Hippa et al. 2005 (phylogeny); Jaschhof and Hippa 2005 (identification key to Costa Rican species); Hippa and Huerta 2009 (new species from Mexico); Hippa and Kurina 2013 (new species from Ecuador, French Guyana, Nicaragua, Argentina and Peru); Oliveira and Amorim 2014 (catalogue of Neotropical Mycetophilidae); Hippa et al. 2017 (new species from Peru).

Holotype. Male, COLOMBIA, Risaralda, SFF Otún Quimbaya Cuchilla Camino, 04°43'N, 75°35'W, 2050 m, Malaise trap, 03–19.i.2003, G. López Leg. M. 3702 (on slide, IAvH).

Figure 1. 

Manota clava sp. n. (holotype). A Hypopygium, ventral view B Hypopygium, dorsal view C Aedeagus and hypoproct, ventral view D Juxtagonostylar megasetae with associated parts, mediodorsal view. Scale bar 0.10 mm. Abbreviations: aaed = apex of aedeagus, aeda = aedeagal apodeme,ce = cercus, dlb = plates posteriorly at dorsal medial margin of gonocoxa, dm = dorsal medial margin of gonocoxa, gs = gonostylus, gx = gonocoxa, gxa = gonocoxal apodeme, hpr = hypoproct, jxs = juxtagonostylar megaseta, psl = parastylar lobe, st9 = sternite 9, tg9 = tergite 9, vm = ventral medial margin of gonocoxa.

Laterotergite setose; anterior basalare non-setose; sternite 9 posteriorly broadly concave and laterally free from gonocoxa; parastylar lobe large and apically broadened; gonocoxa without a remarkable posterolateral lobe; gonostylus subrectangular, posterolaterally drawn out; two juxtagonostylar megasetae, ventral one flame-shaped and pointed, dorsal one bilobed.

Male. Colour. Head brown, face somewhat paler. Antenna light brown, including scape and pedicel. Clypeus and mouthparts pale yellow. Thorax light brown. Legs yellowish. Wing with light brownish tinge because of microtrichia; halter brownish with blackish knob. Abdomen with tergites dark brown to blackish, sternites light brown to yellowish. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 2 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 3 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 ca. 1.1 times longer than palpomere 4. Number of strong postocular setae, 9. Thorax. Anepisternum with 52 setae; anterior basalare and preepisternum 2 non-setose; laterotergite with 13 setae; metepisternum with 9 setae. Legs. Mid and hind tibial organs absent. Wing. R₁ meeting C within basal half of costal margin; sclerotized part of M₂ extending to level of tip of R₁; wing length, 2.4 mm. Hypopygium (Fig. 1A–D). Sternite 9 ca. 2/3 as long as gonocoxa with delimited lateral margins, broadly concave posteriorly, anteriorly incised; posterior half covered with setae which are slightly stronger than adjacent ventral setae of gonocoxa, anterior half non-setose. Ventral medial margin of gonocoxa simple. Parastylar lobe large, apically broadened, club-like, with 3-5 fine posteriorly directed setae medially. No paraapodemal lobe observable. Posterolateral part of gonocoxa not drawn into a remarkable lobe but bearing many long curved setae. Dorsal medial margin of gonocoxa simple, medially bulging, posteromedially almost right-angled. Ventrally from posteromedial corner, there are two plate-like lobes on different levels: more dorsal lobe bears 6 strong apically curved setae laterally, more ventral lobe bears aggregation of fine setae at anteromedial corner and one strong seta at posteromedial corner. Two juxtagonostylar megasetae present, the more anterior and ventral one flame-shaped and pointed, with obscurely discernible seta-like branch marked by broken line in Fig. 1D, the more posterior and dorsal one bilobed, both arising from a large common basal body which is as long as megasetae. Gonostylus sub-rectangular, posterolateral corner drawn out into a rounded lobe, posterior and medial margin with short setae. Aedeagus broadly subtriangular, with lateral shoulders, the apex curved ventrally. Aedeagal apodemes directed laterad. Hypoproct extending posteriorly to apex of gonostyli, each side with ca. 25 normal setae on ventral surface. Cerci medially separated.

Female. Unknown.

