Ingroup "Oniticellini :" "Helictopleurina :" "Helictopleurus quadripunctatus Olivier";Madagascar "Helictopleurus giganteus Harold";Madagascar "Helictopleurus neoamplicollis Krell";Madagascar "Drepanocerina :" "Scaptocnemis segregis Péringuey";"East Africa" "Cyptochirus ambiguous (Kirby)";"South Africa" "Afrodrepanus impressicollis (Boheman)";"South Africa" "Oniticellina :" "Anoplodrepanus reconditus (Matthews)";Jamaica "Tragiscus dimidiatus Klug";"South Africa" "Paroniticellus festivus Steven";Turkey "Oniticellus pictus (Hausmann)";"South Africa" "Euoniticellus intermedius (Reiche)";"South Africa" "Liatongus militaris Castelnau";"South Africa" "Liatongus californicus (Horn)";USA "Attavicinus monstrosus (Bates)";Mexico "Tiniocellus spinipes Roth";"South Africa" "Onthophagini :" "Alloscelina :" "Alloscelus paradoxus Boucomont";"Haut-Uele, Moto, DRC" "Haroldius convexus (Philips & Scholtz)";"South Africa" "Subtribe undefined:" "Amietina eburnea Cambefort";"Ivory Coast" "Caccobius ferrugineus Fahraeus";Botswana "Cassolus humeralis Arrow";"Mangpo, Darjeeling, India" "Cleptocaccobius schaedlei (d’Orbigny)";Kenya "Diastellopalpus thomsoni Bates";Malawi "Digitonthophagus gazella (Fabricius)";"South Africa" "Euonthophagus sp.";"South Africa" "Eusaproecius tinantae (Boucomont)";DRC "Heteroclitopus annamariae Branco";Nigeria "Hyalonthophagus alcyon (Klug)";"South Africa" "Megaponerophilus megaponerae Brauns";"Lulua, Sandoa, DRC" "Milichus apicalis Fahraeus";"South Africa" "Mimonthophagus hinnulus (Klug)";Madagascar "Onthophagus depressus Harold";"South Africa" "Onthophagus hecate (Panzer)";"North America" "Onthophagus capella Kirby";Australia "Phalops sp.";"South Africa" "Pinacotarsus dohrni Harold";"Haut-Uele, Paulis, DRC" "Proagoderus lanista Castelnau";"South Africa" "Pseudosaproecius validicornis Quedenfeldt";Tanzania "Sukelus jessopi (Branco)";Kenya "Stiptopodius doriae Harold";"Gemu-Gofa Prov., Ethiopia" "Strandius lenzi (Harold)";Japan "Tomogonus crassus (d’Orbigny)";"Masinga (Mayombe), DRC?" Outgroups "Eurysternini :" "Eurysternus sp.";"Costa Rica" "Sisyphini :" "Sisyphus sp.";"South Africa" "Onitini :" "Onitis fulgidus Klug";"South Africa"