Odynerus elegans Wesmael, 1833, designated by Richards (1935).

Holotype. ♀, China, Yunnan Prov., Dali City, Yunlong County, Tianchi National Nature Reserve, 25°52'13.05"N, 99°17'14.33"E, 2579 m, 9.VII.2011, Tingjing Li, No. 1001516 (CQNU); Paratypes: 1♀1♂, the same as holotype, Nos. 1001524, 1001517 (CQNU); 1♀, China, Yunnan Prov., Nujiang City, Lanping County, Jinding Town, Xinshengqiao National Forest Park, 26°26'56.36"N, 99°23'04.69"E, 2412 m, 12. VII.2011, Tingjing Li, No. 1004036 (CQNU).

Female (Fig. 1): body length 9.5–10.5 mm, forewing length 10.0–10.5 mm. Black; with the following parts orange-red to red: dorsal pronotal spot, apical border of T1, and subapical border of T2, S2 and T4 (absent in one female paratype); interantennal spot and post-ocular dot orange-yellow; fore tibia inside brown. Wings lightly infuscate. Hairs pale brown; mesosoma with sparse lengthened hairs, in addition to short pubescence.

Head. Clypeus sparsely punctate-puncticulate to foveolate-puncticulate, apex broadly and moderately emarginated, evenly convex, with apical teeth, and slightly reflexed anteriorly (Fig. 3); frons and vertex foveolate-puncticulate, major punctures closely spaced; interantennal carina prominent; cephalic fovea somewhat larger than post ocellus, and foveal interval somewhat less than postocellar distance, cephalic foveal carina indistinct (Fig. 5); occipital carina without submedial incisions.

Mesosoma. Pronotum, except anterior face, foveolate-puncticulate, major punctures closely spaced, more or less costate laterally, anterior face smooth and polish, pronotal carina complete, humeral angle barely projected. Mesoscutum foveolate-puncticulate, major punctures widely spaced medially, moderately spaced anteriorly and posteriorly; notaulus complete; mesepisternum with epicnemial carina dorsally obsolete and ventrally faint (Fig. 6); anterior margin of pseudosternum without high reflexed margin; mesoscutellum foveolate-puncticulate, similar to those on mesoscutum. Dorsal mesepisternum sparsely foveolate-puncticulate, space between punctures smooth and polish, ventral mesepisternum sparsely foveolate-puncticulate, major punctures widely spaced, minor punctures obscure, space between punctures alutaceous; dorsal mesepimeron sparsely foveolate, space between punctures smooth and polish, ventral mesepimeron dull, and coarsely alutaceous. Metanotum foveolate-puncticulate dorsally, obscurely puncticulate ventrally, metanotum nearly vertical, dorsal surface narrow. Propodeal dorsum and posterior face coarse, areolate-rugose, propodeal superior shelf moderately long (2.2 times trans-scutal sulcus), lateral face of propodeum striately rugose ventrally, areolate-rugose dorsally, propodeal submedian carina present ventrally, faint and irregular dorsally (Fig. 10), propodeal valvula short posteriorly, fused distally to submarginal carina, propodeal orifice somewhat elliptic dorsally (Fig. 10).

Metasoma. Metasomal tergum 1 with postcarinal area foveolate-puncticulate to foveate-puncticulate (Figs 7–8), postcarinal length of T1/apical width=0.73, carinal width/apical width=0.79, postcarinal sides slightly convergent toward base, transverse carina laterally faint (Fig. 7), longitudinal furrow narrowly and shallowly depressed, with deeper narrow medial sulcus (Fig. 8), apical margin indistinctly depressed; S1 flat and without basal carina anteriorly, areolate-rugose medially and posteriorly, without median longitudinal ridge, and lateral oblique ridges strongly prominent (Fig. 9); segment 2 foveolate-puncticulate basally, major punctures more closely spaced; T3–5 sparsely punctate-puncticulate to foveolate-puncticulate, major punctures subapically; segment 2 except base and S3–6 with evenly minor punctures.

Male. Body length 7.5 mm, forewing length 8.5 mm. Sculpture, punctuation, hairs, and coloration similar to female except as follows: clypeus entirely light yellow to yellow (Fig. 4), except apical margin brown red; antennal segments 3–13 pale brown ventrally (Fig. 2); dorsal pronotal spot, apical border of T1, and subapical border of both T2 and S2 orange-red; mesepisternum dorsally with small yellow spot; fore tibia inside pale brown, fore tarsus brown apically; antennal tyloids absent, segment 13 length/width=1.56 (Fig. 2).

