Molecular clock dating of mitochondrial COI gene with BEAST ver. 1.8.0. The oldest divergence between A. yushanensis and the other Apenetretus species occurred at 1.81 million years ago (mya); the divergence between A. hsueshanensis and the group of A. smetanai and A. nanhutanus occurred at 0.94 mya; and the divergence between A. smetanai and A. nanhutanus occurred at 0.53 mya.

 
 
  Part of: Weng Y-M, Yeh W-B, Yang M-M (2016) A new species of alpine Apenetretus Kurnakov from Taiwan: evidences from DNA barcodes and morphological characteristics (Coleoptera, Carabidae, Patrobini). ZooKeys 584: 121-134. https://doi.org/10.3897/zookeys.584.6320