Neotype ♂, present designation: “China: Beijing, ca. 1400 m, Dongling Mts, 15.Vi.2001, Xiaolongmen, Liu Lang Yu / N39°97, E115°43 [sic], Mixed woodland litter, Leg. J. Cooter + P. Hlavá [sic] / Neotypus ♂ Domene chenpengi Li desig. B. Feldmann & Z. Peng 2014 / Domene chenpengi Li, det. B. Feldmann 2014“ (MNHUB).

The original description is based on a single male specimen from Chang Chun [ca. 43°45'N, 125°27'E], Jing Yue, collected on 30.VII.1985 by Peng Chen (Li et al. 1990). Inquiries into the whereabouts of the holotype at the Northeast Normal University, where the holotype should be deposited, yielded no results. It was looked for in the respective collection, but not found (personal communication Xiu-Qing Yin, one of the authors in Li et al. 1990 and director of the biogeographical office of Northeast Normal University, e-mail 5 May, 2014; personal communication Jing-Ke Li, author of D. chenpengi and guest professor of the Harbin Normal University, e-mail 5 May, 2014). Thus, the type specimen must be regarded as lost. The insufficient description of D. chenpengi, which fails to provide any illustration whatsoever, is in agreement with examined material previously identified as D. dersuuzalai from the Russian Far East and China, particularly regarding the habitus and the characteristic shape of the male sternite VIII with its shallow posterior excision. Moreover, the type locality of D. chenpengi accords with the known distribution of D. dersuuzalai. In the interest of stability of nomenclature, a neotype designation is deemed necessary to stabilize the present interpretation of D. chenpengi and the synonymy with D. dersuuzalai. To this end, a male from the Dongling mountains in Beijing, a locality reasonably close to the type locality, is designated as the neotype. Based on the detailed description of D. dersuuzalai (Gusarov 1992), the species is doubtlessly conspecific with the neotype of D. chenpengi; hence the synonymy proposed above.

(60 exs.). Russian Far East: Primorskiy Kray: 2 exs., Vladivostok env., Sedanka, 28.VII.1992, leg. Beloborodov (NHMB, cFel); 3 exs., Vladivostok, 11.VII.1993, leg. Pütz & Wrase (cSch); 6 exs., N Vladivostok, “Seitengraben des Parwaja Rjetschka Tales”, 1918–1920, leg. Frieb (NHMW, cFel); 1 ex., Kamenushka, 14.–15.VII.1992, leg. Beloborodov (NHMB); 7 exs., Partisansky district, Alexeyevskiy khrebet, 20 km E Sergeyevka, forests near Andreyevka river, 400–800 m, 26.–29.VII.1993, leg. Pütz & Wrase (cSch, cFel); 1 ex., S Artyom town, Ozernyy Kluytch river, 100–300 m, 10.V.–5.VI.2002, leg. Plutenko (cSch); 2 exs., Lazovskiy reserve, 9 km SE Kievka, lodge Petrova env., 3.–8.VI.1994, leg. Sundukov (cPüt); 1 ex., same data, but 9.–13.VI.1995 (cPüt); 1 ex., Lazovskiy district, Kovarinovo, 5 km NE Lazo, spring valley, 1.–5.VI.1995, leg. Sundukov (cFel); 1 ex., Lazovskiy reserve, Kordon Amerika, 134°03'01"E, 43°16'16"N, 14.–17.V.1999, leg. Sundukov (cSch); 2 exs., Lazovskiy reserve, Kordon Amerika, 18.–19.VI.1997, leg. Sundukov (cSch); 1 ex., Lazovskiy reserve, Kordon Petrova, 133°47'55"E, 42°52'14"N, 19.–20.IX.1999, leg. Sundukov (cSch); 1 ex., Lazovskiy reserve, Kordon Proselochny, 134°07'43"E, 43°00'34"N, 4.–6.X.1999, leg. Sundukov (cSch); 3 exs., Sikhote-Alin reserve, Jasnaya estuary, 26.VI.–4.VII.1998, leg. Sundukov (cAss, cSch); 1 ex., Siniy khrebet, 4 km E Evseevki, 7.–9.VIII.1999, leg. Shavrin (cSch); 1♂, 2 ♀♀ [identified by Gusarov 1995 as D. dersuuzalai], Arsenev env., 27.V.–5.VII.1991, leg. Sausa (NHMW). Khabarovskiy Kray: 3 exs., SE Boitsovo, 12 km NE Bikin, 250–350 m, 26.V.–4.VI.1990, leg. Schawaller (cSch). Sakhalin: 5 exs., Moneron Island, 15.VI.–6.VII.2002, leg. Plutenko (cSch, cFel). China: Beijing: 1 ♂, 5 ♀♀, Xiaolongmen, Yan Shan, Dongling Mts, 1400 m, 15.–16.VI.2001, leg. Hlavač & Cooter (cAss, cSch, cFel); 2 ♂♂, Xiaolongmen, Liu Lang Yu, Dongling Mts, 39°58'N, 115°26'E, ca. 1400 m, under fungoid Juglans bark, 15.VI.2001, leg. Cooter & Hlavač (cRou); 1 ♀, Xiaolongmen, Liu Lang Yu, Dongling Mts, 39°58'N, 115°26'E, ca. 1400 m, mixed forest litter, 15.VI.2001, leg. Cooter & Hlavač (cRou); 2 ♂♂, 1 ♀, Xiaolongmen, Mei Yao Yu, Dongling Mts, 39°58'N, 115°26'E, ca. 1400 m, mixed forest litter, 16.VI.2001, leg. Cooter & Hlavač (cRou, cFel); 1 ♂, 2 ♀♀, Miyun County, Wulin Shan, 40°36'N, 117°23'E, 750–850 m, 8.–9.VII.2006, leg. Shen & Tang (SNUC). South Korea: 1 ♂, Gangwon-do, Seorak-san, 1.5 km S Han-gyeryeong pass rest station, roadside forest, 38°05'26"N, 128°24'03"E, 790 m, from wet, fungusy leaf litter, under trunk, rocks, sifted, 9.IX.2010, leg. Makranczy & al. (HNHM).