China (Yunnan).

This species is similar to Symmorphus (Symmorphus) sichuanensis by S1 without basal carina, T1 with transverse carina laterally faint to obsolete, and mesepisternum with epicnemial carina dorsally obsolete; but can be easily distinguished from the similar species and other members of the genus by the combination of the following characters: body moderately long; in both female and male, dorsal pronotal spot orange-red (Figs 1, 5); in female, mesepisternum with epicnemial carina ventrally faint; in male, antenna without tyloids (Fig. 2), and subapical border of both T2 and S2 orange-red.

It is named after the type locality of the species, Tianchi National Nature Reserve in Dali city, Yunnan of China.

Holotype. ♀, China, Yunnan Prov., Xishuangbanna, Jinghong City, Jinuo mountain, 22°02'17.81"N, 101°00'15.36"E, 901 m, 12.IV, 2010, Rui Zhang, No. 1004037 (YNAU).

Female (Fig. 11): body length 6.5 mm, forewing length 7.0 mm. Black; with the following parts pale brown: basal transverse band of clypeus, post-ocular dot, medially uninterrupted dorsal pronotal band, large dorsal mesepisternal spot, tegula, large mesoscutellar spot apically, apex of T1, apical margin of both T2 and S2, apex of fore femur, and fore tibia largely; fore and mid tarsi dark brown. Wings lightly infuscate. Hairs pale brown, mesosoma without sparse lengthened hairs, in addition to short pubescence.

Head. Clypeus moderately punctate-puncticulate to foveolate-puncticulate, punctures sparse medioapically, space between punctures smooth and polish; clypeal apex truncated, clypeus evenly convex, without apical teeth (Fig. 12). Frons and vertex foveolate-puncticulate, major punctures closely spaced; interantennal carina prominent; cephalic fovea somewhat smaller than post ocellus, foveal interval subequal to postocellar distance, without cephalic foveal carina; occipital carina with 2 submedial incisions.

Mesosoma. Pronotum, except anterior face, foveolate-puncticulate, with major punctures closely spaced, more or less costate laterally; anterior face distinctly imbricate; pronotal carina dorsally obsolete; humeral angle slightly projected. Mesoscutum foveolate-puncticulate, major punctures closely spaced anteriorly and posteriorly, widely spaced laterally; notaulus complete; epicnemial carina complete; pseudosternum anterior margin without high reflexed margin. Mesoscutellum foveolate-puncticulate, major punctures closely spaced, with shallowly medial furrow. Mesepisternum with complete epicnemial carina (Fig. 15), dorsally punctate-puncticulate and minor punctures distinct basally, other parts foveolate-puncticulate, major punctures widely spaced, minor punctures obscure. Mesepimeron dull and densely striate. Metanotum primarily oblique and not vertical, dorsally dull, coarse and areolate-rugose. Propodeum dull, densely striate laterally, areolate-rugose dorsally, posterior face deeply heart-shaped hollowed, margin reflexed, complete and sharply defined throughout (Fig. 16), propodeal superior shelf short, 2 times trans-scutal sulcus; propodeal submedian carina entirely absent, propodeal valvula short posteriorly, fused distally to posterior margin; propodeal orifice small and indistinct.

Metasoma. Metasomal tergum 1 with postcarinal area foveolate-puncticulate, major punctures densely spaced, postcarinal length short, postcarinal length/apical width=0.68, carinal width/apical width=0.85, postcarinal sides barely convergent toward base, transverse carina complete, longitudinal furrow narrowly and shallowly depressed, with deeper narrow medial sulcus, apical margin indistinctly depressed (Fig. 14); S1 rugose anteriorly, basal carina inflated and raised posteriorly, fused to lateral oblique ridges, with median longitudinal ridge, lateral oblique ridges slightly prominent, median longitudinal ridge flanked by longitudinal carinate rugae; segment 2 foveolate-puncticulate, major punctures widely spaced from base to apex, minor punctures connected by obscure imbricate subsculpture; T3–T5 and S3 with densely foveolate toward apex; S3–S6 with imbricate subsculpture.

Male. Unknown.

This species is easily distinguished at once from all other species of Symmorphus by the combination of the following characters: propodeal posterior face deeply hollowed (Fig. 16), and occipital carina with 2 submedial incisions, in other species of the genus propodeal posterior face not hollowed.