The setose laterotergite, non-setose anterior basalare, sternite 9 laterally free from gonocoxa, and gonocoxa without a remarkable posterolateral lobe group together Manota clava sp. n. with M. caribica Jaschhof & Hippa, 2005 (Costa Rica) and M. micula Hippa & Kurina, 2013 (Ecuador, Peru). All three species have also the sternite 9 posteriorly broadly concave and a similar aggregation of setae on plate-like lobe ventrally from dorsal medial margin of gonocoxa. Parastylar lobe is distinct between all three species: large, apically broadened with 3–5 posterior setae in M. clava, large, subtriangular with three posterior setae in M. micula and small, stout with 2–3 setae posteroapically in M. caribica. The gonostylus of M. clava is subrectangular and posterolaterally drawn out while it is oval or almost circular in case of the two other species. Manota clava and M. micula have the juxtagonostylar megasetae complex with transverse and leaf-like expansions, while they are simple and pointed in M. caribica.

The specific epithet is Latin, clava [club or mace], referring to the prominent club-shaped parastylar lobe, and is a noun used as in apposition.

Holotype. Male, COLOMBIA, Valle de Cauca, PNN Farallones de Cali Cgto., La Meseta, 03°34'N, 76°40'W, 2,200 m, Malaise trap, 27. viii–10.ix.2003, S. Sania & M. Losso col., M 4570 (on slide, IAvH).

Figure 2. 

Manota multilobata sp. n. (holotype). A Hypopygium, ventral view B Hypopygium, dorsal view C Aedeagus and hypoproct, ventral view. Scale bar 0.10 mm. Abbreviations: jxs = juxtagonostylar megaseta, psl = parastylar lobe, sla = setigerous finger-like lobe anteriorly from juxtagonostylar megasetae, slp = setigerous finger-like lobe posteriorly from juxtagonostylar megasetae.

Laterotergite non-setose; anterior basalare non-setose; sternite 9 posteriorly broadly and deeply concave, anterior half fused to gonocoxa; parastylar lobe transversally oblong, with ca 20 setae; gonocoxa drawn into a short and broad posterolateral lobe; gonostylus widening apically, somewhat sunken into gonocoxa; two juxtagonostylar megasetae, ventral one flame-shaped, dorsal one twisted; two and one apically setose finger-like lobes anteriorly and posteriorly from juxtagonostylar megasetae, respectively.

Male. Colour. Head brown, face somewhat paler. Antenna light brown, including scape and pedicel. Clypeus and mouthparts yellowish. Thorax light brown. Legs yellowish, basal third of femur 3 infuscated. Wing with brownish tinge because of microtrichia; halter yellow with blackish knob. Abdomen with tergites brownish, sternites somewhat lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 2.1 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, its curved sensilla not discernible; palpomere 4 with parasegment; palpomere 5 not measurable on holotype. Number of strong postocular setae 9. Thorax. Anepisternum with 26 setae; anterior basalare, preepisternum 2 and laterotergite non-setose; metepisternum with 8 setae. Legs. Mid and hind tibial organs absent. Wing. R₁ meeting C within basal half of costal margin; sclerotized part of M₂ extending to level of tip of R₁; wing length, 2.5 mm. Hypopygium (Fig. 2A–C). Sternite 9 broad, extending to the middle of gonocoxa, anterior half fused to gonocoxa, posterior half free, posterior margin broadly and deeply concave, anterior margin shallowly and angularly incised, laterally with bare narrow area, medially setose with setae similar to the adjacent ventral setae of gonocoxa. Ventral medial margin of gonocoxa simple. Posterior margin with two long setae having prominent sockets. Parastylar lobe transversally oblong, well exposed in ventral view, bearing ca 20 setae. No paraapodemal lobe observable. Posterolateral part of gonocoxa not drawn into a remarkable lobe. Dorsal medial margin of gonocoxa simple, bulging medially, contiguous with the dorsal posterior margin. Two juxtagonostylar megasetae arising from separate basal bodies, dorsal megaseta somewhat twisted, with a basal body ca. one fourth of seta’s length, ventral megaseta flame-shaped, with a basal body slightly less than seta’s length. Two finger-like lobes anteriorly from juxtagonostylar megasetae: more anterior lobe subequal to basal body of dorsal juxtagonostylar megaseta with one seta apically, more posterior lobe subequal to basal body of ventral juxtagonostylar seta with three setae apically. Posteriorly from the juxtagonostylar megasetae, a lobe, subequal to basal body of ventral juxtagonostylar seta, bearing one strong apical seta and one weak subapical seta. Gonostylus somewhat sunken into gonocoxa, apically widening, with 5 strong and long setae at posterior and posterolateral margins, other setosity similar to that on gonocoxa ventrally. Aedeagus elongate, narrowly subtriangular, the lateral sides slightly concave, apex curved ventrally. Hypoproct extending posteriorly to level of apex of gonostyli, each side with 6 setae on apical third ventrally. Cerci broad, medially separated.

