(C) 2012 Stjepan Krčmar. This is an open access article distributed under the terms of the Creative Commons Attribution License 3.0 (CC-BY), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
For reference, use of the paginated PDF or printed version of this article is recommended.
The present paper is based on original and literature data. In Croatia the first studies on the occurrence of ixodid species were made about 80 years ago. The number of tick species recorded in Croatia considerably increased during the 1950s, 60s, 70s and 80s of the past century. A total of 21 species of hard tick belonging to 5 genera have been recorded in Croatia. Ixodes is the best represented genus, with seven species recorded. Haemaphysalis is represented by six species, followed by Rhipicephalus with four species. Dermacentor and Hyalomma are represented by two species each. The ticks were collected on 47 different host species. Eleven tick species were collected on Bos taurus and Ovis aries, followed by Capra hircus and Equus caballus with 8 species and Canis lupus familiaris with 6 species. On the remaining 42 host species one, two or three tick species were collected. The most widespread tick is Ixodes ricinus which was found on 25 different host species.
Ixodidae, hard ticks, Croatia, Europe
Hard ticks (Acari: Ixodidae) are obligate hematophaguos ectoparasites and important vectors of viruses, bacteria and protozoa (
The following lists of species include all available literary records and new, still unpublished data. The samplings of ticks in Slavonia and Baranja were carried out during 2011 from the beginning of March to mid-September. Ticks were sampled by the flagging method, and were picked by hand from domestic animals and humans. The flagging method was used for tick sampling in two different vegetation types around the village of Mikleuš (YL 15). A 1 m2 white flag was dragged over the vegetation for about 50 m at each locality. The first locality was in the mixed forests of durmast oak and hornbeam (as. Epimedio-Carpinetum betuli /Horvat 1938/ Borhidi 1963), whereas the second locality was in open biotopes on grassland. In the veterinary clinic in Đakovo (BR 92), ticks were sampled from dogs during vaccination or during normal veterinary examinations, while sampling from cats was performed in the village of Zmajevac (CR 37). Also, some ticks were sampled from humans in the village of Zmajevac. Identification was carried out using standard keys for European ticks (
Phylum: Chordata
Subphylum: Vertebrata
Class: Mammalia
Order: Artiodactyla
Family: Bovidae
Genus: Bos Linnaeus, 1758
Bos taurus Linnaeus, 1758
Tick species: Dermacentor marginatus (Sulzer, 1776), Dermacentor reticulatus (Fabricius, 1794), Haemaphysalis punctata Canestrini and Fanzago, 1878, Haemaphysalis inermis Birula, 1895, Hyalomma scupense Schulze, 1919, Hyalomma marginatum Koch, 1844, Rhipicephalus annulatus (Say, 1821), Rhipicephalus bursa Canestrini and Fanzago, 1878, Rhipicephalus sanguineus (Latreille, 1806), Ixodes hexagonus Leach, 1815, Ixodes ricinus (Linnaeus, 1758)
Genus: Capra Linnaeus, 1758
Capra hircus (Linnaeus, 1758)
Tick species: Dermacentor marginatus, Haemaphysalis punctata, Haemaphysalis sulcata Canestrini and Fanzago, 1878, Hyalomma marginatum, Rhipicephalus bursa, Rhipicephalus sanguineus, Rhipicephalus turanicus Pomerantsev, 1940, Ixodes ricinus
Genus: Ovis Linnaeus, 1758
Ovis aries Linnaeus, 1758
Tick species: Dermacentor marginatus, Haemaphysalis punctata, Haemaphysalis inermis, Haemaphysalis sulcata, Hyalomma scupense, Hyalomma marginatum, Rhipicephalus bursa, Rhipicephalus sanguineus, Rhipicephalus turanicus, Ixodes gibbosus Nuttal, 1916, Ixodes ricinus
Family: Cervidae
Genus: Capreolus Gray, 1821
Capreolus capreolus (Linnaeus, 1758)
Tick species: Haemaphysalis concinna Koch, 1844, Rhipicephalus turanicus, Ixodes ricinus
Order: Carnivora
Family: Canidae
Genus: Canis Linnaeus, 1758
Canis aureus (Linnaeus, 1758)
Tick species: Rhipicephalus sanguineus
Canis lupus familiaris Linnaeus, 1758
Tick species: Dermacentor marginatus, Haemaphysalis inermis, Rhipicephalus sanguineus, Rhipicephalus turanicus, Ixodes hexagonus, Ixodes ricinus
Family: Mustelidae
Genus: Martes Pinel, 1792
Martes foina (Erxleben, 1777)
Tick species: Ixodes frontalis (Panzer, 1798)
Martes martes (Linnaeus, 1758)
Tick species: Ixodes hexagonus, Ixodes ricinus
Family: Felidae
Genus: Felis Linnaeus, 1758
Felis catus Linnaeus, 1758
Tick species: Dermacentor marginatus, Rhipicephalus turanicus, Ixodes ricinus
Order: Chiroptera
Family: Rhinolophidae
Genus: Rhinolophus Lacepede, 1799
Rhinolophus ferrumequinum (Schreiber, 1774)
Tick species: Ixodes vespertilionis Koch, 1844
Rhinolophus hipposideros (Bechstein, 1800)
Tick species: Ixodes vespertilionis
Family: Vespertilionidae
Genus: Miniopterus Bonaparte, 1837
Miniopterus schreibersii (Kuhl, 1817)
Tick species: Ixodes vespertilionis
Genus: Myotis Kaup, 1829
Myotis myotis (Borkhausen, 1797)
Tick species: Ixodes vespertilionis
Order: Insectivora
Family: Erinaceidae
Genus: Erinaceus Linnaeus, 1758
Erinaceus concolor Martin, 1838
Tick species: Ixodes hexagonus
Erinaceus europaeus Linnaeus, 1758
Tick species: Hyalomma marginatum
Family: Soricidae
Genus: Neomys Kaup, 1829
Neomys sp.