Measurements (in mm) and ratios: TL 7.15–7.60, FL 3.80–3.95, HL 1.1–1.15, HW 1.00–1.08, AnL 2.55–2.65, NW 0.35–0.38, PL 1.19–1.23, PW 0.95–1.03, EL 0.98–1.03, EW 1.03–1.05, TiL 1.20–1.25, TaL 0.90–0.98, AW 1.08–1.13, AL 0.85–0.90, HL/HW 1.05–1.13, HW/PW 1.02–1.08, HL/PL 0.90–0.95, NW/HW 0.35–0.36, PL/PW 1.17–1.25, EL/PL 0.82–0.85.

Habitus as in Fig. 2A. Head and pronotum blackish brown; elytra brownish with anterior and posterior portions more or less extensively reddish brown; abdomen brownish; legs yellowish brown, except for the slightly paler tarsi; antennae light brown to yellowish brown.

Figure 1. 

Distributions of Domene species in China: D. chenpengi (filled squares; type locality: open square); D. firmicornis (filled and open diamonds); D. immarginata (filled stars); D. malaisei (open triangles); D. reitteri (filled diamond); D. chenae (open star); D. cultrata (filled triangles); D. cuspidata (open circles); D. reducta (filled circles).

Figure 2. 

Habitus of Domene spp., A D. chenpengi B D. firmicornis. Scales: 1.0 mm.

Head orbicular, widest behind eyes; punctation (Fig. 3A) fine and very dense. All antennomeres longer than broad; antennomeres IV–X of equal length; antennomere I 1.9 times, II 1.2 times, III 1.3 times, XI 1.3 times as long as IV. Maxillary palpus slender, preapical joint about 2.5 times as long as broad.

Figure 3. 

Domene chenpengi. A median dorsal portion of head B median portion of pronotum C female tergite VIII D female sternite VIII E female tergites IX–X. F male sternite VII G male sternite VIII H aedeagus in ventral view I aedeagus in lateral view J aedeagus in dorsal view. Scales: A–B, J 0.2 mm; C–I 0.5 mm.

Pronotum about as broad as head, widest in anterior third; lateral margins slightly convex in dorsal view; punctation (Fig. 3B) very fine with interstices forming narrow, longitudinal ridges.

Elytra without distinct longitudinal ridges; disc often more or less impressed; suture elevated in posterior three-fourths; macropunctation coarse, more or less dense and irregular on disc, interstices with shallow and irregular micropunctation; in lateral and posterior portions with distinctly finer and denser punctation. Hind wings present. Protarsomeres I–IV moderately dilated.

Abdomen with fine and very dense punctation on tergites III–VIII; interstices with microreticulation; tergite VIII more or less obtusely triangularly produced posteriorly (Fig. 3C); posterior margin of tergite VII with palisade fringe.

Male. Sternites III–VI unmodified; sternite VII (Fig. 3F) distinctly transverse, with shallow postero-median impression, this impression with a few modified short and black setae posteriorly; sternite VIII (Fig. 3G) transverse, with pronounced impression posteriorly, this impression with distinctly modified short and stout black setae, posterior excision very small; aedeagus as in Figs 3H–J, ventral process stout and apically moderately acute in lateral view; apical portion of dorsal fig long and distinctly sclerotized, basal portion short.

Female. Posterior margin of sternite VIII (Fig. 3D) in the middle with shallow concave excision; genital segments with distinctly sclerotized structure (Fig. 3E).

The similar external morphology, the similar chaetotaxy and shape of the male sternites VII and VIII, and especially the similar shape of the ventral process of the aedeagus suggest that D. chenpengi is closely allied to D. curtipennis from Japan. For illustrations of D. curtipennis see Gusarov (1992: figure 4). Besides its conspicuous male sexual characters, D. chenpengi is distinguished from the Chinese species of the D. malaisei and D. scabripennis groups and from D. procera by its smaller size alone, and from species of the D. reitteri group by its coarser and less densely punctate the elytra.

The currently known distribution ranges from the Russian Far East and Northern China (Beijing, Jilin) to South Korea. The specimens were partly sifted from leaf litter in mixed forest habitats or found under bark and rocks. The elevations range from 100 up to 1400 m.

Domene crassicornis was recorded by Li et al. (1990) from Jilin, the only record of this species from China. This record is evidently based on a misidentification and probably refers to D. procera. Based on available evidence, the distribution of D. crassicornis is restricted to Japan and consequently does not include China. All revised material from the Russian Far East belongs to D. procera.

The sole record of D. curtipennis from China is that by Li (1992) from Liaoning. It is almost certainly based on a misidentification. Based on available evidence, the distribution of D. curtipennis is restricted to Japan. All the examined material from the Russian Far East, South Korea and China belongs to D. chenpengi, suggesting that D. curtipennis does not occur in China.

Examined type specimens of this species are listed in an addendum in Assing and Feldmann (2014). The previously undescribed female sexual characters are as follows: female tergite VIII (Fig. 4C) with shallow postero-median impression and distinctly concave posterior excision; female sternite VIII (Fig. 4D) about as long as broad, posterior margin concave in the middle; sclerotized structure in female genital segments (Fig. 4E) symmetric and very weakly sclerotized. For illustrations of D. firmicornis see Figs 2B, 4 and Assing and Feldmann (2014: figures 36–43).

Figure 4. 

Domene firmicornis. A median dorsal portion of head B median portion of pronotum C female tergite VIIwI D female sternite VIII E female tergites IX–X. F male sternite VII G male sternite VIII H aedeagus in ventral view I aedeagus in lateral view J aedeagus in dorsal view. Scales: A–B 0.2 mm; C–J 0.5 mm.