Female. Unknown.

Discussion. Manota multilobata sp. n. groups together with M. setilobata sp. n. by having the non-setose anterior basalare, non-setose laterotergite, indistinct or short posterolateral lobes of the gonocoxa, and the megasetae and aggregations of setae at the dorsal medial margin of the gonocoxa all placed far posteriorly. Both species have the obovate gonostylus, which is somewhat sunken into the gonocoxa, and have 4–5 strong apical and subapical setae deviating from other setae, similar arrangement of small setose lobes around juxtagonostylar setae, and sternite 9 basally fused with the gonocoxa. The species differ as follows: 1) in M. multilobata there are two finger-like lobes close together anteriorly from the juxtagonostylar megaseta, the more anterior one with one, the more posterior one with three strong setae (in M. setilobata there is a plate-like lobe anteriorly bearing one seta widely separated from a posterior group of several setae), 2) in M. multilobata posteriorly from the juxtagonostylar megasetae there is a finger-like lobe with one strong and one weak seta (in M. setilobata a flat lobe with numerous fine setae), 3) in M. multilobata the gonocoxa is drawn into a short and broad posterolateral lobe (in M. setilobata it is drawn into a short and narrow lobe), and 4) in M. multilobata sternite 9 has the posterior margin broadly v-shaped incised (in M. setilobata there is narrow and deep medial cleft).

The specific epithet is Latin, multilobata [many-lobed], referring to the setigerous lobes dorsally on the gonostylus (adjective).

Holotype. Male, COSTA RICA, San Isidro de las Peñas Blancas, Texas A&M Soltis Center, Malaise trap, 400 m, 10°23'00"N, 84°36'58"W, 20.iv–26. v.2010, Wendy Porras col. (on slide, MNCR).

Figure 3. 

Manota perplexa sp. n. (holotype). A Hypopygium, ventral view B Hypopygium dorsal view C Aedeagus and hypoproct, ventral view D Cerci with associated parts, dorsal view. Scale bar 0.10 mm. Abbreviations: flb = finger-like lobe, gs = gonostylus, jxs = juxtagonostylar megasetae, tm = twisted megaseta.

Laterotergite non-setose; anterior basalare non-setose; sternite 9 laterally entirely fused to gonocoxa, posterior margin free with protruding posterolateral corners; parastylar lobe indistinct; gonocoxa with a large plate-like lobe bearing four simple megasetae medioventrally from dorsal medial margin and anteriorly from the juxtagonostylar setae; gnostylus subtriangular, with prominent lateral angle; two juxtagonostylar megasetae, anterior one simple and pointed, posterior one bifurcated.