Tick species: Ixodes trianguliceps Birula, 1895
Genus: Sorex Linnaeus, 1758
Sorex alpinus Schinz, 1837
Tick species: Ixodes trianguliceps
Sorex araneus Linnaeus, 1758
Tick species: Ixodes trianguliceps
Order: Perissodactyla
Family: Equidae
Genus: Equus Linnaeus, 1758
Equus asinus Linnaeus, 1758
Tick species: Rhipicephalus sanguineus, Ixodes ricinus
Equus caballus Linnaeus, 1758
Tick species: Dermacentor marginatus, Dermacentor reticulatus, Haemaphysalis punctata, Hyalomma marginatum, Rhipicephalus annulatus, Rhipicephalus bursa, Ixodes hexagonus, Ixodes ricinus
Order: Rodentia
Family: Muridae
Genus: Apodemus Kaup, 1829
Apodemus agrarius (Pallas, 1771)
Tick species: Dermacentor marginatus, Ixodes ricinus, Ixodes trianguliceps
Apodemus flavicollis (Melchior, 1834)
Tick species: Ixodes ricinus, Ixodes trianguliceps
Apodemus mystacinus (Danford et Alston, 1877)
Tick species: Ixodes ricinus, Ixodes trianguliceps
Apodemus sylvaticus (Linnaeus, 1758)
Tick species: Dermacentor marginatus, Ixodes ricinus, Ixodes trianguliceps
Genus: Clethrionomys Tilesius, 1850
Clethrionomys glareolus (Schreber, 1780)
Tick species: Dermacentor marginatus, Ixodes ricinus, Ixodes trianguliceps
Genus: Microtus, Schrank, 1798
Microtus arvalis (Pallas, 1779)
Tick species: Dermacentor marginatus
Order: Primates
Family: Hominidae
Genus: Homo Linnaeus, 1758
Homo sapiens Linnaeus, 1758
Tick species: Ixodes ricinus
Class: Aves
Order: Apodiformes
Family: Apodidae
Genus: Tachymarptis (Linnaeus, 1758)
Tachymarptis melba (Linnaeus, 1758)
Tick species: Haemaphysalis erinacei Pavesi, 1884
Order: Falconiformes
Family: Accipitridae
Genus: Buteo Lacepede, 1799
Buteo buteo (Linnaeus, 1758)
Tick species: Ixodes ricinus
Order: Passeriformes
Family: Corvidae
Genus: Pica Brisson, 1760
Pica pica (Linnaeus, 1758)
Tick species: Ixodes ricinus
Family: Bombycillidae
Genus: Bombycilla Vieillot, 1808
Bombycilla garrulus (Linnaeus, 1758)
Tick species: Ixodes frontalis
Family: Saxicolidae
Genus: Luscinia Forster, 1817
Luscinia megarhynchos C.L. Brehm 1831
Tick species: Ixodes ricinus
Genus: Phoenicurus Forster, 1817
Phoenicurus ochruros (S. G. Gmelin, 1774)
Tick species: Ixodes frontalis
Family: Turdidae
Genus: Turdus Linnaeus, 1758
Turdus merula Linnaeus, 1758
Tick species: Ixodes arboricola Schulze and Schlottke, 1930, Ixodes ricinus
Turdus philomelos C. L. Brehm, 1831
Tick species: Ixodes ricinus
Family: Motacillidae
Genus: Anthus Bechstein, 1805
Anthus trivialis (Linnaeus, 1758)
Tick species: Ixodes ricinus
Genus: Motacilla Linnaeus, 1758
Motacilla flava Linnaeus, 1758
Tick species: Haemaphysalis punctata
Family: Sylviidae
Genus: Cettia Bonaparte, 1834
Cettia cetti (Temminck, 1820)
Tick species: Ixodes ricinus
Class: Reptilia
Order: Squamata
Family: Lacertidae
Genus: Lacerta Linnaeus, 1758
Lacerta bilineata Daudin, 1802
Tick species: Haemaphysalis concinna
Lacerta trilineata Bedriaga, 1886
Tick species: Haemaphysalis sulcata
Lacerta viridis (Laurenti, 1768)
Tick species: Haemaphysalis sulcata
Genus: Dalmatolacerta Arnold, Arribas & Carranza, 2007
Dalmatolacerta oxycephala (Duméril & Bibron, 1839)
Tick species: Haemaphysalis sulcata
Genus: Podarcis Wagler, 1830
Podarcis melisellensis (Braun, 1877)
Tick species: Haemaphysalis concinna, Haemaphysalis sulcata, Ixodes ricinus
Podarcis muralis (Laurenti, 1768)
Tick species: Haemaphysalis concinna
Podarcis sicula (Rafinesque, 1810)
Tick species: Haemaphysalis sulcata, Ixodes ricinus
Genus: Algyroides (Bibron & Bory de Saint-Vincent, 1833)
Algyroides nigropunctatus (Duméril & Bibron, 1839)
Tick species: Haemaphysalis concinna, Haemaphysalis sulcata, Ixodes ricinus
Family: Colubridae
Genus: Elaphe Fitzinger, 1833
Elaphe longissima (Laurenti, 1768)
Tick species: Haemaphysalis concinna
Sampling sites are identified by a running number referring to the map in Figure 1 and UTM grid 10×10 km given in the third column of Table 1. The exact geographical coordinates of the sampling sites are given in the second column of Table 1 and were determined by (http://www.google.com/earth/index.html , http://www.worldplaces.net , http://www.plsavez.hr ).
Sampling sites of hard ticks (Ixodidae) in Croatia.
List of sampling sites
The tick fauna of Croatia is now known to include 21 species. These species are classified into five genera: Dermacentor Koch, 1844, Haemaphysalis Koch, 1844, Hyalomma Koch, 1844, Rhipicephalus Koch, 1844, and Ixodes Latreille, 1795. The genera Dermacentor and Hyalomma are represented by a two species, followed by Rhipicephalus with four species, Haemaphysalis with six species, and Ixodes with seven species. While studying the territory of Slavonia and Baranja during 2011, a total of 1425 ticks were collected. All the collected ticks were classified into 5 species. The identified species were classified into the following genera: Dermacentor, Haemaphysalis and Ixodes. Ixodes ricinus made up 83.44% of the tick fauna in the researched area, while 16.56% were representatives of other species.
Survey of hard ticks recorded in CroatiaFor every species are mentioned, sex, developmental stage ( ♂ male, ♀ female N = nymph, L = larva), sampling place, month or date, host and data source.
Family IXODIDAE Murray, 1877
Genus Dermacentor Koch 1844
1. Dermacentor marginatus (Sulzer, 1776)
♂♂, ♀ ♀ Cres Island: Hrasta 10.XI.1948, from Ovis aries (
♂♂, ♀ ♀ ibidem: Vodice 6.–11.IV. 1948, from Ovis aries (
♂♂, ♀♀ ibidem: Hrasta 29.IX.–1.X.1948, from Ovis aries (
♂♂, ♀♀ ibidem: Belej 29.IX.–1.X.1948, 4.XI. 1948, from Ovis aries (
♂♂, ♀♀ Pula, months: I. II. III. IV.V. IX. X. XI. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Krk Island, months: I. II. IX. X. XI. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Cres Island, months: I. II. III. IV. VIII. IX. X. XI. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Lošinj Island: Veli Lošinj, months: I. II. III. IV. VIII. IX. X. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ ibidem: Čunski-Like, month: VIII. 1948, from Ovis aries (