Holotype ♀: “N. E. Burma, Kambaiti, 2000 m, 4/6.1934, Malaise / HOLOTYPUS [red label] / TYPUS Domene Malaisei O. Scheerpeltz [red label] / Domene Malaisei nov. spec. det. Scheerpeltz, 1941 / 3884 E91” (RMS).

(5 ♂♂, 9 ♀♀). China: Yunnan: 4 ♂♂, 5 ♀♀, Tengchong County, Mingguang, Zizhi, Donghe, 25°42'N, 98°34'E, 2100–2300 m, 01.V.2013, leg. Peng & Song (SNUC, cAss); 4 ♀♀, same data, but 25°42'N, 98°35'E, 2500 m, 30.IV.2013 (SNUC); 1 ♂, Dehong Dai Autonomous Prefecture, mountain range 31 km E Luxi, 24°29'31"N, 98°52'58"E, 2280 m, secondary pine forest with old deciduous trees, litter sifted, 3.VI.2007, leg. Pütz (cFel).

Measurements (in mm) and ratios: Holotype: TL 8.90, FL 5.20, HL 1.38, HW 1.30, PL 1.45, PW 1.25, EL 1.50, HL/HW 0.94, HW/PW 1.10, HL/PL 0.89, PL/PW 1.16, EL/PL 1.03. Additional material: TL 7.90–9.20, FL 4.70–5.05, HL 1.24–1.33, HW 1.17–1.25, AnL 3.17–3.40, NW 0.40–0.46, PL 1.35–1.50, PW 1.15–1.25, EL 1.28–1.45, EW 1.44–1.53, TiL 1.57–1.65, TaL 1.14–1.33, AW 1.26–1.34, AL 1.07–1.18 HL/HW 1.06–1.07, HW/PW 0.99–1.01, HL/PL 0.89–0.93, NW/HW 0.34–0.38, PL/PW 1.15–1.20, EL/PL 0.95–0.97.

Habitus as in Fig. 5. Body black; legs with blackish brown profemora and brown protibiae, basal halves of metafemora light brown, distal halves gradually infuscate; antennae dark brown to brown.

Figure 5. 

Habitus of Domene malaisei. A male B holotype. Scales: 0.5 mm.

Head orbicular, broadest across eyes; punctation (Fig. 6A) coarse, distinctly umbilicate, and very dense, interstices forming very narrow ridges. All antennomeres longer than broad; antennomeres IV–X of equal length; antennomere I 1.3 times, II 0.9 times, III 1.1 times, XI 1.1 times as long as IV. Maxillary palpus very slender, preapical joint 2.8–3.1 times as long as broad.

Figure 6. 

Domene malaisei. A median dorsal portion of head B median portion of pronotum C female tergite VIII D female sternite VIII E female tergites IX–X. F male sternite VII G male sternite VIII H aedeagus in ventral view I aedeagus in lateral view J aedeagus in dorsal view. Scales: A–B 0.2 mm; C–J 0.5 mm.

Pronotum nearly as broad as head, widest in the middle; lateral margins convex in dorsal view; punctation (Fig. 6B) somewhat coarser than that of head; midline with rudiment of fine glossy line.

Elytra without distinct longitudinal ridges; disc more or less weakly impressed; suture elevated in posterior three-fourths; macropunctation coarse, irregular, partly confluent, and partly somewhat seriate; interstices with shallow and irregular micropunctation. Hind wings fully developed. Protarsomeres I–IV moderately dilated.

Abdomen with fine and dense punctation on tergites III–VI; tergite VIII with dense pubescence, posterior margin of tergite VIII broadly and weakly convex (Fig. 6C); interstices with distinct microreticulation; posterior margin of tergite VII with palisade fringe.

Male. Sternites III–VI unmodified; sternite VII (Fig. 6F) distinctly transverse, with very shallow median impression posteriorly, this impression with sparse modified black setae, posterior margin broadly concave; sternite VIII (Fig. 6G) with shallow median impression posteriorly, this impression with stout black setae, posterior excision deep, almost V-shaped, on either side of the posterior excision with dense cluster of dark setae; aedeagus as in Figs 6H–J, ventral process evenly curved and apically acute in lateral view; dorsal fig long, apical portion distinctly sclerotized and apically acute in lateral view, basal portion short.

Female. Posterior margin of sternite VIII (Fig. 6D) broadly convex; genital segments with an asymmetric and weakly sclerotized structure (Fig. 6E).

The similar chaetotaxy and shape of the male sternite VIII and the similar shape of the ventral process of the aedeagus suggest that D. malaisei is closely allied to D. reducta. Domene malaisei is readily distinguished from other species of the group by on average darker coloration, smaller body size, the deeper posterior excision of the male sternite VIII, the evenly curved ventral process of the aedeagus and by the shape of the sclerotized structure in the female genital segments.

The currently known distribution is confined to the type locality Kambaiti in northeastern Myanmar at the border with Yunnan, and two localities in western Yunnan (Fig. 1). The examined non-type specimens were sifted from forest leaf litter at altitudes of 2000–2500 m.

Lectotype ♂, present designation: “Tienmuschan, N. W China, Rtt. / Type / Domene Reitteri Koch det. C. Koch / Holotype 1956 det. Kamp / Holotypus Domene reitteri Koch / Domene reitteri Koch V. L. Gusarov det. 1993 / Lectotypus ♂, Domene reitteri Koch, desig. B. Feldmann 2010” (NHMB).

Paralectotypes 3 ♀♀: „Tienmuschan, N. W China, Rtt. / Cotype / Paratypus Domene reitteri Koch / Domene reitteri Koch V. L. Gusarov det. 1993“; 1 ex. (abdomen missing): “Tienmuschan, N. W China, Rtt. / Cotype” (NHMB).