Male. Colour. Head brown, face somewhat paler. Antenna light brown, scape, pedicel and two basal flagellomeres slightly paler. Clypeus and mouthparts yellowish. Thorax light brown. Legs yellowish. Wing with light brownish tinge because of microtrichia; haltere yellow with brown knob. Abdomen with tergites dark brown to blackish, sternites yellowish. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than the finer ones. Head. Antennal flagellomere 4 ca. 1.8 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with three apically curved sensilla; palpomere 4 with parasegment; palpomere 5 ca. as long as palpomere 4. Nine strong postocular setae. Thorax. Anepisternum with 46 setae; anterior basalare, preepisternum 2 and laterotergite non-setose; metepisternum with 15 setae on anterior part. Legs. Mid and hind tibial organs absent. Wing. R₁ meeting C within basal half of costal margin; sclerotized part of M₂ extending to level of tip of R₁; wing length 1.6 mm. Hypopygium (Fig. 3A–D). Sternite 9 laterally entirely fused to gonocoxa, posterior margin free with protruding posterolateral corners; setae similar to adjacent ventral setae of gonocoxa. Ventral medial margin of gonocoxa simple, posteromedial corner drawn into a lobe, posterolateral part of gonocoxa not drawn into a lobe. Parastylar lobe indistinct, apparently represented by one seta by ventral medial margin of gonocoxa. No paraapodemal lobe observable. Dorsal medial margin of gonocoxa with a transverse shallow incision medially. A large plate-like lobe bearing four simple megasetae medioventrally from dorsal medial margin and anteriorly from the juxtagonostylar setae, anteriormost with its own basal body ca. 1/3 longer than others. Two juxtagonostylar megasetae present, more anterior one pointed, slightly curved simple megaseta, arising from a basal body which is shorter than the megaseta, more posterior one bifurcate, one of the branches whip-like, the other flat and dilated, arising subapically from a very prominent basal body which is longer than the megaseta itself and ca. as long as gonostylus. Ventrally from the more anterior juxtagonostylar megaseta there is an apically twisted megaseta. Dorsally at the posterior margin of gonocoxa a long finger-like lobe apically bearing a seta (in holotype the seta is broken on both sides). Gonostylus subtriangular, with prominent lateral angle, with one very strong and 2–3 weaker setae at posteromedial corner, and with one curved strong seta at posterior margin. Aedeagus narrowly subtriangular, the apex curved ventrally, otherwise the details not visible in the mount. Hypoproct posteriorly extending to the base of gonostyli, with ca. 35 ventral setae on each side. Cerci medially separated, with their apical parts narrowed.

Female. Unknown.

In the key to Costa Rican species by Jaschhof and Hippa (2005), M. perplexa sp. n. would run into couplet 13, because of non-setose laterotergite and absence of posterolateral lobes of gonocoxa. Due to the structure of male genitalia, M. perplexa is clearly different from the two included species, viz. M multisetosa Jaschhof & Hippa and M. tapantiensis Jaschhof & Hippa. Manota perplexa is distinguished e.g. by the lack of well-developed parastylar lobe and the presence of complicated pattern of lobes and strong setae dorsally at the medial margin of gonocoxa as well as by laterally with the gonocoxa fused sternite 9. The complex dorsomedial armature of gonocoxa is unique and easily distinguishes M. perplexa from any other described Manota species.

The specific epithet is Latin, perplexa [confused, complicated or ambiguous], referring to the very complex gonostylus and its juxtapositional structures (adjective).

Holotype. Male, COLOMBIA, Risaralda, SFF Otún Quimbaya Cuchilla Camino, 04°43'N, 75°35'W, 2050 m, Malaise trap, 08–24.v.2003, G. López Leg. M. 3673 (on slide, IAvH). Paratype. Male, same as holotype except 04–17.ii.2003, M. 3694 (on slide, IAvH).

Figure 4. 

Manota setilobata sp. n. (A, B, D, E and F holotype, C paratype). A Hypopygium, ventral view B Hypopygium, dorsal view C Right gonostylus, ventral view D Aedeagus and hypoproct, ventral view E Left side juxtagonostylar megasetae with associated parts, dorsal view F Right side juxtagonostylar megasetae. Scale bar 0.10 mm. Abbreviation: gxl = posterolateral lobe of gonocoxa.

Laterotergite non-setose; anterior basalare non-setose; sternite 9 posteriorly and anteriorly deeply incised, posterior third laterally free; parastylar lobe indistinct; posterolateral part of gonocoxa drawn into a narrow lobe; dorsomedial margin of gonocoxa with a large plate-like lobe bearing one strong seta at posteromedial corner; gonostylus elongated subquadrangular, slightly sunken into gonocoxa; two juxtagonostylar megasetae, both twisted, the more dorsal one apically flattened and dilated; posteriorly from the juxtagonostylar megasetae a narrow flat apically setose lobe.