♂♂, ♀ ♀ Zadar, months: I. II. III. IV.V.VI. IX. X. XI. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Kaštela, month: VI. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (Mikačić, 1965)
♂♂, ♀♀ Split, months: I. II. III. IV.V.VI. XI. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Brač Island, months: I. II. III. IX. X. XI. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Dubrovnik, months: II. III. IV. IX. X. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Srednja Posavina, months: III. IV. V. VI. XI. XII. from Equus caballus, Bos taurus (
♂♂, ♀♀ Brač Island: Sveti Toma 4.–9.XI. 1965 from Ovis aries (
♂♂, ♀♀ ibidem: Osridke 4.–9.XI.1965 from Ovis aries (
♂♂, ♀♀ ibidem: Blato 4.–9.XI. 1965 from Ovis aries (
♂♂, ♀♀ ibidem: months: I. II. X. XI. XII. 1964/1965 collected on vegetation using the cloth dragging method (
♂♂, ♀♀ ibidem: Sveti Toma, months: I. II. XI. XII. 1964/1965 collected on vegetation using the cloth dragging method (
♂♂, ♀♀ ibidem: Zapadna Laščatna, months: I. X. XI. XII.1964/1965 collected on vegetation and/or the ground by flagging/dragging method (
♂♂, ♀♀ ibidem: Sjeverna Laščatna, months: XI. XII. 1964/1965 collected on vegetation using the cloth dragging method (
♂♂, ♀♀ ibidem: months: I. II. III. IX. X. XI. XII. 1964/1965 from Ovis aries (
NN ibidem: month:VII. 1964/1965 from Ovis aries (
♂♂, ♀♀, NN, LL Srednja Posavina: Stružec from 1970 to 1978 collected on vegetation using the cloth dragging method (
NN Peščenica-Vratovo 17.VII. 1974 from Microtus arvalis (
NN Velebit mountain: Predzid (pod Ćelavcem) 1.VIII. 1975 from Apodemus sylvaticus (
♂♂, ♀♀ Brač Island: Supetar 22.II. 1974 from Ovis aries (
L, NN Repušnica 5., 6.VIII. 1982 from Clethrionomys glareolus (
N ibidem: 6., 7.VIII. 1982 from Apodemus agrarius (
♂♂, ♀♀ Čakovec, months: III. IV. V. 2007 collected on vegetation using the cloth-dragging method (
♂♂, ♀♀ Kaštela Bay, month: X.2000 from Ovis aries, Capra hircus, Bos taurus (Punda -Polić et al. 2002)
♂♂, ♀♀ Sinj, month: X. 2000, from Ovis aries, Capra hircus, Bos taurus (
♂♂, ♀♀ Brač Island, Hvar Island (
♂♂, ♀♀ Đakovo 10.III.–30.III. 2011 from Canis lupus familiaris
♂♂, ♀♀ ibidem: 7.IV.–25.IV. 2011 from Canis lupus familiaris
♂♂, ♀♀ ibidem: 2.V.–25.V.2011 from Canis lupus familiaris
♂, ♀ Zmajevac 15.III.- 30.III. 2011 from Felis catus
♀♀ ibidem: 10.IV.–24.IV. 2011 from Felis catus
♀♀ ibidem: 25.IV.–30.IV. 2011 from Felis catus
♀ ibidem: 7.V.–25.V.2011 from Felis catus
2. Dermacentor reticulatus (Fabricius, 1794)
♂♂, ♀♀ Gornja Posavina, months: III. IV. V. from Equus caballus, Bos taurus (
♀ Kapelački Lug, month: V. 1967 collected on vegetation using the cloth dragging method (
♂♂, ♀♀, NN, LL Srednja Posavina: Stružec -, from 1970 to 1978 collected on vegetation using the cloth dragging method (
♂♂, ♀♀ Čakovec, months: III. IV. V. 2007 collected on vegetation using the cloth dragging method (
♀♀ Mikleuš 27.IV.2011 collected on vegetation using the cloth dragging method
♀♀ ibidem: 14.V.2011 collected on vegetation using the cloth dragging method
Genus Haemaphysalis Koch, 1844
3. Haemaphysalis punctata Canestrini & Fanzago, 1878
♂♂, ♀♀ Cres Island: Hrasta 6.–11.IV.1948 from Ovis aries (
♂♂, ♀♀ ibidem: 29.IX.–1.X. 1948 from Ovis aries (
♂♂, ♀♀ ibidem: 10.XI. 1948 from Ovis aries (
♂♂, ♀♀ ibidem: Vrana 6.–11.IV.1948 from Capra hircus (
♂♂, ♀♀, N ibidem: Vodice 6.–11.IV.1948 from Ovis aries (
♂♂, ♀♀ ibidem: Belej 29.IX.–1.X.1948 from Ovis aries (
♂♂, ♀♀ ibidem: 4.XI. 1948 from Ovis aries (
♂♂, ♀♀ Lošinj Island: Veli Lošinj 29.IX.–1.X. 1948 from Bos taurus (
♂♂, ♀♀ ibidem: Čunski-Like 29.IX.–1.X. 1948 from Ovis aries (
♂♂, ♀♀, NN, LL Pula, months: I. II. , III. , IV. V. VI. , VII. , VIII. IX. , X. XI. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Krk Island, months: I. II. IV. IX. X. XI. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Cres Island, months: I. II. III. IV.V. IX. X. XI. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Lošinj Island: Veli Lošinj, months: I. II. III. IV. IX. X. XI. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Zadar, months: I. II. III. IV. V. VI. IX. X. XI. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀, NN, LL Split, months: I. II. III. IV. X. XI. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Brač Island, months: II. III. VII. X. XI. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ ibidem: months: I. II.III. IV. V. IX. X. XI. XII 1964/1965 from Ovis aries (
♂♂, ♀♀, NN Dubrovnik, months: I. II. III. IV.V. IX. X. XI. XII.1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Gornja Posavina, month: V. from Equus caballus, Bos taurus (
♂♂, ♀♀ Brač Island: Sveti Toma 4–9.XI.1965 from Ovis aries (
♂♂, ♀♀ ibidem: Osridke 4.–9.XI. 1965 from Ovis aries (
♂♂, ♀♀ ibidem: Blato 4.–9.XI. 1965 from Ovis aries (
♂♂, ♀♀ ibidem: months: I. II. IV.V. XII. 1964/1965 collected on vegetation and/or the ground by flagging/dragging method (
♂♂, ♀♀ ibidem: Sveti Toma, months: I. II. IV.V. IX. X. XI. XII. 1964/1965 from Ovis aries (
♂♂, ♀♀ ibidem: Blato 5.XI.1965 from Ovis aries (
♂♂, ♀♀ ibidem: Zapadna Laščatna, months: I. IV. IX. X. XI. XII. 1964/1965 collected on vegetation and/or the ground by flagging/dragging method (
♂♂, ♀♀ ibidem: Sjeverna Laščatna, months: I. IV. XI. XII. 1964/1965 collected on vegetation and/or the ground by flagging/dragging method (
N Metković 23.IV. 1964 from Motacilla flava (
♂♂, ♀♀ Brač Island: Supetar 22.II. 1974 from Ovis aries (
♂♂, ♀♀ Kaštela Bay, month: X.2000 from Ovis aries, Capra hircus, Bos taurus (Punda -Polić et al. 2002)
♂♂, ♀♀ Sinj, month: X. 2000 from Ovis aries, Capra hircus, Bos taurus (
♂♂, ♀♀ Brač Island, Hvar Island (
4. Haemaphysalis inermis Birula, 1895
♂♂, ♀♀ Cres Island, months: I. II.1936–1963 from Ovis aries, Bos taurus, Canis lupus familiaris (
♂♂, ♀♀ Lošinj Island: Veli Lošinj, month: IX. 1936–1963 from Ovis aries, Bos taurus, Canis lupus familiaris (