China (Yunnan).

The specific name is the Latin cavatus (= hollow), which refers to propodeal posterior face of the species deeply hollowed.

Holotype. ♂, China, Tibet, Nyingchi, Medog County, 29°71'N, 95°63'E, 3026 m, 13.VII.2013, Yong Zhou, No. 1004038 (CQNU).

Male (Fig. 17): body length 7.0 mm, forewing length 9.0 mm. Black; apical margin of T1 orange-red, subapical margin of both T2 and S2 orange-yellow. Wings lightly infuscate. Hairs white, mesosoma with sparse lengthened hairs, in addition to short pubescence.

Head. Clypeus moderately punctate-puncticulate, clypeal apex deeply emarginated and with acute apical toothed laterally (Fig. 18). Frons punctate-puncticulate to foveolate-puncticulate, major punctures closely spaced; vertex punctate-puncticulate and barely foveolate-puncticulate, minor punctures closely spaced; interantennal carina prominent; male antennal segments 10–13 with tyloids, segment 13 length/width=1.43 (Fig. 19).

Mesosoma. Pronotum, except anterior face, punctate-puncticulate to foveolate-puncticulate, major punctures widely spaced, more or less costate laterally, minor punctures evenly and closely spaced; anterior face obscurely imbricate; pronotal carina complete; humeral angle moderately projected. Mesoscutum punctate-puncticulate to foveolate-puncticulate, primarily punctate-puncticulate, minor punctures evenly and densely spaced, major punctures great widely spaced; notaulus complete and distinctly deep (Fig. 20); pseudosternum anterior margin without high reflexed margin. Mesoscutellum similar to mesoscutum. Dorsal mesepisternum sparsely punctate-puncticulate to foveolate-puncticulate, major punctures very widely spaced, shallow and indistinct, minor punctures obscure; ventral mesepisternum foveolate-puncticulate, major punctures widely spaced, minor punctures obscure. Mesepimeron striately rugose. Metanotum nearly vertical, dorsal surface narrow, foveolate-puncticulate dorsally, striately rugose ventrally. Propodeum striately rugose laterally, areolate-rugose dorsally; posterior face obscure striately rugose, propodeal superior shelf length 3.1 timestrans-scutal sulcus, propodeal orifice broadly and rounded, propodeal submedian carina complete and sharply defined throughout; propodeal valvula short posteriorly, fused distally to submarginal carina.

Metasoma. Metasomal tergum 1 with postcarinal area foveolate-puncticulate, postcarinal length/apical width=0.71, carinal width/apical width=0.85, postcarinal sides slightly convergent toward base, transverse carina complete, longitudinal furrow broadly depressed, with deeper narrow medial sulcus; apical margin indistinctly depressed (Fig. 21); S1 rugose anteriorly, basal carina deeply curved posteriorly, fused to lateral oblique ridges, median longitudinal ridge strongly prominent and flanked by longitudinal carinate rugae; segment 2 punctate-puncticulate, minor punctures connected by obscure imbricate subsculpture; T3–5 sparsely foveolate-puncticulate, major punctures subapically. Segments 3–6 with imbricate subsculpture.

Female. Unknown.

This species is easily distinguished from all other species of Symmorphus by the combination of the following characters: body black, except apical margin of T1, and subapical margins of both T2 and S2 (Fig. 17); clypeal apex deeply emarginated and with acute apical toothed laterally (Fig. 18).

China (Tibet).

The specific name nigriclypeus is the Latin nigr (= black) + clypeus (=clypeus), which refers to the clypeus in male of the species black.

1♀, China, Yunnan Prov., Dali, North Gucheng, 9.V.2007, Rui Zhang; 1♀, China, Yunnan Prov., Dali, Yunlong, Luodun Town, 10.VII.2011, Tingjing Li; 1♀, China, Yunnan Prov., Nujiang, Lanping, Yingpan Town, 13.VII.2011, Zhenhu Wu; 1♀, China, Yunnan Prov., Lijiang, Ninglang, Daxing Town, 25.VII.2011, Tingjing Li; 6♂♂, China, Yunnan Prov., Dehong, Yingjiang, Tongbiguan Natural reserve, 3.V.2013; 1♀, China, Chongqing, Wansheng, Heishangu, 4.V.2011, Zhenhu Wu.

