Corresponding author: Radim Gabriš (
Academic editor: F. Vitali
The genera
Gabriš R, Trnka F, Wahab RA, Kundrata R (2017) Taxonomic revision of the endemic Bornean genera
To finish a revision of the genera classified in
In this study we examined mounted adults of both sexes. Genitalia were briefly submerged in hot 10% KOH, dissected and transferred to glycerol. Main diagnostics were photographed using a Zeiss Discovery.V12 with ZEN software. The line illustrations were derived from the photographs. All dissected parts were mounted on separate cardboards using Dimethyl Hydantoin Formaldehyde (
This genus is easily recognizable among the Bornean
Body elongate to elongate-oval, small to medium-sized. Body densely clothed with very short pubescence; coloration either more or less uniformly brown or brown with yellowish stripes ranging from vertex through sides of pronotum to basal part of elytra, mouthparts lighter; in some cases antennae reddish brown or black (Figs
Head about the same width as anterior margin of pronotum; genae sub-parallel at frontal view; frontoclypeus with distinct midline running from interantennal groove to labrum, sparsely covered with large, rounded, deep punctures; antennal tubercles prominent with deep narrow depression in between; antennal cavities opened dorsally; anterior margin of anteclypeus shallowly emarginate, with sparse long yellowish semi-erect setae. Labrum free, transverse, glabrous, either with one row of punctures bearing long setae (Figs
Prothorax subcylindrical, about as long as wide, widest before middle, then gradually narrowed towards posterior margin, laterally with one small more or less distinct tubercle; pronotal disc weakly convex, sparsely covered with deep punctures, with indistinct tubercles, anterior and posterior angles obtuse. Prosternum in front of coxae slightly shorter than diameter of coxal cavity, procoxal cavities circular, with lateral extension, narrowly separated. Scutellum transverse, widely rounded apically, about two times as wide as long. Elytra elongate, 1.6–1.8 times as long as wide at widest part, 1.7–2.1 times as long as pronotum in males and 2.0–2.3 times in females, basally slightly wider than posterior pronotal margin, widest near middle, from middle gradually tapered towards apex, fused along suture; each elytron with three rows of tubercles irregular in size, in some cases inner row forming a distinct ridge basally (Figs
Abdomen with five ventrites (Figs
(
Dorsal habitus of
(
Female, “Malaysia, Sabah, / Crocker Range 20- / IV-2007 Cope / collection” (
This species is similar to
Holotype (male).
Prothorax as long as wide, laterally with one distinct tubercle; pronotal disc with a pair of indistinct tubercles near middle and one median at second half; pronotal tubercles punctate. Prosternum in front of coxae 0.9 times shorter than diameter of coxal cavity. Scutellum transverse, two times as wide as long. Elytra elongate, 1.6 times as long as wide at widest part, 1.7 times as long as pronotum, widest near middle; each elytron with three rows of indistinct tubercles, inner row forming a distinct ridge basally; sparsely covered with large deep punctures arranged in slightly irregular rows. Legs long, slender; relative lengths of metatarsomeres 1.0 : 0.5 : 1.0 : 2.0.
Male genitalia with tegmen elongate, widest near middle, basally with very short strut; parameres elongate, 3.5 times longer than wide, apically with long setae (Fig.
Variability in males.
Female. Most characters same as for males.
Malaysia: Borneo (Sabah; Fig.
Holotype, male, “E. MALAYSIA: Sarawak / confl. Sun Oyan and / Mujong riv., E. Kapit / 50m, 18.V.1994, # 5 / Löbl & Burckhardt // Holotype //
Dorsal habitus of
Holotype (male).
Prothorax as long as wide, laterally with one moderately distinct tubercle; pronotal disc with a pair of distinct tubercles near middle and one median at second half; pronotal tubercles punctate. Prosternum in front of coxae 0.9 times shorter than diameter of coxal cavity. Scutellum transverse, two times as wide as long. Elytra elongate, 1.6 times as long as wide at widest part, 1.8 times as long as pronotum, widest near middle; each elytron with three rows of distinct tubercles (Figs
Lateral habitus of
Male genitalia with tegmen elongate, widest near middle, basally with short strut; parameres elongate, 2.9 times longer than wide, apically with long setae (Fig.
Female. Most characters same as for males.
Malaysia: Borneo (Sarawak: Kapit; Fig.
Holotype, male, [Malaysia] “SARAWAK: / 5th Division / Gn. Mulu NP // Camp 5 / Kerangas // Pitfall / trap // iv. 78, N. M. Collins / B.M.1978-11 //
This species is similar to
Holotype (male).