♂♂, ♀♀ Zadar, month: V. 1936–1963 from Ovis aries, Bos taurus, Canis lupus familiaris (
♂♂, ♀♀ Dubrovnik, month: V. 1936–1963 from Ovis aries, Bos taurus, Canis lupus familiaris (
5. Haemaphysalis concinna Koch, 1844
♂ Dekanovec 10.VI. 1977 from Capreolus capreolus (
NN Krk Island: 15.V.2002 from Podarcis muralis, Podarcis melisellensis, Lacerta bilineata, Algyroides nigropunctatus, Elaphe longissima (
♂♂ Mikleuš 14.V.2011 collected on vegetation using the cloth-dragging method
NN ibidem: 8.VI.2011 collected on vegetation using the cloth-dragging method
♀, NN ibidem: 9.VI.2011 collected on vegetation using the cloth-dragging method
♂ ibidem: 15.VI.2011 collected on vegetation using the cloth-dragging method
NN, LL ibidem: 3.VII.2011 collected on vegetation using the cloth-dragging method
♂, NN ibidem: 4.VII.2011 collected on vegetation using the cloth-dragging method
NN, LL ibidem: 2.VIII.2011 collected on vegetation using the cloth-dragging method
NN, LL ibidem: 3.VIII.2011 collected on vegetation using the cloth-dragging method
N ibidem: 13.IX.2011 collected on vegetation using the cloth-dragging method
6. Haemaphysalis sulcata Canestrini & Fanzago, 1878
♂♂, ♀♀ Pula, months: X. XI. 1936–1963 from Ovis aries (
♂♂, ♀♀ Krk Island, months: IV. XI. XII. 1936–1963 from Ovis aries (
♂♂, ♀♀ Cres Island, months: I. II. III. IV. V. X. XI. XII. 1936–1963 from Ovis aries (
♂♂, ♀♀ Lošinj Island: Veli Lošinj, months: I. IV. X. 1936–1963 from Ovis aries (
♂♂, ♀♀ Zadar, months: I. II. III. IV. XI. XII. 1936–1963 from Ovis aries (
♂♂, ♀♀ Split, months: I. II. III. IV. 1936–1963 from Ovis aries (
♂♂, ♀♀ Brač Island, months: I. II. III. IV. V. X. XI. XII. 1964/1965 from Ovis aries (
NN ibidem: months: VII. VIII. 1964/1965 from Ovis aries (
♀ ibidem: Nadsela 1.X.1965 from Ovis aries (
L Krk Island: 23.VII.1956 from Algyroides nigropunctatus (
L ibidem: 23.VII.1956 from Podarcis melisellensis (
N Cres Island: Vrana, month: VII.1949 from Lacerta trilineata (
NN, LL Trstenik Island: 20.VII.1965 from Lacerta viridis (
NN Mali Dolfin reef: 20.IX.1961 from Podarcis sicula (
NN, LL Lutrošnjak Island: 25.VII.1965 from Podarcis sicula (
N, LL Kamenjak Island: 25.VII.1965 from Podarcis sicula (
NN, LL Premuda Island: 1.VIII.1965 from Podarcis sicula (
LL Zapadni Greben Island: 23.VII.1965 from Podarcis sicula (
LL Južni Greben Island: 23.VII.1965 from Podarcis sicula (
NN, LL Silba Island: 13.VIII.1963 from Podarcis sicula (
NN, LL ibidem: 22.VII.1965 from Podarcis sicula (
LL ibidem: 2.VIII.1965 from Podarcis sicula (
NN, LL Morovnik Island: 24.VII.1965 from Podarcis sicula (
NN, LL Olib Island: 3.VIII.1965 from Podarcis sicula (
LL Planičić reef: 3.VIII.1965 from Podarcis sicula (
LL Šip reef: 24.VII.1965 from Podarcis sicula (
L Vela Sestrica Island: 11.VIII.1963 from Podarcis sicula (
N, LL Dužac Island: 11.VIII.1963 from Podarcis sicula (
NN, LL Golac Island: 9.VIII.1965 from Podarcis sicula (
NN Mali Laganj Island: 12.VIII.1963 from Podarcis sicula (
NN, LL Baričevac Island: 6.VIII.1965 from Podarcis sicula (
NN, LL Mežanj Island: 8.VIII.1965 from Podarcis sicula (
LL Kurjak reef: 24.VII. 1965 from Podarcis melisellensis (
NN, LL Sparušnjak Island: 7.VIII.1965 from Podarcis melisellensis (
NN, LL Maslinovac Island: 7.VIII.1963 from Podarcis melisellensis (
NN, LL Vrtlac Island: 8.VIII.1963 from Podarcis melisellensis (
LL Veseljuh Island: 28.VII.1964 from Podarcis sicula (
NN, LL Bisaga Island: 7.VIII.1964 from Podarcis sicula (
N, LL Mali Tetovišnjak Island: 22.VII.1964 from Podarcis sicula (
LL Čerigul Island: 22.VII.1964 from Podarcis sicula (
LL Mali Kamešnjak Island: 23.VII.1964 from Podarcis sicula (
NN, LL Samograd Island: 6.VIII.1963 from Podarcis sicula (
LL Koromašna Island: 20.VII.1964 from Podarcis sicula (
NN, LL Gušteranski Island: 23.VII.1964 from Podarcis sicula (
N, LL Palagruža Island: 8.VI.1960 from Podarcis sicula (
LL Kamik od Tramuntane Island: 9.VI.1960 from Podarcis sicula (
LL Brusnik Island: 7.VI.1960 from Podarcis melisellensis (
NN Pod Mrčaru reef: 31.VII.1971 from Podarcis melisellensis (
NN, LL Božikovac Island: 1.VIII.1964 from Podarcis melisellensis (
N, LL Mala Lavdara Island: 1.VIII.1964 from Podarcis melisellensis (
NN, LL Trimulići Island: 2.VIII.1964 from Podarcis melisellensis (
NN, LL Trstikovac Island: 2.VIII.1964 from Podarcis melisellensis (
LL Vela Skala Island: 2.VIII.1964 from Podarcis melisellensis (
NN, LL Mala Skala Island: 2.VIII.1964 from Podarcis melisellensis (
NN, LL Rončić Island: 1.VIII.1964 from Podarcis melisellensis (
NN, LL Mali Babuljaš Island: 7.VIII.1963 from Podarcis melisellensis (
NN, LL Malo Šilo Island: 6.VIII.1964 from Podarcis melisellensis (
NN, LL Mrtovac Island: 6.VIII.1964 from Podarcis melisellensis (
NN, LL Veli Babuljaš Island:7.VIII.1964 from Podarcis melisellensis (
NN, LL Mali Prišnjak Island: 7.VIII.1964 from Podarcis melisellensis (
NN, LL Vodenjak Island: 7.VIII.1964 from Podarcis melisellensis (
NN, LL Jančar Island: 7.VIII.1964 from Podarcis melisellensis (
NN, LL Purara Island: 7.VIII.1964 from Podarcis melisellensis (
NN, LL Mala Prduša Island: 7.VIII.1964 from Podarcis melisellensis (
NN, LL Babina Guzica Island: 8.VIII.1964 from Podarcis melisellensis (
LL Veli Puh Island: 21.VII.1964 from Podarcis melisellensis (
LL Vodeni Puh Island: 21.VII.1964 from Podarcis melisellensis (
NN, LL Kameni Puh Island: 21.VII.1964 from Podarcis melisellensis (
LL Čavlin Island: 22.VII.1964 from Podarcis melisellensis (
LL Veli Tetovišnjak Island: 22.VII.1964 from Podarcis melisellensis (
LL Veli Kamešnjak Island: 23.VII.1964 from Podarcis melisellensis (
N, LL Žirje Island: 27.VII.1964 from Podarcis melisellensis (
N, L Mikavica Island: 4.VIII.1964 from Podarcis melisellensis (
LL Škrovada reef: 23.VII.1964 from Podarcis melisellensis (
NN, LL Kosmerka Island: 4.VIII.1964 from Podarcis melisellensis (
