Surveyed caves in Du’an Karst from which trechine beetles were discovered. a Qiaoxu Dong (Qibainong) b Lubian Dong (Xia’ao) c Shuiyuan Dong (Longfu) d Jinzhu Dong I and Jinzhu Dong II e Nongguanshang Dong II f Baxian Dong (Chengjiang) g Diaomao Dong (Chengjiang) h Nongzhong Dong I (Chengjiang) i Nongqu Dong (Longwan) j Longhuan Dong (Longwan) k Lapo Dong I (Lalie).