The original description of D. reitteri is based on an unspecified number of syntypes from “Tienmuschan (nordwestliches [sic] China) ex coll. E. Reitter” (Koch 1939). Five syntypes, one male, three females and one unsexed specimen, were located in the Koch collection at the Naturhistorisches Museum Basel. The male syntype is designated as the lectotype.

(87 ♂♂, 59 ♀♀). China: Zhejiang: 11 ♂♂, 7 ♀♀, Anji County, Longwang Shan, 30°23'59"N 119°26'26"E, 1300–1450 m, 14.V.2013, leg. Hu (SNUC); 31 ♂♂, 14 ♀♀, Longwang Shan, 30°24'28"N 119°26'37"E, 1050–1200 m, 15.V.2013, leg. Li & al. (SNUC); 15 ♂♂, 7 ♀♀, Longwang Shan, Qianmutian, 30°24'N 119°26'E, 1050–1250 m, 08.VI.2012, leg. Yin & Hu (SNUC, cAss); 1 ♂, Longwang Shan, 30°24'N 119°26'E, 1250–1450 m, 14.V.2013, leg. Chen & Pan (SNUC); 3 ♂♂, 5 ♀♀, Longwang Shan, Dongguan, 1250 m, 26.V.2009, leg. Feng & al. (SNUC); 3 ♂♂, 6 ♀♀, Longwang Shan, Qianmutian, 1300 m, 24.V.2009, leg. Feng & al.(SNUC); 8 ♂♂, 10 ♀♀, Longwang Shan, 950–1200 m, 25.IV.2006, leg. He (SNUC); 2 ♂♂, 3 ♀♀, Longwang Shan, Qianmutian, 1300 m, 29.V.2009, leg. Feng & al.(SNUC); 1 ♀, Longwang Shan, Qianmutian, 700–1325 m, 28.VII.2011, leg. Pan (SNUC); 1 ♂, 1 ♀, Tianmu Shan, 1200–1300 m, 25.–29.VII.2011, leg. Chen (SNUC); 2 ♂♂, 2 ♀♀, Tianmu Shan, 300 m, 17.V.2006, leg. Hu & Tang (SNUC); 1 ♀, Tianmu Shan, 1100 m, 24.VII.2011, leg. Hu & Tang (SNUC); 1 ♂, Tianmu Shan, 1500 m, 15.VIII.2010, leg. Hu (SNUC); 1 ♂, Tianmu Shan, 300–400 m, 29.V.2010, leg. Wang (SNUC); 1 ♂, East Tianmu Shan, 1050–1150 m, 13. IV.2011, leg. Peng & Zhu (SNUC); 2 ♂♂, Tianmu Shan, 13.VI.2009, leg. Song (SNUC); 2 ♂♂, Tianmu Shan, 1000 m, 2.V.2009, leg. Song (SNUC); 2 ♂♂, 1 ♀, Tianmu Shan, 1500 m, 15.VIII.2010, leg. Hu (SNUC); 2 ♂♂, 1 ♀, West Tianmu Shan N.R., path to peak of immortals (“Blind Alley”), 30°20'34"N, 119°25'51"E, 1100–1200 m, primary mixed forest, litter moss, sifted, 15.VI.2007, leg. Wrase (cSch, cFel).

Measurements (in mm) and ratios: TL 5.78–8.62, FL 4.16–4.43, HL 1.07–1.17, HW 1.05–1.11, AnL 2.78–3.05, NW 0.37–0.41, PL 1.28–1.35, PW 1.02–1.06, EL 1.07–1.13, EW 1.12–1.20, TiL 1.39–1.44, TaL 0.89–0.96, AW 1.12–1.24, AL 1.07–1.17, HL/HW 1.02–1.05, HW/PW 1.03–1.05, HL/PL 0.84–0.87, NW/HW 0.35–0.37, PL/PW 1.25–1.28, EL/PL 0.84–0.89.

Habitus as in Figs 7A, 8A, 8D. Body blackish brown; legs with dark brown profemora and protibiae, basal halves of metafemora light brown, distal halves gradually infuscate; antennae brown to light brown.

Figure 7. 

Habitus of Domene spp., A D. reitteri B D. chenae. Scales: 1.0 mm.

Figure 8. 

Domene reitteri. A lectotype B male sternite VII–VIII C aedeagus of lectotype in lateral view D paralectotype. Scales: A, D 1.0 mm; B–C 0.5 mm.

Head orbicular, broadest across eyes; punctation (Fig. 9A) moderately coarse, weakly umbilicate, and very dense, interstices forming very narrow ridges. All antennomeres longer than broad; antennomeres IV–X of equal length; antennomere I 1.6 times, II 1.1 times, III 1.3 times, XI 1.3 times as long as IV. Maxillary palpus very slender, preapical joint 2.7–3.2 times as long as broad.

Figure 9. 

Domene reitteri. A median dorsal portion of head B median portion of pronotum C female tergite VIII D female sternite VIII E female tergites IX–X. F male sternite VII G male sternite VIII H aedeagus in ventral view I aedeagus in lateral view J aedeagus in dorsal view. Scales: A–B 0.2 mm; C–J 0.5 mm.

Pronotum slightly narrower than head, widest in the middle; lateral margins convex in dorsal view; punctation (Fig. 9B) somewhat coarser than that of head; midline with rudiment of a fine glossy line.

Elytra without distinct longitudinal ridges; suture weakly elevated; punctation very fine, dense and uniform; hind wings reduced. Protarsomeres I–IV dilated in both sexes.

Abdomen with punctation fine and dense on tergites III–VI, finer and somewhat sparser on tergite VIII, posterior margin of tergite VIII weakly convex in the middle (Fig. 9C); interstices with shallow microreticulation; posterior margin of tergite VII with palisade fringe.