L Pod Mrčaru reef: 31.VII.1971 from Dalmatolacerta oxycephala (
♂♂, ♀♀ Southern Croatia from Ovis aries and Capra hircus (
7. Haemaphysalis erinacei Pavesi, 1884
N Dubrovnik 10.VIII. 1964 from Tachymarptis melba (
8. Haemaphysalis parva (Neumann, 1897)
♂♂, ♀♀ Brač Island: (
Genus Hyalomma Koch, 1844
9. Hyalomma scupense Schulze, 1919
♂♂, ♀♀ Pula, months: III. IV. IX. 1936–1963 from Ovis aries, Bos taurus (
♂♂, ♀♀ Krk Island, months: II. III. IV. 1936–1963 from Ovis aries, Bos taurus (
♂♂, ♀♀ Lošinj Island: Veli Lošinj, month: IV. 1936–1963 from Ovis aries, Bos taurus (
♂♂, ♀♀ Zadar, months: II. III. IV. VI. VIII. 1936–1963 from Ovis aries, Bos taurus (
NN ibidem: months: IX. XI. 1936–1963 from Ovis aries, Bos taurus (
♂♂, ♀♀ Split, months: I. II. III. IV. VI. 1936–1963 from Ovis aries, Bos taurus (
NN ibidem: months: XI. XII. 1936–1963 from Ovis aries, Bos taurus (
♂♂, ♀♀ Brač Island, months:V.VI. 1936–1963 from Ovis aries, Bos taurus (
♂♂, ♀♀ ibidem: 1964/1965 from Ovis aries (
♂♂, ♀♀ ibidem: -, (
♂♂, ♀♀, NN Dubrovnik, month: I. 1936–1963 from Ovis aries, Bos taurus (
♂♂, ♀♀ ibidem: months: II. III. IV.V. IX. XI. XII.1936–1963 from Ovis aries, Bos taurus (
10. Hyalomma marginatum Koch, 1844
♂♂, ♀♀ Pula, months: I. II. III. IV. V. VI. VII. VIII. IX. X. XI. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Krk Island, months: II. III. IV. V. VI. VII. VIII. IX. X. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Cres Island, months: IV. VI. VII. X. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Lošinj Island: Veli Lošinj, months: II. III. IV. V. VI. VII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Zadar, months: II. III. IV. V. VI. VII. VIII. X. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Split, months: II. III. IV. V. VI. VII. VIII. IX. X. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ Dubrovnik, months: III. IV. V. VI. VII. VIII. IX. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂, ♀ Petrinja, month: V. from Equus caballus, Bos taurus (
♂♂, ♀♀ Brač Island, months: I. II. III. IV. V. VI. VII. VIII. IX. X. 1936–1963 from Ovis aries, Equus caballus, Bos taurus (
♂♂, ♀♀ ibidem: months: V. VI. VII. VIII. IX. X. 1964/1965 from Ovis aries (
♂, ♀ ibidem: Blato, month: IV.1964/1965 from Ovis aries (
♂♂, ♀♀ ibidem: Zapadna Laščatna, months: IV. VII.1964/1965 collected on vegetation and/or the ground by flagging/dragging method (
♂♂, ♀♀ ibidem: (
NN, LL ibidem: month: VII. 1967 from Erinaceus europaeus (
♂♂, ♀♀ Kaštela Bay, month: X. 2000 from Ovis aries, Capra hircus, Bos taurus (Punda -Polić et al. 2002)
♂♂, ♀♀ Sinj, month: X. 2000 from Ovis aries, Capra hircus, Bos taurus (
Genus Rhipicephalus Koch, 1844
11. Rhipicephalus annulatus (Say, 1821)
♂♂, ♀♀ Lošinj Island: Veli Lošinj 29.IX.–1.X.1948 from Bos taurus (
♂♂, ♀♀ ibidem: months: IV. V. X. 1936–1963 from Equus caballus, Bos taurus (
♂♂, ♀♀ Zadar, months: III. IV.V. VI. VII. VIII. IX. 1936–1963 from Equus caballus, Bos taurus (
NN ibidem: months: IV. VII. VIII. IX. 1936–1963 from Equus caballus, Bos taurus (
♂♂, ♀♀ Split, months: VI. VII. VIII. IX. X. XI. 1936–1963 from Equus caballus, Bos taurus (
NN ibidem: months: IX. XI. 1936–1963 from Equus caballus, Bos taurus (
♂♂, ♀♀ Brač Island: month: VII. 1936–1963 from Equus caballus, Bos taurus (
♂♂, ♀♀ ibidem: (
♂♂, ♀♀ Dubrovnik, months: III. IV. V. VII. 1936–1963 from Equus caballus, Bos taurus (
NN ibidem: months: IV.XI. 1936–1963 from Equus caballus, Bos taurus (
12. Rhipicephalus bursa Canestrini & Fanzago, 1878
♂♂, ♀♀ Lošinj Island: Čunski-Like 6.–11.IV.1948 from Ovis aries (
♂♂, ♀♀ Cres Island, months:, IV. V. 1948 from Ovis aries (
♂♂, ♀♀ ibidem: month: VIII. 1948 from Ovis aries (
LL ibidem: 29.IX.–1.X.1948 from Ovis aries (
♂♂, ♀♀, NN, LL ibidem: Hrasta 6.–11.IV. 1948 from Ovis aries (
♂♂, ♀♀, NN, LL ibidem: 29.IX.–1.X.1948 from Ovis aries (
NN, L ibidem: 10.XI.1948 from Ovis aries (
NN ibidem: Vrana 6.–11.IV. 1948 from Capra hircus (
♂♂, ♀♀, NN ibidem: Vodice 6.–11.IV. 1948 from Ovis aries (
♂♂, ♀♀ ibidem: Orlec 6.–11.IV. 1948 from Ovis aries (
♂♂, ♀♀ ibidem: Belej 29.IX.–1.X.1948 from Ovis aries (
NN ibidem: 4.XI.1948 from Ovis aries (
♂♂, ♀♀ Pula, months: II. III. IV. V. VI. VIII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
NN ibidem: months: I. II. III. IV. X. XI. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
LL ibidem: months: III. IV. XI. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
♂♂, ♀♀ Krk Island, months: II. III. IV. V. VI. VII.VIII. X. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
NN ibidem: months: I. III. IV.VI. X. XI. XII.1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
LL ibidem: months: III. IV. X. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
♂♂, ♀♀ Cres Island: Cres, months: III. IV. V. VI. VII. IX. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
NN ibidem: months: I. II. III. IV. V. XI. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
LL ibidem: months: I. III. XI. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
♂♂, ♀♀ Lošinj Island: Veli Lošinj, months: III. IV. V. VI. VII. VIII. IX. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
NN ibidem: months: II. III. IV.X. XI. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
LL ibidem: months: III. IV. IX. XI. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
♂♂, ♀♀ Zadar, months: III. IV. V. VI.VII. VIII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
NN ibidem: months: I. II. III. IV. V. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
LL ibidem: month: I. III. IV. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
♂♂, ♀♀ Split, months: II. III. V. VI. VII. VIII. IX. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
NN ibidem: months; II. III. X. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
LL ibidem: month: III. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
♂♂, ♀♀ Brač Island, months: II. III. IV. V.VI. VII.VIII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
