Corresponding author: Uwe Kallweit (
Academic editor: V. Blagoderov
The distribution of
The position of
This study is based primarily on material from the Senckenberg Naturhistorische Sammlungen Dresden (SNSD), partially collected by the author. Malaise trap and sweep net samples from the Bavarian Alps, and from the islands of Sakhalin and Honshu were included. The type specimen of the newly described species is deposited in Dresden.
In addition to the two species treated in detail below, one specimen of
Specimens are mounted on microscope slides or kept in 70% ethanol. The amber fossil has been preserved in polyester, following Hoffeins’ preparation method (2001). They were studied with the aid of an Olympus SZH10 stereomicroscope. For light microscope study and the preparation of drawings an Olympus BH2 was used. Photographs were taken with a digital Olympus C-3030 camera attached to Olympus SZH10 and BH-2 microscopes. SEM photos were taken by a Zeiss SmartSEMTM, Type Supra 55VP. Length measurements were taken of slide-mounted specimens only. The lengths of wing veins were assessed with the base of the stem vein as the basalmost point. The wing index, used to describe the shape of the wing, is the ratio of the wing length to its width. Morphological terminology, including abbreviations, follows
(based on Zaitzev’s original genus description, with additions).
The described genus belongs to the tribe
A distinctive species with vein M4 basally detached, thus no complete hind fork present (
Male.
Female.
2 males (on slide and in ethanol), Germany, Bavaria, Allgaeu, Mt. Ponten (Alps), East of Hinterstein, SE of Sonthofen, 1840 m,
Other material, pictures taken by electron-scanning microscope: 1 male. Japan, Hokkaido, Ogusawa-suigenchi area; near city of Otaru;
The species is characterized by a peculiar extended fore basistarsus, which is 1.5 times as long as the fore tibia. A strong basalization of radial veins has led to the nearly horizontal crossvein ta and foreshortened R1. C extending beyond apex of R5 for 1/3 distance to M1. Subcosta ending a short distance before base of ta in R. Crossvein ta not reaching the length of M-stem. A single setum chaeticum of the mid tibial organ has 2 filamentous branches (
Male.
Female unknown.
Details of specialized setae, situated at the upper outer side of tibiae. Abbreviations: Gnor =
Species | Tribe | Figures | Position of tibia | Shape of the single specialized seta | ||||||
---|---|---|---|---|---|---|---|---|---|---|
mid | hind | simple | complex | foliiform | filamentous | branched | peglike | |||
|
Gnor | 31–32 | X | X | X | |||||
|
Gnor | 33–34 | X | X | X | |||||
|
Scio | 49&55 | X | X | ||||||
|
Gnor | 45-48 | X | X | X | |||||
|
Gnor | 37-38 | X | X? | ||||||
|
Scio | 50&54 | X | X | X | |||||
|
Scio | 51&57 | X | X | X | |||||
|
Scio | 52&56 | X | X | X | |||||
|
Scio | 53&58 | X | X | X | |||||
|
Gnor | 35–36 | X | X | X | |||||
|
Gnor | 43–44 | X | X | X | |||||
Gnor | 39–40 | X | X | X | ||||||
Gnor | 41–42 | X | X | X |
The name refers to the origin of this species from Yoshiwa Village, Hiroshima Pref., Honshu, Japan.
1 | Vein M4 detached at base. With ventral gonocoxal projection only. Striking long cercal setae ( |
|
2 | M4 meets vein CuA below base of ta. With ventral & dorsal gonocoxal projection | 3 |
3 | Gonostylus of 2 separated lobes ( |
|
– | Gonostylus simple, tapered |
The study provides new records of
Males of
Despite differences in structure of terminalia, the two Palaearctic
Palaearctic species of
The subcosta in both species is long, ending immediately in front of the base of the anterior transversal vein. Males have a mid tibial organ on the dorsal basal third of the tibia, females do not. Female cerci are directed ventrad. Considering these features as a complex trait, one may conclude that
The legs of
Zaitzev (1989, p. 136) pointed to the similarity between
Zaitzev’s description of the
Terminalia of
Terminalia of
Mid tibial organ. Not on the same scale.
Mid and hind tibial organ. Not on the same scale.
Mid tibial organ. Not on the same scale
This paper is the result of a study, using the facilities of the Senckenberg Naturhistorische Sammlungen Dresden (SNSD). I would like to express my sincere appreciation to Dr Mathias Jaschhof (Senckenberg Deutsches Entomologisches Institut, Müncheberg) for providing specimens. I thank Peter J. Chandler (Melksham, UK) for reading an earlier version of this manuscript. An anonymous reviewer has pointed to the recent record of