Male. Sternites III–VI unmodified; sternite VII (Figs 8B, 9F) distinctly transverse, with median impression of triangular shape posteriorly, this impression with numerous distinctly modified, short and stout black setae; posterior margin distinctly concave in the middle; sternite VIII (Figs 8B, 9G) transverse, with pronounced and symmetric impression posteriorly, this impression with distinctly modified short and stout black setae, posterior excision small and U-shaped; aedeagus as in Figs 8C, 9H–J, ventral process stout and apically acute; dorsal fig with large and lamellate apical portion, and with short, thin basal portion; internal sac with membranous structures.

Female. Posterior margin of sternite VIII (Fig. 9D) broadly convex; genital segments with weakly asymmetric large and moderately sclerotized structure (Fig. 9E).

The fine, dense and uniform punctation of the elytra, and the similar shape and chaetotaxy of the male sternite VII and sternite VIII suggest that D. reitteri is most closely allied to D. chenae. It is distinguished from D. chenae by the finer punctation of the head and pronotum, the numerous distinctly modified, short and stout black setae on the male sternite VII, the stouter ventral process of the aedeagus and by the shape of the sclerotized structure in the female genital segments.

The distribution is confined to several localities in the Tianmu Shan range in the northwest of Zhejiang. The specimens were sifted from leaf litter in broad-leaved and primary mixed forests at altitudes of 300–1500 m.

(2 ♂♂, 1 ♀). Holotype ♂: “China: Guangxi Prov., Lingui County, Huping N. R., Anjiangping, 25°34'N, 109°57'E, 13.VII.2011 1,200 m, Zhu, Chen & Ma leg. / Holotypus ♂ Domene chenae sp. n., det Peng & Li. 2014” (SNUC). Paratypes: 1 ♂: same data as holotype (SNUC); 1 ♀: same data, but “He & Tang leg.” (SNUC).

The species is named after Yan Chen, who collected some of the type specimens.

Measurements (in mm) and ratios: BL 7.95–8.17, FL 4.55–4.73, HL 1.20–1.24, HW 1.14–1.17, AnL 3.17–3.39, NW 0.43–0.46, PL 1.30–1.37, PW 1.07–1.09, EL 1.11–1.13, EW 1.22–1.24, TiL 1.58–1.66, TaL 0.94–1.02, AW 1.26–1.30, AL 1.12, HL/HW 1.05–1.06, HW/PW 1.06–1.07, HL/PL 0.91–0.92, NW/HW 0.38–0.39, PL/PW 1.21–1.26, EL/PL 0.82–0.85.

Habitus as in Fig. 7B. Body black with distinctly paler abdominal apex; legs with blackish brown profemora and dark brown protibiae, basal halves of metafemora light brown, distal halves gradually infuscate; antennae brown to light brown.

Head orbicular, widest across eyes; punctation (Fig. 10A) coarse, distinctly umbilicate, and very dense, interstices forming very narrow ridges. All antennomeres longer than broad; antennomeres IV–X of equal length; antennomeres I 1.6 times, II 0.9 times, III 1.1 times, XI 1.2 times as long as IV. Maxillary palpus very slender, preapical joint 2.8–2.9 times as long as broad.

Figure 10. 

Domene chenae. A median dorsal portion of head B median portion of pronotum C female tergite VIII D female sternite VIII E female tergites IX–X. F male sternite VII G male sternite VIII H aedeagus in ventral view I aedeagus in lateral view J aedeagus in dorsal view. Scales: A–B 0.2 mm; C–J 0.5 mm.

Pronotum narrower than head, widest in the middle; lateral margins weakly convex in dorsal view; punctation (Fig. 10B) similar to that of head; midline with rudiment of a fine glossy line.

Elytra without distinct longitudinal ridges; suture elevated in posterior two thirds; punctation fine, dense and uniform; interstices without micropunctation. Hind wings probably present. Protarsomeres I–IV moderately dilated.

Abdomen with fine and dense punctation on tergites III–VIII; posterior margin of tergite VIII broadly and weakly convex (Fig. 10C); interstices with shallow microreticulation; posterior margin of tergite VII with palisade fringe.

Male. Sternites III–VI unmodified; sternite VII (Fig. 10F) distinctly transverse, with median impression of triangular shape posteriorly, this impression with moderately modified dark setae, posterior margin broadly concave in the middle; sternite VIII (Fig. 10G) transverse, with shallow median impression posteriorly, this impression with distinctly modified stout and black setae, posterior excision small and U-shaped; aedeagus as in Figs 10H–J, ventral process more slender and curved, apically acute; dorsal fig with distinctly sclerotized apical portion, basal portion short.

Female. Posterior margin of sternite VIII (Fig. 10D) broadly convex; genital segments with weakly asymmetric, large and sclerotized structure (Fig. 10E).

The fine, dense and uniform punctation of the elytra, and the similar shape and chaetotaxy of the male sternite VII and sternite VIII suggest that D. chenae is allied to D. reitteri. The species is distinguished from D. reitteri by the coarser punctation of the head and pronotum, the somewhat shorter elytra, the moderately modified dark setae of the male sternite VII, the differently shaped ventral process of the aedeagus and the more distinctly sclerotized structure in the female genital segments.

The type locality is situated in Anjiangping to the northwest of Guilin, northern Guangxi (Fig. 1). The specimens were sifted from leaf litter and grass in broad-leaved forests at an altitude of 1200 m.