NN ibidem: months: I. II. III. IV. X. XI. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
♂♂, ♀♀ ibidem: 1964/1965 from Ovis aries (
♂♂, ♀♀ ibidem: (
NN ibidem: Blato 4.–9.XI. 1965 from Ovis aries (
♂, ♀ ibidem: Blato, month: IX.1964/1965 collected on vegetation and/or the ground by flagging/dragging method (
♂♂, ♀♀ ibidem: Sveti Toma, months: IX. X. 1964/1965 collected on vegetation and/or the ground by flagging/dragging method (
♂♂, ♀♀ ibidem: Zapadna Laščatna, months: VII. VIII. 1964/1965 collected on vegetation and/or the ground by flagging/dragging method (
♂♂, ♀♀ Dubrovnik, months: III. IV.V. VI.VII. VIII. IX. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
NN ibidem: months: II. IV. V. X. XI. XII. 1936–1963 from Ovis aries, Equus caballus, Bos taurus, Capra hircus (
♂♂, ♀♀ Kaštela Bay, month: X. 2000 from Ovis aries, Capra hircus, Bos taurus (
♂♂, ♀♀ Sinj, month: X. 2000 from Ovis aries, Capra hircus, Bos taurus (
13. Rhipicephalus sanguineus (Latreille, 1806)
♂♂, ♀♀ Mali Lošinj Island: Čunski-Like 6.–11.IV. 1948 -, V. 1948 from Ovis aries (
♂♂, ♀♀ Cres Island: Hrasta 6.–11.IV. 1948 from Ovis aries (
♂♂, ♀♀ ibidem: Belej 6.–11.IV. 1948 from Ovis aries (
♂♂, ♀♀ ibidem: Vrana 6.–11.IV. 1948 from Capra hircus (
♂♂, ♀♀ ibidem: Vodice 6.–11.IV. 1948 from Ovis aries (
♂♂, ♀♀ ibidem: Orlec. months: IV. V. 1948 from Ovis aries (
♂♂, ♀♀ ibidem: Cres, months: IV. V. VII. 1936–1963 from Ovis aries, Bos taurus, Canis lupus familiaris (
♂♂, ♀♀ Pula, months: V.VI. 1936–1963 from Ovis aries, Bos taurus, Canis lupus familiaris (
♂ ibidem: 18.V.1973 from Canis lupus familiaris (
♂♂, ♀♀ Zadar, months: III. IV. V. VI. 1936–1963 from Ovis aries, Bos taurus, Canis lupus familiaris (
♂♂, ♀♀ Split, months: III. IV. V. VI. 1936–1963 from Ovis aries, Bos taurus, Canis lupus familiaris (
♂♂, ♀♀ Dubrovnik, months: III. IV. V. VI. VII. VIII. 1936–1963 from Ovis aries, Bos taurus, Canis lupus familiaris (
♂♂, ♀♀ Brač Island, months: II. III. IV. 1936–1963 from Ovis aries, Bos taurus, Canis lupus familiaris (
♂♂, ♀♀ ibidem: from 1964/1965 from Ovis aries (
♂♂, ♀♀ ibidem: Blato, months; IV. V. VI. 1964/1965 collected on vegetation and/or the ground by dragging/flagging method (
♂♂, ♀♀ ibidem: Sveti Toma, months: IV. V. VI. VII. 1964/1965 collected on vegetation and/or the ground by dragging/flagging method (
♂♂, ♀♀ ibidem: Zapadna Laščatna, months: IV. VI. VII.1964/1965 collected on vegetation and/or the ground by dragging/flagging method (
♂♂, ♀♀ ibidem: Sjeverna Laščatna, months: IV. VI. 1964/1965 collected on vegetation and/or the ground by dragging/flagging method (
♂♂, ♀♀ ibidem: 1.–4.V. 1964 and 28.III.–1.IV.1965 collected on vegetation by dragging method (
♂♂, ♀♀ ibidem: 28.III. , 27.IV., 1.VI., 4.VI. 1965 from Ovis aries, (Tovornik and Vesenjak -Hirjan 1988)
♂♂, ♀♀ ibidem: (
♂♂, ♀♀ Umag 27.V.1978 from Canis lupus familiaris (
♀ Slano (Dubrovnik) 10.V.1953 from Canis aureus (Tovornik and Vesenjak -Hirjan 1988)
♂♂, ♀♀ Ugljan Island: Kali 9.–20.V.1955 from Ovis aries (Tovornik and Vesenjak -Hirjan 1988)
♂♂, ♀♀ Rovanjska 23.IV.1958 from Ovis aries (
♂♂, ♀♀ Zadar 25.IV.1958 from Ovis aries (
♂♂ Pašman Island: 2.V.1985 from Ovis aries (
♂, ♀ Dugi otok Island: 29.IV.1986 from Ovis aries (
♂♂, ♀♀ Mljet Island: Babino polje 8.–9.IV.1981, 16.–17.V. 1981 from Capra hircus, Ovis aries, Canis lupus familiaris, Equus asinus (Tovornik and Vesenjak -Hirjan 1988)
♂♂, ♀♀ ibidem: Zadublje 8.–9.IV.1981 from Capra hircus, Ovis aries, Canis lupus familiaris, Equus asinus (
♂♂, ♀♀ ibidem: Veliko jezero 8.–9.IV.1981 from Capra hircus, Ovis aries, Canis lupus familiaris, Equus asinus (
♂♂, ♀♀ ibidem: Soline 8.–9.IV.1981 from Capra hircus, Ovis aries, Canis lupus familiaris, Equus asinus (
♂♂, ♀♀ ibidem: Babino polje 16.–17.V. 1981 from Capra hircus, Ovis aries, Canis lupus familiaris, Equus asinus (
♂♂, ♀♀ ibidem: Blato 16.–17.V. 1981 from Capra hircus, Ovis aries, Canis lupus familiaris, Equus asinus (
14. Rhipicephalus turanicus Pomerantsev, 1940
♂♂, ♀♀ Ugljan Island: Kali 9.–20.V.1955 from Ovis aries (Tovornik and Vesenjak -Hirjan 1988)
♂ Zadar 25.IV. 1958 from Ovis aries (
♂ Lokrum Island: 9.IX.1963 from Capreolus capreolus (
♂ ibidem: 4.IX. 1963 from Capreolus capreolus (
♂♂, ♀ Brač Island: 1.–4.V.1964 and 28.III.–1.IV.1965 collected on vegetation by dragging method (
♂♂, ♀♀ ibidem: 28.III., 27.IV., 1.VI., 4.VI., 1965 from Ovis aries (
♂♂, ♀♀ Pašman Island: 2.V.1985 from Ovis aries (
♂♂, ♀ Dugi otok Island: 29.IV.1986 from Ovis aries (Tovornik and Vesenjak -Hirjan 1988)
♂♂, ♀♀ Mljet Island: 8.–9.IV. 1981, 16–17.V. 1981 from Capra hircus and other domestic animals (
♂♂, ♀♀ ibidem: Babino polje 8.–9.IV.1981 from Capra hircus, Ovis aries, Felis catus, Canis lupus familiaris (
♂♂, ♀♀ ibidem: Zadublje 8.–9.IV.1981 from Capra hircus, Ovis aries, Canis lupus familiaris, Felis catus (
♂♂, ♀♀ ibidem: Veliko jezero 8.–9.IV.1981 from Capra hircus, Ovis aries, Canis lupus familiaris, Felis catus (
♂♂, ♀♀ ibidem: Soline 8.–9.IV.1981 from Capra hircus, Ovis aries, Canis lupus familiaris, Felis catus (
♂♂, ♀♀ ibidem: Babino polje 16.–17.V. 1981 from Capra hircus, Ovis aries, Canis lupus familiaris, Felis catus (
♂♂, ♀♀ ibidem: Blato 16.–17.V. 1981 from Capra hircus, Ovis aries, Canis lupus familiaris, Felis catus (
Genus Ixodes Latreille, 1795
15. Ixodes arboricola Schulze & Schlottke, 1930
♀♀ Zagreb 29.IX. 1962 from Turdus merula (
16. Ixodes frontalis (Panzer, 1798)
♀ Velebit mountain: Donja Klada, 1975 from Martes foina (
♀ Dubrovnik 22.II.1962 from Bombycilla garrulus (
♀ ibidem: 1.XII. 1967 from Phoenicurus ochruros (
♀ Zagreb, vicinity, 20.IV. 1975 from Martes foina (
17. Ixodes gibbosus Nuttal, 1916
♂♂, ♀♀ Brač Island: Sveti Toma 4–9. XI. 1965 from Ovis aries (
♂♂, ♀♀ ibidem: Osridke 4–9.XI.1965 from Ovis aries (
♂♂, ♀♀ ibidem: Blato 4.–9.XI. 1965 from Ovis aries (