Prothorax as long as wide, laterally with one indistinct tubercle; pronotal disc with a pair of very indistinct tubercles near middle and one median at second half; pronotal tubercles punctate. Prosternum in front of coxae 0.9 times shorter than diameter of coxal cavity. Scutellum transverse, about two times as wide as long. Elytra elongate, 1.8 times as long as wide at widest part, 2.1 times as long as pronotum, widest near middle; each elytron with three rows of only slightly elevated tubercles (Fig.
Male genitalia with tegmen elongate, widest near middle, basally with short strut; parameres elongate, 2.3 times longer than wide, apically with tufts of short setae (Fig.
Paratype is smaller (
Female unknown.
Malaysia: Borneo (Sarawak: Gn. Mulu NP; Fig.
The specific name is a matronym in honor of Ms. Hana Gabriš Syptáková (Salisov, Czech Republic).
Holotype, male, “BRUNEI, Ulu Temburong NP / Kuala Belalong FSC /
This species is similar to
Holotype (male).
Prothorax as long as wide, laterally with one distinct tubercle; pronotal disc with a pair of moderately distinct tubercles near middle and one median at second half and one indistinct median at anterior half; pronotal tubercles punctate. Prosternum in front of coxae 0.9 times shorter than diameter of coxal cavity. Scutellum transverse, two times as wide as long. Elytra elongate, 1.8 times as long as wide at widest part, 1.9 times as long as pronotum, widest near middle; each elytron with three rows of distinct, longitudinally elongate tubercles (Figs
Male genitalia with tegmen elongate, widest before middle, basally with short strut; parameres elongate, 3.3 times longer than wide, apically with long setae (Fig.
The male paratype is larger (body length 9.4 mm, body width 2.8 mm).
Female. Most characters same as for males.
Brunei (Fig.
This species is named after Mr. I. H. Tuf (
Body slender, elongate, densely clothed with very short pubescence; coloration brown, with some parts paler, yellowish, antennae and legs reddish-brown (Figs
Head about the same width as anterior margin of pronotum; genae convex at frontal view; frontoclypeus with distinct midline running from interantennal groove to labrum, sparsely punctured; antennal tubercles prominent with narrow, deep depression in between; antennal cavities opened dorsally; anterior margin of anteclypeus shallowly emarginate, with sparse long yellowish semi-erect setae. Labrum free, transverse, glabrous, with a row of distinct punctures and sparsely and irregularly distributed additional less distinct punctures, with sparse long semi-erect setae (Fig.
Prothorax about as long as wide, subparallel-sided at anterior half, widest slightly medially, then gradually narrowed towards posterior margin, laterally with one very weakly developed tubercle; pronotal disc weakly convex, sparsely covered with deep punctures, not smooth, without tubercles (Fig.
Abdomen with five visible ventrites, first ventrite (excluding intercoxal process) almost 1.5 times longer than second; intercoxal process subparallel-sided basally, narrowed and broadly rounded apically. Fifth ventrite with apex truncate, margin with sparse semi-erect pubescence. Male genitalia with tegmen elongate, widest at apical third, basally with long strut; parameres moderately long, setose apically. Penis relatively long, with dorsal struts diverged from about two fifths of penis length (Fig.
Holotype, male, “Samarahan / June 1906 [handwritten] // Type. //
Holotype (male).
Prothorax 1.1 times as long as wide, laterally with one very weakly developed tubercle; pronotal disc without tubercles (Fig.
Male genitalia with tegmen elongate, widest at apical third, basally with long strut; parameres moderately long. Penis long, apically broadly rounded; dorsal struts diverged from about two fifths of penis length (Fig.
Malaysia: Borneo (Sarawak: Samarahan; Fig.
1 | Body uniformly brown (Figs |
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– | Body brown with yellowish stripes dorsally (Figs |
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2 | Body length 7.0–8.0 mm; length ratio of antennomeres I–III 4.4–4.6 : 1.7–1.9 : 1.0 (Fig. |
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– | Body length 9.2–9.9 mm; length ratio of antennomeres I–III 6.3–6.5 : 2.5–2.7 : 1.0 (Figs |
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3 | Apex of the scape with distinct lateral hook-shaped projection (Figs |
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– | Apex of the scape simple (Figs |
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Genus
Genus
Genus
Genus
Genus
Genus
In 2013, two Czech universities (Palacky University in Olomouc, University of Ostrava) and the Universiti Brunei Darussalam established a collaboration which resulted in the biodiversity survey of the Ulu Temburong National Park in Brunei (
The distribution of
We thank M. Barclay, M. Geiser (