(10 ♂♂, 12 ♀♀). Holotype: ♂, “China (Shaanxi) Qin Ling Shan, 110.06 E, 34.27 N, Hua Shan, 118 km E Xian, N valley, 1200–1400 m, leafy wd.sifted, 18./20.VIII.1995, Wrase / Sammlung M. Schülke Berlin / Holotypus ♂ Domene cultrata sp. n., det. B. Feldmann & Z. Peng 2014 ” (cSch). Paratypes: 2 ♂♂, 3 ♀♀ (4 specimens are teneral): same label data as holotype (cSch, cRou, cFel);; 1 ♂: “China [28] S-Shaanxi, 34 km S Hanzhong, 32°44'22"N, 106°51'55"E, 1460 m, 14.VIII.2012, V. Assing” (cAss); 1 ♂: “China [27a] S-Shaanxi, Micang Shan, 42 km S Hanzhong, 32°40'52"N, 106°49'16"E, 1090 m, 14.VIII.2012, V. Assing” (cFel); 2 ♀♀: “China (S. Shaanxi), Micang Shan, 42 km S Hanzhong, 32°40'43"N, 106°48'33"E, 1090 m, (stream valley, shady S. slope, sec. mixed for., raked from roots of perennials, soil, under stones) 17.VIII.2012, D.W. Wrase (32)” (cSch, cFel); 1 ♂, 1 ♀: “China: Shaanxi, Qin Ling Shan, 110.06 E, 34.27 N, Hua Shan Mt. N Valley, 1200–1400 m, 118 km E Xian, sifted, 18./20.VIII.1995, leg. M. Schülke” (cSch); 1 ♀: “China: border Shaanxi–Sichuan [today Chongqing], Daba Shan pass, 20 km SSE Zhenping, 1700–1800 m, 31°44'N, 109°35'E, 9.VII.2001, A. Smetana [C96b]” (cSme); 1 ♂, 1 ♀: “China: Shaanxi Prov., Zhouzhi County, Houzhenzi, Qinling, West Sangongli Gou, N33.50.613 E107.48.524 / 17–19.V.2008 alt. 1,336 m, Hao Huang & Xu Wang leg.” (SNUC); 1 ♂, 2 ♀♀: “China [17] S-Gansu, S Longnan, Min Shan, macchia, 33°05'24"N, 104°45'13"E, 1500 m, 6.VIII.2012, V. Assing”(cAss); 2 ♂♂, 2 ♀♀: “China (W-Hubei) Daba Shan, creek vall. 8 km NW Muyuping, 31°29'N, 110°22'E, 1540 m, (edge of small creek), 18.VII.2001, Wrase (16)” (cSch, cFel).

The specific epithet is an adjective derived from the Latin noun culter (knife) and alludes to the shape of the ventral process of the aedeagus.

Measurements (in mm) and ratios: BL 8.90–10.2, FL 5.38–5.50, HL 1.31–1.50, HW 1.22–1.39, AnL 3.22–3.62, NW 0.46–0.50, PL 1.48–1.62, PW 1.17–1.40, EL 1.46–1.63, EW 1.50–1.78, TiL 1.65–1.92, TaL 1.18–1.42, AW 1.37–1.53, AL 1.33–1.48, HL/HW 1.04–1.14, HW/PW 0.99–1.07, HL/PL 0.88–0.96, NW/HW 0.36–0.38, PL/PW 1.19–1.26, EL/PL 0.99–1.01.

Habitus as in Fig. 11A. Body dark brown; legs brownish yellow, with brown profemora and protibiae; antennae brown to light brown.

Figure 11. 

Habitus of Domene spp., A D. cultrata B D. cuspidata C D. reducta. Scales: 1.0 mm.

Head orbicular, widest behind eyes; punctation (Fig. 12A) coarse, umbilicate and dense, interstices forming very narrow ridges. All antennomeres longer than broad; antennomeres IV–X of equal length; antennomeres I 1.6 times, II 0.9 times, III 1.3 times, XI 1.4 times as long as IV. Maxillary palpus very slender, preapical joint 2.8–3.0 times as long as broad.

Figure 12. 

Domene cultrata. A median dorsal portion of head B median portion of pronotum C female tergite VIII D female sternite VIII E female tergites IX–X. F male sternite VII G male sternite VIII H aedeagus in ventral view I aedeagus in lateral view J aedeagus in dorsal view. Scales: A–B 0.2 mm; C–J 0.5 mm.

Pronotum about as wide as head, widest in the middle; lateral margins convex in dorsal view; punctation (Fig. 12B) similar to that of head; midline with rudiment of a fine glossy line.

Elytra without distinct longitudinal ridges; suture elevated in posterior two thirds; macropunctation coarse, irregular, partly confluent, and partly somewhat seriate; interstices rugose, rendering elytra matt, with irregular and mostly barely visible micropunctation (visible in posterior part of elytra). Hind wings fully developed. Protarsomeres I–IV distinctly dilated.

Abdomen with fine and dense punctation on tergites III–VIII; posterior margin of tergite VIII broadly convex (Fig. 12C); interstices with distinct microreticulation; posterior margin of tergite VII with palisade fringe.

Male. Sternites III–VI unmodified; sternite VII (Fig. 12F) distinctly transverse, with shallow median impression posteriorly, this impression with sparse, strongly modified, short and stout black setae, posterior margin broadly concave; sternite VIII (Fig. 12G) with shallow median impression, this impression with distinctly modified stout black setae, posterior excision moderately deep and V-shaped, on either side of the posterior excision with dense cluster of dark setae; aedeagus as in Figs 12H–J, ventral process nearly straight and apically acute; dorsal fig with long, large and distinctly sclerotized apical portion, basal portion short and lamellate; internal sac with small sclerotized spines and with distinct membranous structures.

Female. Sternite VIII (Fig. 12D) distinctly oblong, posterior margin strongly convex; genital segments with asymmetric, slender and moderately sclerotized structure (Fig. 12E).

Domene cultrata is subject to rather pronounced intraspecific variation of size, body proportions and coloration of the legs.

Based on the similar chaetotaxy and shape of the male sternite VIII, and the shape of the ventral process of the aedeagus, D. cultrata belongs to the D. malaisei species group and is allied to D. cuspidata. It can be distinguished from other species of the group by the distinctly coarser macropunctation of the elytra, the differently shaped ventral process of the aedeagus, and the slender sclerotized structure in the female genital segments, from D. malaisei and D. reducta also by the shallower impression and the less deep posterior excision of the male sternite VIII.

