 1901—III, 188
succineoides Petit de la Saussaye, 1840—※I, 24
sufflatus Gould, 1859—※III, 101
sugillatus Pfeiffer, 1857—III, 188
sulcosus Pfeiffer, 1841—III, 188
sulphureus Pfeiffer, 1857—※III, 45; III, 189
superfasciatus Gassies, 1871—II, 41
superstriatus Sowerby III, 1890—I, 40
swainsoni Pfeiffer, 1845—III, 189
sykesi da Costa, 1906—III, 190
taquinensis Pfeiffer, 1855—I, 41
tasmanicus Pfeiffer, 1853—II, 41
taunaisii Férussac, 1822—※IV, 5
taylori Pfeiffer, 1861†—IV, 44
taylorianus Reeve, 1849—I, 42
tenuilabris Pfeiffer, 1866—III, 190
terebra Reibisch, 1892—III, 191
terebralis Pfeiffer, 1842—III, 191
thamnoica d’Orbigny, 1835—III, 192
thompsonii Pfeiffer, 1845—IV, 35
tigrinus da Costa, 1898—III, 193
tigris Broderip in
toleratus Fulton, 1903—II, 42
torallyi d’Orbigny, 1835—※III, 128; III, 194
translucens Broderip in
transparens Reeve, 1849—III, 195
trichoda d’Orbigny, 1835—III, 195
tricinctus Reeve, 1848—III, 196
trigonostomus Jonas, 1844—※III, 103
trimarianus Martens, 1893—III, 196
trinitarius E.A. Smith, 1896—III, 197
tristis Pfeiffer, 1855—III, 197
tropicalis Morelet, 1849—III, 198
trujillensis Philippi, 1867—※III, 105
tumidulus Pfeiffer, 1842—III, 198
tupacii d’Orbigny, 1835—III, 199
turbinatus Pfeiffer, 1845—II, 42
turneri Pfeiffer, 1860—II, 43
turritella d’Orbigny, 1835—III, 199
turritellatus Beck, 1838—※III, 200
turritus Broderip in
umbilicaris Souleyet, 1842—III, 200
umbraticus Reeve, 1850—※III, 77
umbricatus Reeve, 1849—III, 201
undulosus Martens, 1893†—IV, 44
unicolor Sowerby I, 1833—※III, 92; ※III, 98; ※III, 149
unifasciatus Sowerby I, 1833—III, 202
ustulatus Sowerby I, 1833—※III, 150; III, 202
varians Broderip in
varicosus Pfeiffer, 1853—III, 203
variegata Pfeiffer, 1842—II, 43
venezuelensis Pfeiffer, 1856†—IV, 44
venosus Reeve, 1848†—IV, 44
ventricosus Preston, 1907†—IV, 44
ventrosus Reibisch, 1892—III, 203
veranyi Pfeiffer, 1848—※I, 38; I, 44
verrucosus Pfeiffer, 1855—III, 203
versicolor Broderip in
veruculum Morelet, 1860—III, 204
vesicalis Pfeiffer, 1853—III, 205
vespertinus Pfeiffer, 1858—III, 205
vexillum Broderip in
vexillum Wood, 1828†—※III, 103; IV, 44
vicaria Fulton, 1896—IV, 35
vicinus Preston, 1907†—IV, 44
victor Pfeiffer, 1854—IV, 36
vilchezi Weyrauch, 1960—III, 206
vincentina E.A. Smith, 1895—IV, 37
vincentinus Pfeiffer, 1846—III, 206
virginalis Pfeiffer, 1856—III, 207
virgultorum Morelet, 1863—III, 207
vitrinoides Reeve, 1848¶—IV, 40
vittatus Broderip in
voithianus Pfeiffer, 1847—III, 208
volsus Fulton, 1907—III, 208
wintlei Finch, 1929—III, 209
wolfi Reibisch, 1892—III, 209
woodwardi Pfeiffer, 1857—III, 210
xanthostoma d’Orbigny, 1835—III, 210
yanamensis Morelet, 1863—IV, 37
yatesi Pfeiffer, 1855—※IV, 36; IV, 38
yungasensis d’Orbigny, 1837—III, 211
zhorquinensis Angas, 1879—III, 211
ziczac da Costa, 1898—III, 212
ziegleri Pfeiffer, 1847—※III, 37; ¶III, 212
ziegleri Reeve, 1849†—IV, 44
zilchi Breure, 1977—I, 45
zilchi Weyrauch, 1958—※III, 83
zoographica d’Orbigny, 1835—III, 213