Corresponding authors: Yongjie Wang (
Academic editor: S. Winterton
A new species of osmylid (
Dong M, Xu H, Wang Y, Jia C, Liu Z (2016) Revision of the genus
Herein the genus
All the described specimens are deposited in the Entomological Museum of
India: Sikkim.
Head brown or dark brown; compound eyes black; antennae shorter than half length of forewing; prothorax black and length longer than width, with yellow setae; meso- and metathorax dark brown with long setae; legs yellow with short brown setae; forewings oblong and subacute at apex, with few brown spots; nygmata clear surrounded by light brown spot; veins thickened; costal cross-veins simple and occasionally bifurcate; forewing Rs with 8–15 branches, distal to the base of wing; cross-veins among Rs branches forming two or three series of gradates; forewing M branching more basally than the divergence of basal branch of Rs; no more than four cross-veins present between the two branches of M; forewing Cu bifurcating near the base of wing; CuA and CuP with numerous pectinate branches; CuP longer than half length of CuA; hind wings similar to forewings in size and shape without any spot apart from pterostigma; hindwing M branching near the base of wing; hindwing CuA with numerous pectinate branches; CuP simple and shorter than half length of CuA; male genitalia with 9th tergite narrow, and sternite approximately quadrate; ectoproct relatively large, callus cerci rounded and located at the middle or underside of ectoproct; male genitalia composed of gonarcus, entoprocesses and mediuncus, arched gonarcus similar to other genera of
Although the type species of
Distribution of
1 | Head and thorax with spots or stripes |
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– | Head and thorax without any spot and stripe; pronotum yellowish brown without any stripe, meso- and metanotum dark brown; cross-veins |
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2 | Mediuncus with a process at base (Fig. |
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– | Mediuncus without any process at base, membrane of wings only with a few spots |
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3 | Forewings with many spots including along the gradate cross-veins |
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– | Forewings with few spots in the membrane (Fig. |
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4 | More than 11 branches of Rs |
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– | 11 or fewer branches of Rs |
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5 | An oblique brown stripe present from pterostigma to the outer margin, Cu with four brown spots; pronotum with two pale yellow longitudinal stripes, mesonotum brown anteriorly |
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– | No oblique stripes present from pterostigma to the outer margin |
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6 | Pronotum black with a black longitudinal stripe medially; meso- and metanotum black; Rs with 14–17 branches; Cu edged with 3–4 yellow spots; apex of gonarcus bent upwards and relative long in lateral view |
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– | Pronotum brown with a yellowish longitudinal stripe medially, meso- and metanotum brown; vertex with a Y-shaped mark |
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7 | Membrane of wings yellow and gradate cross-veins not edged with spots (Fig. |
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– | Membrane of wings hyaline and gradate cross-veins edged with brown |
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8 | Apex of mediuncus short, broad and flat in dorsal view (Figs |
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– | Apex of mediuncus long, acute and protuberant in dorsal view (Figs |
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Holotype Male. CHINA: Shaanxi (Province): Taibai, [
Pronotum with four brown spots at anterior margin, two round brown spots in middle and two brown spots at posterior margin; membrane hyaline, pterostigma yellow, without dark spots besides the pterostigma; veins light yellow at base but brown from middle to the end; mediuncus C-shaped in lateral view with a basal process and boat-shaped in dorsal view.
Body length 8.6 mm.
Wings of
China (Shaanxi).
The specific name refers to the process at the base of the mediuncus.
This species is known from its type locality, Shaanxi province. It is easily distinguished from other species in the genus because of the body coloration, morphology and the genital characters.
Holotype male. CHINA: Yunnan (Province): Lushuixian Gangfang, [
Pronotum black with two yellowish longitudinal stripes at anterior margin; mesonotum with two yellow spots in middle; apex of gonarcus bent upwards and relative long in lateral view.
Body length 9–11 mm.
Wings of
China (Tibet, Yunnan, Sichuan)
Although the original identification of both
Holotype Male, CHINA: Tibet: Yadong, [
Pronotum with two yellow stripes in middle, an oblique brown mark presenting from the pterostigma to the posterior outer margin of forewing; base of mediuncus slender and apex dilated as lobe-shape in lateral view.
Body length 8.0 mm.
Wings of
China (Tibet).
Holotype Male. CHINA: Sichuan (Province): Wolong, [
Pronotum with two yellow narrow stripes in middle; metanotum with two brown spots on anterior margin; gonarcus sclerotized distally and bent upward and hook-shaped in lateral view; base of mediuncus approximately finger-shaped in lateral view.
Body length 9–11 mm.
Wings of
China (Sichuan, Shaanxi, Gansu, Henan).
Holotype Female. CHINA: Yunnan (Province): Lushui, Yaojiapin, [
Pronotum with a dark brown longitudinal stripe medially; metanotum with a brown spot in middle; forewing light yellow; mediuncus C-shaped in lateral view and scoop-shaped in dorsal view.
Body length 7–9 mm.
Wings of
China (Yunnan).
Holotype Male. CHINA: Hubei (Province): Shennongjia, [
Pronotum with two narrow longitudinal dark brown marks in middle; ectoproct with a dorsal coniform process in lateral view; gonarcus with a short finger-like process distally in lateral view.
Body length 8–10 mm.
Wings of
China (Henan, Shaanxi, Hubei, Chongqing).
Holotype Male. CHINA: Fujian (Province): Wuyishan, Huanghuacong, [
Two yellow longitudinal stripes present from pronotum to mesonotum. mesoscutellum bright yellow, metanotum with a central yellow stripe; apex of gonarcus slightly dilated, short and curved dorsally.
Body length 7–10 mm.
Wings of
China (Fujian).
We wish to express our gratitude to Timothy R. New and Valdimir Makarkin for providing literature necessary to this project. This research was supported by the National Natural Science Foundation of China (Gants 31272352, 31301905 and 41372013), the Beijing Natural Science Foundation (Grant 5132008), and Ph.D. Programs Foundation of Ministry of Education of China (Grant 20131108120005).