This species has been recorded from the Qinling Shan and Daba Shan, as well as from adjacent mountain ranges (Fig. 1). The specimens were sifted from leaf litter in forests or raked from roots of perennials and soil, or found under stones at altitudes of 1090–1800 m. Four specimens found in August are teneral.

(10 ♂♂, 24 ♀♀). Holotype: ♂: “China: Shaanxi Prov., Hanzhong City, Nanzheng County, Yuanba Town, Liping National Forest Park / 32°50'N, 106°36'E, 15.VII.2012 1,400–1,600 m, Chen, Li, Ma, & Zhao leg. / Holotypus ♂ Domene cuspidata sp. n., det. B. Feldmann & Z. Peng 2014” (SNUC). Paratypes: 2 ♂♂, 11 ♀♀: same data as holotype (SNUC, cAss); 1 ♀: same data, but “16.VII.2012, Yu-Hong Pan leg.” (SNUC); 1 ♀: same data, but “16.VII.2012, Li-Zhen Li leg.” (SNUC); 3 ♀♀: same data, but “16.VII.2012” (SNUC); 1 ♀: “China, Shaanxi, Qinling Shan above Houzhenzi, 115 km WSW Xi’an, 1450 m, 33°50'N, 107°47'E, 5.VII.2001, A. Smetana [C95b]” (cSme); 1 ♂: “China [3] S-Gansu, N Chengxian, W-Qinling Shan, 34°08'24"N, 105°46'43"E, 1750 m, 28.VII.2012, V. Assing” (cAss); 1 ♂: “China: S-Gansu [CH 12-03], W Qinling Shan, 43 km N Chengxian, 34°08'24"N, 105°46'43"E, 1750 m, moist valley with creek and ponds, meadow with Artemisia, 28.VII.2012, leg M. Schülke” (cSch); 1 ♀: “China: S-Gansu [CH 12-05], W Qinling Shan, 47 km N Chengxian, 34°10'17"N, 105°42'56"E, 1850 m, mixed secondary forest margin, litter sifted, 29.VII.2012, leg M. Schülke” (cSch); 1 ♂, 1 ♀: “China, S-Gansu [CH 12-05], W. Qinling Shan, 47 km N Chengxian, 34°10'20"N, 105°42'19"E, 1830 m, (creek valley, loam deposit on meadow with tall herbaceous vegetation,raked/dug, 29.VII.2012, D. W. Wrase” (cSch, cFel); 1 ♂, 3 ♀♀: “China: W-Sichuan, Ya’an Pref., Shimian Co., Daxue Shan, road betw. Anshunchang–Wanba, 12 km W Shimian, 1300 m, 9.VII.1999, leg. A. Pütz” (cPüt, cFel).

The specific epithet is an adjective derived from the Latin noun cuspis (cusp) and refers to the apically acute ventral process of the aedeagus.

Measurements (in mm) and ratios: BL 8.89–9.56, FL 5.12–5.34, HL 1.26–1.39, HW 1.20–1.30, AnL 3.17–3.61, NW 0.45–0.50, PL 1.48–1.57, PW 1.20–1.28, EL 1.42–1.50, EW 1.48–1.62, TiL 1.72–1.79, TaL 1.20–1.33, AW 1.35–1.49, AL 1.32–1.65, HL/HW 1.04–1.08, HW/PW 1.00–1.03, HL/PL 0.85–0.89, NW/HW 0.38–0.39, PL/PW 1.21–1.24, EL/PL 0.94–0.97.

External characters (Fig. 11B) as in D. cultrata, distinguished only by the distinctly less coarse macropunctation and less rugose interstices of the elytra rendering the elytra more shiny in D. cuspidata, and by the primary and secondary sexual characters:

Male. Sternites III–VI unmodified; sternite VII (Fig. 13F) distinctly transverse, with shallow postero-median impression, this impression with sparse strongly modified, short and stout black setae, posterior margin concave in the middle; sternite VIII (Fig. 13G) with extensive median impression, this impression with distinctly modified stout black setae, posterior excision less deep, V-shaped, on either side of the posterior excision with a dense cluster of dark setae; aedeagus as in Figs 13H–J, ventral process distinctly sclerotized, with slender and very acute apical portion; dorsal fig with long, large and distinctly sclerotized apical portion, basal portion short; internal sac with membranous structures.

Figure 13. 

Domene cuspidata. A median dorsal portion of head B median portion of pronotum C female tergite VIII D female sternite VIII E female tergites IX–X. F male sternite VII G male sternite VIII H aedeagus in ventral view I aedeagus in lateral view J aedeagus in dorsal view. Scales: A–B 0.2 mm; C–J 0.5 mm.

Female. Sternite VIII (Fig. 13D) oblong, posterior margin broadly convex; genital segments with asymmetric, large and moderately sclerotized structure (Fig. 13E).

Based particularly on the similar chaetotaxy and shape of the male sternite VIII and the shape of the ventral process of the aedeagus, D. cuspidata belongs to the D. malaisei species group and is closely allied to D. cultrata. It is distinguished from other species of the group by the apically more acute ventral process of the aedeagus and by the large, moderately sclerotized structure in the female genital segments, from D. malaisei and D. reducta also by the shallower impression and less deep posterior excision of the male sternite VIII.

The species was recorded from the Qinling Shan and Dalou Shan (Fig. 1). The specimens were sifted from forest leaf litter and a loamy meadow with tall herbaceous vegetation at altitudes of 1300–1850 m. Six paratypes found in July are teneral.