Male. Colour. Head brown, face somewhat paler. Antenna light brown, including scape and pedicel. Clypeus and mouthparts yellowish. Thorax light brown. Legs yellowish. Wing with light brownish tinge because of microtrichia; haltere yellow with blackish knob. Abdomen with tergites dark brown to blackish, sternites light brown to yellowish. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 2.3 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with three apically curved sensilla; palpomere 4 with parasegment; palpomere 5 missing in both known specimens. Number of strong postocular setae, 10. Thorax. Anepisternum with 29–33 setae; anterior basalare, preepisternum 2 and laterotergite non-setose; metepisternum with 3–5 setae. Legs. Mid and hind tibial organs absent. Wing. R₁ meeting C within basal half of costal margin; sclerotized part of M₂ extending to level of tip of R₁; wing length, 2.4 mm. Hypopygium (Fig. 4A–F). Sternite 9 ca. 2/3 as long as gonocoxa, anterior 2/3 laterally fused to gonocoxa, posterior 1/3 free, posterior and anterior margins with deep incisions which separate the sclerite almost into two halves, covered with setae similar to adjacent ventral setae of gonocoxa. Ventral medial margin of gonocoxa simple. Parastylar lobe not identifiable with certainty, possible fused with gonocoxa and in Fig. 4A comprising the part visible between the posterior margin of sternite 9 and the gonostylus. No paraapodemal lobe observable. Posterolateral part of gonocoxa drawn into a narrow lobe. Dorsomedial margin of gonocoxa simple. In a more ventral level, a large plate-like lobe bearing one strong seta at posteromedial corner. Two juxtagonostylar megasetae present, both twisted, the more dorsal one apically flattened and dilated, both arising from basally fused basal bodies which are as long as the megasetae. Dorsally from juxtagonostylar megasetae, a flat apically setose lobe, connected with a thin, one seta bearing plate-like lobe anteriorly from it. Posteriorly from the juxtagonostylar megasetae a narrow flat apically setose lobe. Gonostylus elongated subquadrangular, slightly sunken into gonocoxa, with 4–5 strong and long setae at posterior margins, other setosity similar to that on gonocoxa ventrally, dorsal side non-setose. Aedeagus subtriangular, lateral sides slightly concave, apex curved ventrally. Hypoproct extending posteriorly over apex of gonostyli, each side with 4–5 strong setae apically and ca. 30 fine setae on ventral surface. Cerci medially separated.

Female. Unknown.

Manota setilobata sp. n. resembles M. multilobata sp. n. For a more detailed discussion on distinguishing characters, see above.

The specific epithet is Latin, setilobata [with seta-bearing lobes], referring to the apically setose lobes dorsally on the gonocoxa (adjective).

Holotype. Male, COLOMBIA, Valle de Cauca, PNN Farallones de Cali Cgto., La Meseta, 03°34'N, 76°40'W, 2200 m, Malaise trap, 27.viii–10.ix.2003, S. Sania & M. Losso col., M 4570 (on slide, IAvH). Paratypes. 2 males, same as holotype (on slides, MZUSP); 1 male, same as holotype except 24.xii.2003–27.i.2004, M 4564 (on slide, IAvH); 1 male, same as holotype except 27.i–10.ii.2004, M 4563 (on slide, IAvH); 1 male, same as holotype except 10–25.ii.2004, M 4555 (on slide, IZBE); 1 male, COLOMBIA, Huilla, PNN Cueva de los Guácharos, Alto el Mirador, 01°38'N, 76°06'W, 1980 m, Malaise trap, 6-21.iv.2002, J. Fonseca col., M 3127 (on slide, IAvH); 1 male, COLOMBIA, Cauca, PNN Gorgona, El Saman, 02°58'N, 78°11'W, 5 m, Malaise trap, 28.ix–22.x.2001, H. Torres col., M 2457 (on slide, IZBE); 1 male, same as previous except 06–23.iii.2002, R. Duque col., M 3088 (on slide, IAvH); 1 male, COLOMBIA, Risaralda, SFF Otún Quimbaya, El Molinillo, 04°43'N, 75°34'W, 2200 m, Malaise trap, 17.ii–4.iii.2003, G. López col., M 3696 (on slide, IAvH).

Figure 5. 

Manota subaristata sp. n. (A and C holotype B paratype). A Hypopygium, ventral view B Hypopygium, dorsal view C Right gonostylus, ventral view. Scale bar 0.10 mm. Abbreviations: flb = finger-like setigerous lobe, gxl = posterolateral lobe of gonocoxa.