♂♂, ♀♀, NN, LL ibidem: Zapadna Laščatna 24.–25.XI. 1964 collected on the vegetation by dragging method (
♂♂, ♀♀ ibidem: Blato, months: I. II. IV. V. XII. 1964/1965 collected on vegetation and/or the ground by dragging/flagging method (
♂♂, ♀♀ ibidem: Sveti Toma, months: I. II. IV. V. XI. XII. 1964/1965 collected on vegetation and/or the ground by dragging/flagging method (
♂♂, ♀♀ ibidem: Zapadna Laščatna, months: I. IV. XI. XII. 1964/1965 collected on vegetation and/or the ground by dragging/flagging method (
♂♂, ♀♀ ibidem: Sjeverna Laščatna, months: I. IV. XI. XII. 1964/1965 collected on vegetation and/or the ground by dragging/flagging method (
♂♂, ♀♀ ibidem: Supetar 22.II. 1974 from Ovis aries (
♂♂, ♀♀ ibidem: (
18. Ixodes hexagonus Leach, 1815
♂♂, ♀♀ Gornja Posavina, month: III. from Equus caballus, Bos taurus (
NN Velika Kapela mountain: Razvala 16.VIII. 1969 from Erinaceus concolor (
♀ Dugo Selo 5.IV. 1959 from Martes martes (
♀♀ Đakovo 7.IV.–25.IV. 2011 from Canis lupus familiaris
♂♂ ibidem: 2.V.–25.V. 2011 from Canis lupus familiaris
19. Ixodes ricinus (Linnaeus, 1758)
♂♂, ♀♀, NN, LL Cres Island: Hrasta 6.–11.IV.1948 from Ovis aries (
♂, ♀♀ ibidem: 10.XI.1948 from Ovis aries (
♂♂, ♀♀, NN ibidem: Vrana 6.–11.IV.1948 from Capra hircus (
♂♂, ♀♀, NN, LL ibidem: Vodice 6.–11.IV.1948 from Ovis aries (
NN ibidem: Cres 6.–11.IV.1948 from Capra hircus (
♂♂, ♀♀, NN ibidem: Belej 4.XI.1948 from Ovis aries (
♂♂, ♀♀ Pula, months: I. II. III. IV. V. VI. VIII. IX. X. XI. XII. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
NN ibidem, months: I. II. III. IV. X. XII. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
LL ibidem, months: IV. VII. XII. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
♂♂, ♀♀ Krk Island, months: I. II. III. IV. V. IX. XI. XII. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
NN ibidem, months: I. II. III. IV. IX. X. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
LL ibidem, months: I. II. III. IV. X. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
♂♂, ♀♀ Cres Island, months: I. II. III. IV.V. IX. X. XI. XII. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
NN ibidem, months: I. II. III. IV. IX. XI. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
LL ibidem, months: II. III. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
♂♂, ♀♀ Lošinj Island: Veli Lošinj, months: I. II. III. IV. V. IX. X. XI. XII. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
NN ibidem: months: I. II. III. IV. V. IX. X. XI. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
LL ibidem: months: II. III. IV. X. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
♂♂, ♀♀ Zadar, months: I. II. III. IV. VI. IX. X. XI. XII. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
NN ibidem: months: I. III. IV. IX. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
LL ibidem: months: I. IV. IX. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
♂♂, ♀♀ Split, months: I. II. III. IV. X. XI. XII. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
NN ibidem: months: I. II. III.1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
LL ibidem: month: II. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
♂♂, ♀♀ Brač Island, months: I. III. IX. X. XI. XII. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
NN ibidem: months: I. II. III. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
♂♂, ♀♀ Dubrovnik, months: I. II. III. IV.V. IX. X. XI. XII. 1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
NN ibidem: months: I. II. IV. XI. XII.1936–1963 from Ovis aries, Bos taurus, Capra hircus, Equus caballus, Equus asinus (
N, L Rovinj 7.IV.1959 from Algyroides nigropunctatus (
N, L ibidem: 7.IV.1959 from Podarcis melisellensis (
L ibidem: 10.IV. 1959 from Podarcis melisellensis (
NN, LL Tovarjež Island: 7.V.1959 from Podarcis sicula (
N, L Lakal Island: 7.V.1959 from Podarcis sicula (
NN, LL Lunga Island: 6.IV.1959 from Podarcis sicula (
NN, LL Koversada Island: 4.IV.1959 from Podarcis sicula (
N, L Figarola Island: 9.IV.1959 from Podarcis sicula (
N, L Banjol Island: 11.IV.1959 from Podarcis sicula (
N Veliki Piruzi reef: 4.IV.1959 from Podarcis sicula (
LL Sturag Island: 12.VIII.1959 from Podarcis sicula (
LL Sveti Ivan Island: 12.VIII.1959 from Podarcis sicula (
L Sveti Ivan na Pučini reef: 4.IV.1959 from Podarcis sicula (
NN, LL Pulari Island: 8.V.1959 from Podarcis sicula (
LL Revera Island: 8.V.1959 from Podarcis sicula (
N, LL Veštar Island: 8.IV.1959 from Podarcis sicula (
NN, LL Velika Sestrica Island: 8.IV.1959 from Podarcis sicula (
NN, LL Pisulj Island: 8.V.1959 from Podarcis sicula (
NN, LL Gustinja Island: 8.V.1959 from Podarcis sicula (
NN, LL Unije Island: 17.VII.1960 from Podarcis melisellensis(
NN, LL ibidem: 17.VII. 1960 from Podarcis sicula (
N, LL Zeča Island: 18.VIII.1963 from Podarcis sicula (
N, LL Vele Srakane Island: 19.VII.1960 from Podarcis sicula (
LL Susak Island: 19.VII.1960 from Podarcis sicula (
NN, LL Oruda Island: 19.VII.1960 from Podarcis sicula (
NN, LL Palacol Island: 19.VII.1960 from Podarcis sicula (
N, LL Male Orjule Island: 19.VII.1960 from Podarcis sicula (
N, LL Školjić Island: 19.VII.1960 from Podarcis sicula (
NN, LL Sveti Petar Island: 29.VII.1965 from Podarcis sicula (
LL Ilovik Island: 29.VII.1965 from Podarcis sicula (
LL Grujica Island: 28.VII.1965 from Podarcis sicula (
♂♂, ♀♀, NN, LL Srednja Posavina: Stružec, 1970–1978 collected on vegetation and/or the ground by dragging/flagging method (
♂♂, ♀♀ Gornja Posavina, months: III. IV. V. VI. VII. IX. X. XI. (
NN ibidem, months: III. IV. V. VI. (
LL ibidem, months: IV. V. (
♂♂, ♀♀ Stara Ves 22.III.1963 collected in the forest biotopes (
♀ Zagreb 5.IV. 1959 from Martes martes (
LL Velika Kapela mountain: Razvala 16.VIII.1968 from Clethrionomys glareolus (
N, LL Lika: Plitvice, Crna rijeka 12.VIII. 1968 from Apodemus sylvaticus (
N, LL ibidem: 12.VIII. 1968 from Apodemus flavicollis (