China (Sichuan, Yunnan, Chongqing); Nepal; Korea.

1♀, China, Jilin Prov., Tonghua, Mehekou, 22.VIII.1993, Zhihong Li.

China (Jilin); Norway; Sweden; Finland; Denmark; France; Germany; Austria; Greece; Turkey; Belarus; Russia (to Primorskij Krai); Kazahkstan; Mongolia; the Korean Peninsula; Japan.

1♀, China, Beijing, Haidian, 29.IV.1952. Jikun Yang; 1♀, China, Beijing, Gongzhufen, 12.V.1952. Jikun Yang; 1♂, China, Shaanxi Prov., Baoji, Fengxian, Jialing River, 26.V.2007, Xiaoyu, Jiang.

China (Beijing, Shaanxi, Fujian, Guangdong, Jiangsu, Sichuan); Russia; South Korea; Japan.

No specimens examined.

China (Jiangsu).

1♂, China, Chongqing, Jiangjin, Simianshan National Nature Reserve, 6.V.2012, Ju You.

China (Jiangsu, Chongqing); Europe, eastward to northeastern Siberia and Far Eastern Russia; Kyrgyzstan; Kazahkstan; Korea; Japan.

6♀♀2♂♂, China, Sichuan Prov., Panzhihua, Renhe Town, 28.VII.2011, Tingjing Li.

China (Sichuan); Russia; the Korean Peninsula; Japan.

2♀♀, China, Liaoning Prov., Liaoyang, Gongchangling, Anping, 7.VII.2012.Ju You; 1♀, China, Jilin Prov., Changchun, Dehui, Xiajiadian, 28.VI.2012, Ju You; 1♀, China, Jilin Prov., Baishan, Linjiang, Naozhi Town, 7.VII.2012, Ju You.

China (new record: Liaoning, Jilin); Norway; Sweden; Finland; Netherlands; Germany; Austria; Belarus; Mongolia; Russia.

No specimens examined.

China (Taiwan).

2♀♀, China, Inner Mongolia, Helan Mountain, Gulamuxiaosong Hill, 30.VII.2010, Jian Li & Junzhe Xue; 1♀, China, Inner Mongolia, Helan Mountain, Halawu Ravine, 20.VII.2006, Ming Luo; 1♀, China, Inner Mongolia, Helan Mountain, Yushuwan, 27.VII.2010, Fangzhou Ma.

China (new record: Inner Mongolia), Russia: southern Siberia to Sakhalin; Korea; Japan.

2♀♀4♂♂, China, Yunnan Prov., Diqing, Deqin, Near the county, 19.VII.2011, Tingjing Li; 4♀♀, China, Yunnan Prov., Dali, Yunlong, Tianchi, 9.VII.2011, Tingjing Li.

China (Yunnan, Sichuan, Taiwan); Russia: Irkutsk, Primorskij Krai; Kazahkstan; North Korea; Japan.

No specimens examined.

China (Taiwan).

3♀♀: China, Sichuan Prov., Leshan City, Emeishan, Gaoqiao Town, Yanshi Village, 11.VIII.2011, Tingjing Li.

China (Sichuan); Thailand.

5♂♂, China, Ningxia, Jingyuan, Xixia Forest, 15–16.VII.2008, Xiumin Li; 1♂, China, Ningxia, Longde, Sutai Forest, 1–2, VII.2008, Xinpu Wang; 1♂, China, Ningxia, Guyuan, Lvyan Forest, 9–10.VII.2008, Guodong Ren.

China (new record: Ningxia); Kyrgyzstan; Kazahkstan.

10♀♀, China, Yunnan Prov., Dali, Yunlong, Tianchi, 9.VII.2011, Tingjing Li; 9♀♀, China, Yunnan Prov., Nujiang, Lanping, Yingpan Town, 12.VII.2011, Zhenhu Wu; 1♀, China, Sichuan Prov., Kangding, Paoma Mountain, 7.VII.2005, Hu Zhou.

China (new record: Yunnan, Sichuan); India; Nepal.

No specimens examined.

China (Shaanxi).

5♀♀11♂♂, China, Yunnan Prov., Diqing, Deqin, Near the county, 19.VII.2011, Tingjing Li; 1♂, China, Tibet, Changdu, Mangkang, 3508 m, 5.VII.2013, Yong Zhou.

China (Yunnan, Tibet, Fujian).