Laterotergite non-setose; anterior basalare non-setose; sternite 9 laterally fused to gonocoxa except for posterior fifth; parastylar lobe indistinct; posterolateral part of gonocoxa drawn into a lobe; dorsomedial margin of gonocoxa with a plate-like lobe bearing one anterior and two posterior simple megasetae; gonostylus in dorsal and ventral view narrow, crescent-shaped; two juxtagonostylar megasetae, more dorsal one subbasally geniculate and apically bifurcate, more ventral one simple, slightly flattened whip-like; posteriorly from juxtagonostylar megasetae a short finger-like lobe with 3–4 strong setae.

Male. Colour. Head brown, face somewhat paler. Antenna light brown, including scape and pedicel. Clypeus and mouthparts yellowish. Thorax brown. Legs yellowish. Wing with brownish tinge because of microtrichia; halter yellow with blackish knob. Abdomen with tergites brown to dark brown, sternites somewhat lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 1.6–1.7 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 3 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 ca. 1.4–1.5 times longer than palpomere 4. Number of strong postocular setae 9–11. Thorax. Anepisternum with 42–47 setae; anterior basalare, preepisternum 2 and laterotergite non-setose; metepisternum with 8–14 setae. Legs. Mid and hind tibial organs absent. Wing. R₁ meeting C within basal half of costal margin; sclerotized part of M₂ extending to level of tip of R₁; wing length, 1.9–2.3 mm. Hypopygium (Fig. 5A–C). Sternite 9 laterally fused to gonocoxa except for posterior 1/5, extending to middle of gonocoxa, posterior margin slightly concave, anterior margin deeply incised. Posterior 1/3 of sternite 9 setose, otherwise non-setose, setae similar to adjacent ventral setae of gonocoxa. Medioventral margin of gonocoxa roundly angled. Parastylar not identifiable, apparently membranous and covered by gonocoxa, possibly with one seta visible at the gonocoxal margin in Fig. 5A. No paraapodemal lobe observable. Posterolateral part of gonocoxa drawn into a lobe ca. 1/3 length of gonocoxa. Mediodorsal margin of gonocoxa simple. A plate-like lobe with one anterior and two posterior simple megasetae medioventrally from dorsal medial margin and anteriorly from juxtagonostylar setae. Two juxtagonostylar megasetae present: more dorsal megaseta subbasally geniculate, apical part bifurcate, one of the branches whip-like, the other apically flattened and dilated; the more ventral megaseta simple, slightly flattened whip-like. Posteriorly from juxtagonostylar megasetae a short finger-like lobe with 3–4 strong setae. Gonostylus in dorsal and ventral view narrow, crescent-shaped (in some slides apically pointed), with 3–4 setae dorsally near lateral margin and 1 ventral seta near the medial margin. Aedeagus elongate subtriangular, without lateral shoulders, apex curved ventrally. Hypoproct extending posteriorly to level of base of gonostyli or slightly over, with ca. 25 ventral setae on each side. Cerci medially separate, apically slightly widened.

Female. Unknown.

Manota subaristata sp. n. is similar to M. aristata Hippa & Kurina, 2013 in having the dorsal juxtagonostylar megaseta with a long whip-like branch. Manota subaristata, however, has the megaseta subbasally geniculate, arising from a separate basal body, while it is basally straight and arising from apical half of the common basal body with the ventral juxtagonostylar seta in M. aristata. Manota subaristata has 3–4 strong setae on a finger-like setose lobe posteriorly from the juxtagonostylar megaseta, which are absent in M. aristata. By the latter character, the species resembles M. acutistylus Jaschhof & Hippa, 2005, but the megasetae at the dorsal medial margin of gonocoxa in M. subaristata are longer and there are two of them in the posterior group, not three as in M. acutistylus (see also the discussion for M. aristata in Hippa and Kurina 2013: 109). In M. subaristata, the juxtagonostylar megasetae are subequal in length, while M. acutistylus has the dorsal juxtagonostylar megaseta remarkably shorter, ca. half of the length of ventral one. All these three species have the gonostylus apically tapering in dorsal and ventral view, and sternite 9 largely fused to the gonocoxa, and may compose together a small clade within the Neotropical diversity of the genus.

The specific epithet is Latin, formed from the specific epithet of M. aristata by the prefix sub- [somewhat], as a reference to the similarity of the two species (adjective).

Figure 6. 

Distribution of the new species of Manota.