(5 ♂♂, 9 ♀♀). Holotype ♂: “China: Sichuan Prov., Tianquan County, Labahe N. R., 30°09'N, 102°27'E, 29.VII.2006 1,900 m, Hu & Tang leg. / Holotypus ♂ Domene reducta sp. n., det. B. Feldmann & Z. Peng 2014” (SNUC). Paratypes: 1 ♂, 5 ♀♀ [all teneral], same label data as holotype (SNUC); 2 ♂♂, 1♀ [1 ♂, 1 ♀ teneral]: same data, but “Liangluxiang, 29°56'N, 102°23'E, alt. 1,500–1,700 m / 10.VII.2012, Dai, Peng & Yin leg.” (SNUC); 1 ♀ [teneral]: same data, but “Liangluxiang, 29°56'N, 102°23'E, alt. 1,900–2,000 m, 10.VII.2012, Dai, Peng & Yin leg leg.” (SNUC); 1 ♂: “China, Sichuan: Quing-cheng-Shan [ca. 30°53'N, 103°35'E], 1400–1700 m, 22.VI.1996, D. Erber” (cFel); 1 ♀: “China, W.Sichuan, (Ya’an Pref. Tianquan Co.), Jiajin Shan, valley above Labahe, N.R.ST., 57 km W Ya’an, 30°06'N, 102°25'E (light forest), 1800 m, 12.VII.1999, D.W. Wrase” (cFel); 1 ♀: “China: W-Sichuan, Ya’an Prefecture, Tianquan Co., Jiajin Shan, Tal oberh. Labahe, N.R.St., 57 km W. Ya’an, 30°06'N 102°25'E, Streu, Rinde, Pilze, 1800 m, 12.VII.1999, leg. M. Schülke” (cSch).

The specific epithet (Latin, adjective: reduced) alludes to the minute sclerotized structure in the female genital segments.

Measurements (in mm) and ratios: BL 8.95–10.84, FL 5.37–5.48, HL 1.42–1.48, HW 1.35–1.41, AnL 3.36–3.56, NW 0.50–0.55, PL 1.57–1.66, PW 1.28–1.35, EL 1.41–1.48, EW 1.63–1.70, TiL 1.81–1.87, TaL 1.28–1.31, AW 1.51–1.57, AL 1.52–1.54, HL/HW 1.04–1.06, HW/PW 1.03–1.05, HL/PL 0.89–0.91, NW/HW 0.37–0.39, PL/PW 1.22–1.24, EL/PL 0.88–0.91.

Habitus as in Fig. 11C. Body dark brown; legs light brown with darker profemora and protibiae; antennae brown to light brown.

Head orbicular, widest behind eyes; punctation (Fig. 14A) moderately coarse, distinctly umbilicate, and very dense, interstices forming very narrow ridges. All antennomeres longer than wide; antennomeres IV–X of equal length; antennomeres I 1.7 times, II 1.1 times, III 1.4 times, XI 1.2 times as long as IV. Maxillary palpus slender, preapical joint 3.2–3.5 times as long as broad.

Figure 14. 

Domene reducta. A median dorsal portion of head B median portion of pronotum C female tergite VIII D female sternite VIII E female tergites IX–X. F male sternite VII G male sternite VIII H aedeagus in ventral view I aedeagus in lateral view J aedeagus in dorsal view. Scales: A–B 0.2 mm; C–J 0.5 mm.

Pronotum slightly narrower than head, widest in the middle; lateral margins convex in dorsal view; punctation (Fig. 14B) somewhat coarser than that of head; midline with rudiment of a fine glossy line.

Elytra without distinct longitudinal ridges; suture elevated in posterior three-fourths; macropunctation moderately coarse, irregular, partly confluent, and partly somewhat seriate; interstices with irregular micropunctation. Hind wings fully developed. Protarsomeres I–IV distinctly dilated.

Abdomen with fine and dense punctation on tergites III–VIII; posterior margin of tergite VIII broadly and weakly convex (Fig. 14C); interstices with shallow microreticulation; posterior margin of tergite VII with palisade fringe.

Male. Sternites III–VI unmodified; sternite VII (Fig. 14F) distinctly transverse, with median impression of triangular shape posteriorly, this impression with strongly modified, short and stout black setae, posterior margin weakly concave in the middle; sternite VIII (Fig. 14G) with shallow and extensive median impression, this impression with stout black setae, posterior excision moderately deep and V-shaped, on either side of the posterior excision with a dense cluster of dark setae; aedeagus as in Figs 14H–J, ventral process long, slender, evenly curved and apically acute; dorsal fig with long and distinctly sclerotized apical portion, basal portion long and lamellate.

Female. Posterior margin of sternite VIII (Fig. 14D) broadly convex; genital segments with a small symmetric, weakly sclerotized structure (Fig. 14E).

Based particularly on the similar chaetotaxy and shape of the male sternite VIII, and the shape of the ventral process of the aedeagus, D. reducta belongs to the D. malaisei species group and is closely related to D. malaisei. Domene reducta is distinguished from other species of the group by on average larger body size (especially from D. malaisei), the shape of the impression on the male sternite VIII, the long, slender, evenly curved ventral process of the aedeagus and by the symmetric, small and moderately sclerotized structure in the female genital segments.

The species is known from the Qingcheng Shan and Hengduan Shan, central Sichuan (Fig. 1). The specimens were sifted from leaf litter and soil in evergreen broad-leaved forests at altitudes of 1400–1900 m. Nine paratypes found in July are teneral.

5♀♀, Sichuan, Emei Shan, 29°34'N, 103°21'E, 1800–2400 m, sifted, 27.VI.–5.VII.2009, leg. Grebennikov (cSme, cAss).

The above brachypterous females undoubtedly represent an undescribed species distinguished from the other species known from China by the conspicuously large head and the distinctly impressed sutural portion of the elytra, from most species also by the short and narrow elytra and by the absence of a palisade fringe at the posterior margin of the male tergite VII.