Onthophagini topology, with characters and states shown and based on unweighted data. Inserted clades are the resolved topologies found with Piwe weighting using K values of 11-30.

 
 
  Part of: Philips TK (2016) Phylogeny of the Oniticellini and Onthophagini dung beetles (Scarabaeidae, Scarabaeinae) from morphological evidence. ZooKeys 579: 9-57. https://doi.org/10.3897/zookeys.579.6183