N Vukomeričke gorice: Prkovec 19.VII. 1974 from Apodemus flavicollis (
L Kozjak mountain: Malačka 29.IV. 1974 from Apodemus mystacinus (
♀♀, N Dugo Selo 9.X.1964 from Capreolus capreolus (
♂, ♀♀, NN Dekanovec from Capreolus capreolus (
♂♂, ♀♀, NN Nedelišće 3.V.1978 from Capreolus capreolus (
N Dubrovnik 12.II. 1966 from Buteo buteo (
♀, NN, LL Stara Ves 21.IV. 1964 from Anthus trivialis (
NN, LL ibidem: 10.V. 1964 from Anthus trivialis (
NN, LL ibidem: 24.VI.1964 from Pica pica (
ibidem: 15., 16. VII. 1964 from Pica pica (
N Baćinska jezera 30.IV. 1976 from Cettia cetti (
L Vrana-Biograd 25.IV. 1974 from Luscinia megarhynchos (
NN ibidem: 25.IV.1974 from Turdus merula (
NN Metković-Gabela 14.X. 1964 from Turdus philomelos (
L Peščenica-Vratovo 26–27.IV.1984 from Clethrionomys glareolus (
N ibidem: month: V. 1984 from Apodemus flavicollis (
N, LL ibidem: 3.VI. 1984 from Apodemus flavicollis (
NN ibidem: 1.VII.1984 from Apodemus flavicollis (
LL ibidem: 2.VII.1984 from Apodemus flavicollis (
N Peščenica-Vratovo 22.–23.III.1984 from Apodemus agrarius (
N ibidem: 2.VII.1984 from Clethrionomys glareolus (
♂♂, ♀♀ Northern Croatia (
♂♂, ♀♀ Lukovdol, Lipov vrh months: V.- VIII. 2008 from Capreolus capreolus (
♂♂, ♀♀, NN Zmajevac 5.III.–30.III. 2011 from Felis catus
♂♂, ♀♀, NN ibidem: 10. IV.–24.IV. 2011 from Felis catus
♂♂, ♀♀, NN ibidem: 25.IV.- 30.IV. 2011 from Felis catus
♂♂, ♀♀, N ibidem: 7.V.- 25.V. 2011 from Felis catus
LL ibidem: 31.VII.2011 from Homo sapiens
♀ ibidem: 26.X. 2011 from Felis catus
♂, ♀♀, NN Đakovo 10.III.–30.III. 2011 from Canis lupus familiaris
♂♂, ♀♀, NN ibidem: 7.IV.- 25.IV. 2011 from Canis lupus familiaris
♀♀ ibidem: 2.V.–25.V. 2011 from Canis lupus familiaris
♀, NN Sungerski Lug 25.VI. 2011 from Canis lupus familiaris
♂♂, ♀♀, NN Mikleuš 27.IV. 2011 collected on vegetation using the cloth-dragging method
♂♂, ♀♀, NN, L ibidem: 14.V. 2011 collected on vegetation using the cloth-dragging method
♂♂, ♀♀, NN ibidem: 8.VI. 2011 collected on vegetation using the cloth-dragging method
♂, ♀, NN ibidem: 9.VI.2011 collected on vegetation using the cloth-dragging method
♂♂, ♀♀, NN ibidem: 15.VI.2011 collected on vegetation using the cloth-dragging method
♂♂, ♀♀, NN ibidem: 3.VII.2011 collected on vegetation using the cloth-dragging method
♂♂, ♀♀, NN ibidem: 4.VII.2011 collected on vegetation using the cloth-dragging method
♂♂, ♀♀, NN, LL ibidem: 2.VIII.2011 collected on vegetation using the cloth-dragging method
♂, ♀♀, NN, LL ibidem: 3.VIII.2011 collected on vegetation using the cloth-dragging method
♀♀, N ibidem: 13.IX.2011 from Canis lupus familiaris
20. Ixodes trianguliceps Birula, 1895
♂♂, ♀♀ Rijeka (
♂♂, ♀♀ Split (
♂♂, ♀♀ Kozjak mountain: Malačka (
NN, LL ibidem: 29.IV. 1974 from Apodemus mystacinus (
♂♂, ♀♀ Velebit mountain (
L ibidem: Veliki Zavižan 28.IX. 1975 from Sorex araneus (
♀ ibidem: Crni padež 1.VIII. 1975 from Clethrionomys glareolus (
L ibidem: Baške Oštarije 11.VIII. 1968 from Apodemus sylvaticus (
L ibidem: Predzid (pod Ćelavcem) 1.VIII.1975 from Apodemus sylvaticus (
♂♂, ♀♀ Istra: Učka mountain (
LL ibidem: 12.VII. 1969 from Apodemus flavicollis (
♂♂, ♀♀ Gorski Kotar: Risnjak mountain (
L ibidem: 9.VIII. 1967 from Sorex alpinus (
♀, L ibidem: 1.IX. 1978 from Clethrionomys glareolus (
♀ Velika Kapela mountain: Razvala 16.VIII. 1968 from Clethrionomys glareolus (
LL Lika: Plitvice, Crna rijeka 12.VIII. 1968 from Apodemus sylvaticus (
LL ibidem: 12.VIII. 1968 from Apodemus flavicollis (
N ibidem: Mukinje 29.V. 1977 from Clethrionomys glareolus (
♀ ibidem: 29.V. 1975 from Sorex araneus (
♀, N, L Medvednica mountain: Sljeme 25.VI. 1975 from Apodemus flavicollis (
N Vukomeričke gorice: Prkovec 19.VII.1974 from Apodemus flavicollis (
N ibidem: 19.VII. 1974 from Apodemus agrarius (
♀♀ Peščenica-Vratovo 5.X. 1977 from Apodemus flavicollis (
N, LL ibidem: 28.X. 1976 from Sorex araneus (
♀ Križevci, Stara Ves 20.IV.1963 from Apodemus sylvaticus (
♀ Peščenica-Vratovo 26.–27.IV. 1984 from Apodemus flavicollis (
N ibidem: 2.VII.1984 from Apodemus flavicollis (
NN ibidem: 3.VI. 1984 from Apodemus flavicollis (
N ibidem: 1.VIII.1984 from Sorex araneus (
♀ ibidem: 20.–24.IX. 1983 from Apodemus agrarius (
L Peščenica-Vratovo 29.–30.XI.1983 from Neomys sp., (
21. Ixodes vespertilionis Koch, 1844
♀, N Dubrovnik, Močiljska špilja 31.III.1953 from Rhinolophus ferrumequinum, Rhinolophus hipposideros, Myotis myotis, Miniopterus schreibersii and from cave habitats (
♀, L ibidem: 6.V.1961 from Rhinolophus ferrumequinum, Rhinolophus hipposideros, Myotis myotis, Miniopterus schreibersii and from cave habitats (
♂ Krbavsko polje: 17.IX.1963 cave habitats (
♀ Cres Island: Petrićevi-špilja Čampari 31.III.1975 from Rhinolophus ferrumequinum, Rhinolophus hipposideros, Myotis myotis, Miniopterus schreibersii and from cave habitats (
♂ Buzet, Rabakova peć, Ročko polje, month: VII.1961 from Rhinolophus ferrumequinum, Rhinolophus hipposideros, Myotis myotis, Miniopterus schreibersii and from cave habitats (
♂♂, ♀♀ Brač Island (
Creating a list of ticks present in the Croatian fauna is somewhat complicated because the territory of Croatia was formerly part of the Kingdom of Yugoslavia and later part of the Socialist Federal Republic of Yugoslavia. Before the disintegration of Yugoslavia, nearly all of the records were simply summarized as being from Yugoslavia and the faunas of the new European states were not differentiated. For instance,
I would like to thank Dr. Tomi Trilar (Slovenian Museum of Natural History, Ljubljana, Slovenia) for confirming the collected materials and for literature data. Furthermore, I would like to thank Dr. Đurđica Stubičan for assistance in collecting the literature data (Faculty of Veterinary Medicine, University of Zagreb, Croatia) and thank the students Marko Teni for help in field work. Also, I thank Dr. Lawrence J. Hribar of the Florida Keys Mosquito Control District, USA for all improvements in this manuscript.